The Echinoid Directory

• Test rigid with ambulacra firmly sutured to interambulacra.
• Apical disc and peristome about the same size; both ca. 50% test diameter; plating of both systems unknown.
• Primary interambulacral tubercles perforate and crenulate with incised areoles; areoles discrete and separate except adorally.
• Scrobicular tubercles hardly differentiated; extrascrobicular zones with scattered secondary tubercles.
• Ambulacra weakly sinuous; pore zones weakly incised; pore-pairs conjugate with the two pores widely separated.
• Ambital and adapical ambulacral plating simple, but with every third tubercle enlarged. Adoral plating with large primary tubercle on every second or third element.
• Spines; none definitely attached to tests. Possibly long, slender and cylindrical; neck short and shaft ornamented with beaded ribs.

Jurassic (Pliensbachian to Kimmeridgian); Europe.

Cidaris parandieri Agassiz, 1840, p. 58 [=Cidarites blumenbachii Munster, in Goldfuss, 1829], by original designation.

• Vadet (1988, 1991) included the following in this genus:

   

• N. parandieri (Agassiz, 1840); Oxfordian, western Europe [N. blumenbachi (Munster, in Goldfuss, 1829); Oxfordian, Europe (see Radwanska 2003)].
• N. moeschi (de Loriol, 1869); Oxfordian-Kimmeridgian, Europe.
• N. smithii (Wright, 1857): Oxfordian - Kimmeridgian, UK, France.
• N. pichauriensis (Lambert, 1925); Pliensbachian, France.
• N. zschokkei (Cotteau, 1875); (Aalenian?) Bajocian, western Europe.
• N. desori (Cotteau, 1857); Bathonian-Callovian, France.
• N. poucheti (Desor, 1858); Oxfordian-Kimmeridgian, France.
• N. spinulosa (Cotteau, 1875); Bajocian, France, UK.
• N. microstoma (Cotteau, 1875); Bathonian-Callovian, France, UK.
• N. leberti (Vadet, 1989); Callovian, France.

Cidaroida, Polycidaridae.

Subjective junior synonym of Paracidaris (Anisocidaris) Thiery, 1928.

Very similar to Paracidaris especially in ambulacral tuberculation, but ambulacra straight rather than strongly sinuous, and with pore-pairs subconjugate with the two pores widely separate by a raised partition. Differs from Alpicidaris only in having crenulate rather than non-crenulate tuberculation. Polycidaris has got somewhat imbricate adradial sutures and its primary tubercles have confluent areoles on ambital and adoral plates. Histocidaris has simple, uniform ambulacral tuberculation throughout and Zbindenicidaris has simple ambulacral tuberculation ambitally and adapically.

Anisocidaris Thiery, 1928, is a genus based on Cidaris bajociensis, whose neotype and only known specimen is a 15 mm individual in the Lambert collection. This has the same ambulacral structure as Nenoticidaris parandieri, the type species of Nenoticidaris, and rather similar large, circular areoles. However, the small size of the neotype makes definitive assignment very difficult. Vadet (1991) claimed that Anisocidaris bajociensis was simply a juvenile Cidaris spinulosa Cotteau (although this is by no means certain, since the two look very different). Since the defining characters of Nenoticidaris Vadet, 1988 rest on its heterogeneous ambulacral tuberculation, subconjugate pore-pairs and large primary tubercles with small perforate mamelons and crenulate platforms, Anisocidaris Thiery, 1928, is clearly part of this group and the senior subjective synonym.

Vadet, A. 1988. Revision des "Cidaris" de l\'Oxfordien et du Kimmeridgien Europeens. Memoires de la Societe academique du Boulonnais 4, 1-148, pls 1-25.

Vadet, A. 1991. Revision des "Cidaris" du Lias et du Dogger Europeens. Memoires de la Societe academique du Boulonnais 10, 1-176.

Radwanska, U. 2003. The Kimmeridgian echinoids of Poland. Acta Geologica Polonica 53, 143-165.