Dr Harald Schneider PhD habil.

Harald Schneider
  • Researcher, Pteridophytes
  • Life Sciences department
  • Plants Division
Natural History Museum
Cromwell Road


Employment history

Researcher Band 3, The Natural History Museum, 2006.

Hochschulassistent, Department of Systematic Botany, Albrecht-von-Haller Institute of Plant Sciences, Georg-August University, Goettingen, Germany, 2002-2006.

Postdoctoral Research Associate, Duke University, Durham, NC, USA. 2001.

Postdoctoral Research Associate, University of California, Berkeley, CA, USA. 2000.

Postdoctoral Research Associate, The Field Museum, Chicago, IL, USA. 1999.

Postdoctoral Research Associate, Rijksherbarium, University of Leiden, Netherlands. 1996-1998.

Research Assistant, University of Zurich, Switzerland. 1992-1996.


Habilitation at the Georg-August University Goettingen, Germany (Venia Legendi), 2006.

PhD, Biological Sciences, University of Zurich, Switzerland. 1996.

Diploma, Biological Sciences, University of Kaiserslautern, Germany, 1992.

Professional Roles


Host of Marie-Curie Intraeuropean Fellowship “Mascarene Ferns” awarded to S Hennequin. 2008.

Courant Research Centre Geobiology funded by Excellence Initiative of the Germany Science Foundation and the Ministry of Science, Third Supporting Track (Concepts for the Future). (PI) 2008.

German Science Foundation Grant (HE 3584/2-2). Molecular phylogeny and historical biogeography of Porellales (co-PI). 2008.

German Science Foundation Grant (SCHN 758/4-1). Scaly tree ferns of Madagascar: exploring island radiations in ferns. (PI). 2006.

German Science Foundation Grant (SCHN 758/2-2). Exploring phylogenetic aspects of radiations in derived ferns: using the epiphytic polygrammoid ferns as a model lineages. (PI). 2006.

German Science Foundation Grant (HE 3584/2-1). Molecular phylogeny and historical biogeography of Porellales (co-PI). 2005.

German Science Foundation Grant (SCHN 758/2-1). Exploring phylogenetic aspects of radiations in derived ferns: using the epiphytic polygrammoid ferns as a model lineages. (PI). 2003.

National Science Foundation Grant (DEB 0089909) Collaborative Research: Using phylogeny to investigate the history of morphological evolution in heterosporous ferns. (co-PI). 2001.


Member of the Linnean Society, London.

Marie-Curie Fellowship, European Commission Research Grant “Diversity and phylogeny of selected genera in the brake fern family (Pteridaceae) in Malesia.


Royal Society Conference Travel Awards (2007, 2008, 2010)

Edgar T Wherry Award of the Pteridological Section of the Botanical Society America (2000)

Mellon Systematic training grant (2001)

Robery O Bass Visiting Scientist Award (2001)

Thomas J Dee Fellowship award (1999)

Travel Award XIV International Botanical Conference (1999)

Travel Award, Green Plant Phylogeny Research Co-ordination Group (1999)

Travