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Universal Chalcidoidea Database

Taxonomic tree

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show families in superfamily <Serphitoidea> Serphitoidea
show families in superfamily <Mymarommatoidea> Mymarommatoidea
hide families Chalcidoidea
show taxa in family <Agaonidae> Agaonidae
show taxa in family <Aphelinidae> Aphelinidae
show taxa in family <Azotidae> Azotidae
show taxa in family <Chalcididae> Chalcididae
show taxa in family <Cynipencyrtidae> Cynipencyrtidae
show taxa in family <Diversinitidae> Diversinitidae
show taxa in family <Encyrtidae> Encyrtidae
show taxa in family <Eriaporidae> Eriaporidae
show taxa in family <Eucharitidae> Eucharitidae
hide Eulophidae
show species in genus <Acanthala> Acanthala
show species in genus <Aceratoneura> Aceratoneura
show species in genus <Aceratoneuromyia> Aceratoneuromyia
show species in genus <Achrysocharoides> Achrysocharoides
show species in genus <Acrias> Acrias
show species in genus <Afrotroppopsis> Afrotroppopsis
show species in genus <Agmostigma> Agmostigma
show species in genus <Aleuroctonus> Aleuroctonus
show species in genus <Alibertia> Alibertia
show species in genus <Allocerastichus> Allocerastichus
show species in genus <Alophomorphella> Alophomorphella
show species in genus <Alophomyia> Alophomyia
show species in genus <Alveoplectrus> Alveoplectrus
show species in genus <Ambocybe> Ambocybe
show species in genus <Ametallon> Ametallon
show species in genus <Anaprostocetus> Anaprostocetus
show species in genus <Anselmella> Anselmella
show species in genus <Anumanniola> Anumanniola
show species in genus <Aoridus> Aoridus
show species in genus <Apleurotropis> Apleurotropis
show species in genus <Apotetrastichus> Apotetrastichus
show species in genus <Aprostocetus> Aprostocetus
show species in genus <Aprostoporoides> Aprostoporoides
show species in genus <Apterastichus> Apterastichus
show species in genus <Arabiola> Arabiola
show species in genus <Arachnolophus> Arachnolophus
show species in genus <Arachnoobius> Arachnoobius
show species in genus <Aranobroter> Aranobroter
show species in genus <Aroplectrus> Aroplectrus
show species in genus <Ascotolinx> Ascotolinx
show species in genus <Asecodes> Asecodes
show species in genus <Astichomyiia> Astichomyiia
show species in genus <Astichus> Astichus
show species in genus <Aulogymnus> Aulogymnus
show species in genus <Austeulophus> Austeulophus
show species in genus <Australsecodes> Australsecodes
show species in genus <Awara> Awara
show species in genus <Baeoentedon> Baeoentedon
show species in genus <Baryscapus> Baryscapus
show species in genus <Bellerus> Bellerus
show species in genus <Benoitius> Benoitius
show species in genus <Beornia> Beornia
show species in genus <Borneoastichus> Borneoastichus
show species in genus <Boucekastichus> Boucekastichus
show species in genus <Boucekelimus> Boucekelimus
show species in genus <Boucekiola> Boucekiola
show species in genus <Bridarolliella> Bridarolliella
show species in genus <Cabeza> Cabeza
show species in genus <Careostrix> Careostrix
show species in genus <Carlyleia> Carlyleia
show species in genus <Ceranisus> Ceranisus
show species in genus <Ceratoneura> Ceratoneura
show species in genus <Ceratoneuronella> Ceratoneuronella
show species in genus <Ceratoneuropsis> Ceratoneuropsis
show species in genus <Chaenotetrastichus> Chaenotetrastichus
show species in genus <Chouioia> Chouioia
show species in genus <Chrysocharis> Chrysocharis
show species in genus <Chrysocharodes> Chrysocharodes
show species in genus <Chrysonotomyia> Chrysonotomyia
show species in genus <Chytrolestes> Chytrolestes
show species in genus <Cirrospiloidelleus> Cirrospiloidelleus
show species in genus <Cirrospilopsis> Cirrospilopsis
show species in genus <Cirrospilus> Cirrospilus
show species in genus <Citrostichus> Citrostichus
show species in genus <Closterocerus> Closterocerus
show species in genus <Clotildiella> Clotildiella
show species in genus <Clypecharis> Clypecharis
show species in genus <Clypomphale> Clypomphale
show species in genus <Cobarus> Cobarus
show species in genus <Colpixys> Colpixys
show species in genus <Colpoclypeus> Colpoclypeus
show species in genus <Comastichus> Comastichus
show species in genus <Cornugon> Cornugon
show species in genus <Crataepus> Crataepus
show species in genus <Cristelacher> Cristelacher
show species in genus <Cryptastichus> Cryptastichus
show species in genus <Cucarastichus> Cucarastichus
show species in genus <Dahlbominus> Dahlbominus
show species in genus <Dapsilothrix> Dapsilothrix
show species in genus <Dasyeulophus> Dasyeulophus
show species in genus <Dasyomphale> Dasyomphale
show species in genus <Davincia> Davincia
show species in genus <Dentalion> Dentalion
show species in genus <Dermatopelte> Dermatopelte
show species in genus <Derostenoides> Derostenoides
show species in genus <Derostenus> Derostenus
show species in genus <Deutereulophus> Deutereulophus
show species in genus <Deutereulophus> Deutereulophus
show species in genus <Diaulinopsis> Diaulinopsis
show species in genus <Diaulomorpha> Diaulomorpha
show species in genus <Dichatomus> Dichatomus
show species in genus <Dicladocerus> Dicladocerus
show species in genus <Diglyphomorpha> Diglyphomorpha
show species in genus <Diglyphomorphomyia> Diglyphomorphomyia
show species in genus <Diglyphus> Diglyphus
show species in genus <Dimmockia> Dimmockia
show species in genus <Dineulophus> Dineulophus
show species in genus <Dinopteridion> Dinopteridion
show species in genus <Driopteron> Driopteron
show species in genus <Dubeyiella> Dubeyiella
show species in genus <Dzhanokmenia> Dzhanokmenia
show species in genus <Elachertomorpha> Elachertomorpha
show species in genus <Elachertus> Elachertus
show species in genus <Elasmus> Elasmus
show species in genus <Emersonella> Emersonella
show species in genus <Emersonella> Emersonella
show species in genus <Encyrtomphale> Encyrtomphale
show species in genus <Enneastichus> Enneastichus
show species in genus <Entedon> Entedon
show species in genus <Entedonomphale> Entedonomphale
show species in genus <Entedononecremnus> Entedononecremnus
show species in genus <Epichrysoatomus> Epichrysoatomus
show species in genus <Epichrysocharis> Epichrysocharis
show species in genus <Eprhopalotus> Eprhopalotus
show species in genus <Eriastichus> Eriastichus
show species in genus <Euceratoneura> Euceratoneura
show species in genus <Euderomphale> Euderomphale
show species in genus <Euderus> Euderus
show species in genus <Eulophinusia> Eulophinusia
show species in genus <Eulophomorpha> Eulophomorpha
show species in genus <Eulophomyia> Eulophomyia
show species in genus <Eulophoscotolinx> Eulophoscotolinx
show species in genus <Eulophus> Eulophus
show species in genus <Euplectromorpha> Euplectromorpha
show species in genus <Euplectrophelinus> Euplectrophelinus
show species in genus <Euplectrus> Euplectrus
show species in genus <Eupronotius> Eupronotius
show species in genus <Eurycephaloplectrus> Eurycephaloplectrus
show species in genus <Exalarius> Exalarius
show species in genus <Exastichus> Exastichus
show species in genus <Farooqiella> Farooqiella
show species in genus <Galeopsomyia> Galeopsomyia
show species in genus <Gallastichus> Gallastichus
show species in genus <Gallowayia> Gallowayia
show species in genus <Gasterichus> Gasterichus
show species in genus <Gattonia> Gattonia
show species in genus <Gautamiella> Gautamiella
show species in genus <Ginsiella> Ginsiella
show species in genus <Goetheana> Goetheana
show species in genus <Goethella> Goethella
show species in genus <Grassator> Grassator
show species in genus <Grotiusomyia> Grotiusomyia
show species in genus <Gyrolasomyia> Gyrolasomyia
show species in genus <Hadranellus> Hadranellus
show species in genus <Hadrotrichodes> Hadrotrichodes
show species in genus <Hakuna> Hakuna
show species in genus <Hamonia> Hamonia
show species in genus <Hayatiola> Hayatiola
show species in genus <Hemiptarsenus> Hemiptarsenus
show species in genus <Henryana> Henryana
show species in genus <Holarcticesa> Holarcticesa
show species in genus <Holcotetrastichus> Holcotetrastichus
show species in genus <Hoplocrepis> Hoplocrepis
show species in genus <Horismenoides> Horismenoides
show species in genus <Horismenus> Horismenus
show species in genus <Horismenus> Horismenus
show species in genus <Hubbardiella> Hubbardiella
show species in genus <Hyssopus> Hyssopus
show species in genus <Idukkiella> Idukkiella
show species in genus <Iniostichus> Iniostichus
show species in genus <Inti> Inti
show species in genus <Ionympha> Ionympha
show species in genus <Itahipeus> Itahipeus
show species in genus <Janicharis> Janicharis
show species in genus <Kiggaella> Kiggaella
show species in genus <Klyngon> Klyngon
show species in genus <Kocaagizus> Kocaagizus
show species in genus <Kocourekia> Kocourekia
show species in genus <Kokandia> Kokandia
show species in genus <Kolopterna> Kolopterna
show species in genus <Kostjukovius> Kostjukovius
show species in genus <Kostjurixia> Kostjurixia
show species in genus <Lasalleola> Lasalleola
show species in genus <Leprosa> Leprosa
show species in genus <Leptocybe> Leptocybe
show species in genus <Lisseurytomella> Lisseurytomella
show species in genus <Makarora> Makarora
show species in genus <Megaceratoneura> Megaceratoneura
show species in genus <Melittobia> Melittobia
show species in genus <Melittobiopsis> Melittobiopsis
show species in genus <Meruacesa> Meruacesa
show species in genus <Mesofrons> Mesofrons
show species in genus <Mestocharella> Mestocharella
show species in genus <Mestocharis> Mestocharis
show species in genus <Metaplectrus> Metaplectrus
show species in genus <Microdonophagus> Microdonophagus
show species in genus <Microlycus> Microlycus
show species in genus <Minotetrastichus> Minotetrastichus
show species in genus <Miotropis> Miotropis
show species in genus <Mischotetrastichus> Mischotetrastichus
show species in genus <Mohaniella> Mohaniella
show species in genus <Monteithius> Monteithius
show species in genus <Monterrondo> Monterrondo
show species in genus <Moona> Moona
show species in genus <Myrmobomyia> Myrmobomyia
show species in genus <Myrmokata> Myrmokata
show species in genus <Narendrania> Narendrania
show species in genus <Naumanniola> Naumanniola
show species in genus <Necremnoides> Necremnoides
show species in genus <Necremnus> Necremnus
show species in genus <Neoaceratoneura> Neoaceratoneura
show species in genus <Neochrysocharis> Neochrysocharis
show species in genus <Neogasterichus> Neogasterichus
show species in genus <Neohyperteles> Neohyperteles
show species in genus <Neomestocharella> Neomestocharella
show species in genus <Neopediobopsis> Neopediobopsis
show species in genus <Neopomphale> Neopomphale
show species in genus <Neotrichoporoides> Neotrichoporoides
show species in genus <Nesolynx> Nesolynx
show species in genus <Nesympiesis> Nesympiesis
show species in genus <Notanisomorphella> Notanisomorphella
show species in genus <Noyesius> Noyesius
show species in genus <Obesulus> Obesulus
show species in genus <Ogmoelachertus> Ogmoelachertus
show species in genus <Omphale> Omphale
show species in genus <Omphalentedon> Omphalentedon
show species in genus <Oncastichus> Oncastichus
show species in genus <Oomyzus> Oomyzus
show species in genus <Opeuderus> Opeuderus
show species in genus <Ophelimus> Ophelimus
show species in genus <Oradis> Oradis
show species in genus <Oxycantha> Oxycantha
show species in genus <Oxypracetus> Oxypracetus
show species in genus <Palmistichus> Palmistichus
show species in genus <Paphagus> Paphagus
show species in genus <Parachrysocharis> Parachrysocharis
show species in genus <Paracrias> Paracrias
show species in genus <Paragaleopsomyia> Paragaleopsomyia
show species in genus <Parahorismenus> Parahorismenus
show species in genus <Paraolinx> Paraolinx
show species in genus <Parasecodella> Parasecodella
show species in genus <Parasecodes> Parasecodes
show species in genus <Paraspalangia> Paraspalangia
show species in genus <Paratetrastichus> Paratetrastichus
show species in genus <Pareulophites> Pareulophites
show species in genus <Parzaommomyia> Parzaommomyia
show species in genus <Pasohstichus> Pasohstichus
show species in genus <Pauahiana> Pauahiana
show species in genus <Peckelachertus> Peckelachertus
show species in genus <Pediobius> Pediobius
show species in genus <Pediobopsis> Pediobopsis
show species in genus <Pediocharis> Pediocharis
show species in genus <Pelorotelus> Pelorotelus
show species in genus <Pentastichodes> Pentastichodes
show species in genus <Pentastichus> Pentastichus
show species in genus <Perditorulus> Perditorulus
show species in genus <Perinetia> Perinetia
show species in genus <Perthiola> Perthiola
show species in genus <Petalidion> Petalidion
show species in genus <Petiolacus> Petiolacus
show species in genus <Phymastichus> Phymastichus
show species in genus <Piekna> Piekna
show species in genus <Planotetrastichus> Planotetrastichus
show species in genus <Platocharis> Platocharis
show species in genus <Platyplectrus> Platyplectrus
show species in genus <Platytetracampe> Platytetracampe
show species in genus <Pleurotropopseus> Pleurotropopseus
show species in genus <Pleurotroppopsis> Pleurotroppopsis
show species in genus <Pnigalio> Pnigalio
show species in genus <Pomphale> Pomphale
show species in genus <Pracetus> Pracetus
show species in genus <Proacrias> Proacrias
show species in genus <Pronotalia> Pronotalia
show species in genus <Propodeochertus> Propodeochertus
show species in genus <Pseudiglyphus> Pseudiglyphus
show species in genus <Pseudiglyphus> Pseudiglyphus
show species in genus <Pseudosecodes> Pseudosecodes
show species in genus <Puklina> Puklina
show species in genus <Quadrastichodella> Quadrastichodella
show species in genus <Quadrastichus> Quadrastichus
show species in genus <Renaniana> Renaniana
show species in genus <Rhicnopelte> Rhicnopelte
show species in genus <Ryhonos> Ryhonos
show species in genus <Sanyangia> Sanyangia
show species in genus <Sarasvatia> Sarasvatia
show species in genus <Schizocharis> Schizocharis
show species in genus <Selitrichodes> Selitrichodes
show species in genus <Semielacher> Semielacher
show species in genus <Setelacher> Setelacher
show species in genus <Seyrigina> Seyrigina
show species in genus <Shardiella> Shardiella
show species in genus <Sifraneurus> Sifraneurus
show species in genus <Sigmoepilachna> Sigmoepilachna
show species in genus <Sigmophora> Sigmophora
show species in genus <Skoka> Skoka
show species in genus <Sphenolepis> Sphenolepis
show species in genus <Sporrongia> Sporrongia
show species in genus <Stenomesius> Stenomesius
show species in genus <Stenopetius> Stenopetius
show species in genus <Stepanovia> Stepanovia
show species in genus <Stipecarinata> Stipecarinata
show species in genus <Styotrichia> Styotrichia
show species in genus <Sureshanella> Sureshanella
hide species Sympiesis
abalisis Narendran
show synonyms of <Sympiesis aburiana> aburiana (Waterston)
show synonyms of <Sympiesis acalle> acalle (Walker)
acicus Narendran
acrobasidis Miller
show synonyms of <Sympiesis albiflava> albiflava Fahringer
show synonyms of <Sympiesis albiscapus> albiscapus (Erdos)
albiventris Storozheva
ampla Storozheva
ancylae Girault
show synonyms of <Sympiesis angustipennis> angustipennis (Erdos)
aperta Storozheva
argenticoxae Girault
show synonyms of <Sympiesis asecta> asecta Delucchi
aureolus Szelenyi
show synonyms of <Sympiesis australiensis> australiensis (Girault)
bardisis Narendran
show synonyms of <Sympiesis biroi> biroi (Erdos)
show synonyms of <Sympiesis boasi> boasi (Girault)
bukobensis Ferriere
caliginosa Storozheva
show synonyms of <Sympiesis campbellensis> campbellensis (Kerrich and Yoshimoto)
show synonyms of <Sympiesis capeki> capeki Boucek
chaliloides Yao, Yang and Li
chenopodii Ashmead
clausenis Narendran
closterae Yao, Yang and Li
comosus Kerrich
comperei Crawford
show synonyms of <Sympiesis consona> consona (Girault)
corrugata Szelenyi
show synonyms of <Sympiesis cosmopterygi> cosmopterygi (Risbec)
crinita Storozheva
show synonyms of <Sympiesis cyaneipurpurea> cyaneipurpurea (Girault)
deminuta Storozheva
derogatae Kamijo
show synonyms of <Sympiesis dolichogaster> dolichogaster Ashmead
dzhebelicus Myartseva and Kurashev
earina Storozheva
elaeagni Myartseva
show synonyms of <Sympiesis elatisis> elatisis Narendran
enargiae Miller
show synonyms of <Sympiesis euspilapterygis> euspilapterygis (Erdos)
extera Storozheva
factitata Storozheva
show synonyms of <Sympiesis feketei> feketei Gyorfi
femorata Gijswijt
festiva Storozheva
show synonyms of <Sympiesis flavopicta> flavopicta Boucek
fragariae Miller
show synonyms of <Sympiesis fucosa> fucosa (Girault)
show synonyms of <Sympiesis gordius> gordius (Walker)
show synonyms of <Sympiesis gordius> gordius (Walker)
show synonyms of <Sympiesis gracillariae> gracillariae (Chambers)
grahami Erdos
show synonyms of <Sympiesis gregori> gregori Boucek
show synonyms of <Sympiesis grotii> grotii (Girault)
grunini Storozheva
show synonyms of <Sympiesis gyorfii> gyorfii Erdos
harithi Narendran
show synonyms of <Sympiesis hawaiiensis> hawaiiensis (Ashmead)
helvetica Szelenyi
show synonyms of <Sympiesis hirticula> hirticula Kamijo
show synonyms of <Sympiesis hyblaeae> hyblaeae Surekha
hyplosis Narendran
show synonyms of <Sympiesis ibseni> ibseni (Girault)
japonica Kamijo
show synonyms of <Sympiesis javanica> javanica (Ferriere)
show synonyms of <Sympiesis javensis> javensis (Gahan)
jeolikotensis Khan, Agnihotri and Sushil
show synonyms of <Sympiesis kampalana> kampalana (Ferriere)
karagiosis Myartseva and Kurashev
kazmii Narendran
kelebiana Erdos
konae Ashmead
kottiyooricus Narendran
kyurendagensis Myartseva and Kurashev
laetus Masi
laevifrons Kamijo
lehri Storozheva
lepida Storozheva
lesnei Risbec
lindbergi Ferriere
show synonyms of <Sympiesis longfellowi> longfellowi (Girault)
show synonyms of <Sympiesis longiventris> longiventris (Girault)
lucida Storozheva
maculata Erdos
show synonyms of <Sympiesis marilandia> marilandia Girault
show synonyms of <Sympiesis marilandica> marilandica Girault
show synonyms of <Sympiesis mauiensis> mauiensis (Ashmead)
mishi Yefremova and Shroll
show synonyms of <Sympiesis myartsevai> myartsevai Ozdikmen
show synonyms of <Sympiesis noncarinata> noncarinata (Girault)
show synonyms of <Sympiesis notata> notata (Zetterstedt)
show synonyms of <Sympiesis obscura> obscura (Ashmead)
oditidis Kerrich
ornatula Storozheva
show synonyms of <Sympiesis oryzae> oryzae Risbec
show synonyms of <Sympiesis particola> particola (Girault)
show synonyms of <Sympiesis polygoniae> polygoniae (Risbec)
show synonyms of <Sympiesis pulcherrima> pulcherrima (Dodd)
show synonyms of <Sympiesis punctifrons> punctifrons (Thomson)
show synonyms of <Sympiesis pustacola> pustacola Szelenyi
qinghaiensis Liao
show synonyms of <Sympiesis ringoniellae> ringoniellae Kamijo
rugithorax Crawford
show synonyms of <Sympiesis saintpierrei> saintpierrei (Girault)
santelinae Szelenyi
show synonyms of <Sympiesis sergeyi> sergeyi Storozheva
show synonyms of <Sympiesis sericeicornis> sericeicornis (Nees)
show synonyms of <Sympiesis smaragdina> smaragdina Storozheva
solitaria Szelenyi
show synonyms of <Sympiesis spadiceipes> spadiceipes (Girault)
sparsus Yoshimoto
show synonyms of <Sympiesis speciosa> speciosa (Girault and Dodd)
show synonyms of <Sympiesis stigmata> stigmata Girault
show synonyms of <Sympiesis stigmatipennis> stigmatipennis Girault
show synonyms of <Sympiesis striatipes> striatipes (Ashmead)
show synonyms of <Sympiesis superba> superba (Girault)
swezeyi Yoshimoto and Ishii
tebennae Myartseva and Kurashev
show synonyms of <Sympiesis tenuis> tenuis Storozheva
thapsianae Boucek
thyrsisae Gupta, Gawas and Bhambure
show synonyms of <Sympiesis triclada> triclada (Provancher)
trjapitzini Storozheva
tschulensis Myartseva and Kurashev
show synonyms of <Sympiesis turcica> turcica Fahringer
show synonyms of <Sympiesis turkmenica> turkmenica Ozdikmen
show synonyms of <Sympiesis vagans> vagans (Timberlake)
show synonyms of <Sympiesis viridula> viridula (Thomson)
vizri Myartseva and Kurashev
volgensis Yefremova and Shroll
show synonyms of <Sympiesis xanthostoma> xanthostoma (Nees)
yuekseli Doganlar
zygis Myartseva and Kurashev
show species in genus <Sympiesomorpha> Sympiesomorpha
show species in genus <Tachinobia> Tachinobia
show species in genus <Tamarixia> Tamarixia
show species in genus <Tanava> Tanava
show species in genus <Tatiana> Tatiana
show species in genus <Tetrasta> Tetrasta
show species in genus <Tetrastichomphale> Tetrastichomphale
show species in genus <Tetrastichomyia> Tetrastichomyia
show species in genus <Tetrastichus> Tetrastichus
show species in genus <Thripastichus> Thripastichus
show species in genus <Thripobius> Thripobius
show species in genus <Thymus> Thymus
show species in genus <Tooloomius> Tooloomius
show species in genus <Trichospilus> Trichospilus
show species in genus <Trielacher> Trielacher
show species in genus <Trigonopterites> Trigonopterites
show species in genus <Trisecodes> Trisecodes
show species in genus <Trjapitzinichus> Trjapitzinichus
show species in genus <Tropicharis> Tropicharis
show species in genus <Tylomischus> Tylomischus
show species in genus <Uroderostenus> Uroderostenus
show species in genus <Uroentedon> Uroentedon
show species in genus <Wichmannia> Wichmannia
show species in genus <Xanthella> Xanthella
show species in genus <Xenaprostocetus> Xenaprostocetus
show species in genus <Xenopomphale> Xenopomphale
show species in genus <Xiphentedon> Xiphentedon
show species in genus <Zagrammosoma> Zagrammosoma
show species in genus <Zaommomentedon> Zaommomentedon
show species in genus <Zaommomyiella> Zaommomyiella
show species in genus <Zasympiesis> Zasympiesis
show species in genus <Zealachertus> Zealachertus
show species in genus <Zeastichus> Zeastichus
show species in genus <Zeytinus> Zeytinus
show taxa in family <Eupelmidae> Eupelmidae
show taxa in family <Eurytomidae> Eurytomidae
show taxa in family <Khutelchalcididae> Khutelchalcididae
show taxa in family <Leucospidae> Leucospidae
show taxa in family <Megastigmidae> Megastigmidae
show taxa in family <Mymaridae> Mymaridae
show taxa in family <Ormyridae> Ormyridae
show taxa in family <Perilampidae> Perilampidae
show taxa in family <Pteromalidae> Pteromalidae
show taxa in family <Rotoitidae> Rotoitidae
show taxa in family <Signiphoridae> Signiphoridae
show taxa in family <Tanaostigmatidae> Tanaostigmatidae
show taxa in family <Tetracampidae> Tetracampidae
show taxa in family <Torymidae> Torymidae
show taxa in family <Trichogrammatidae> Trichogrammatidae


NB: This list represents valid Chalcidoid taxa. Family- and genus-group synonyms are not included.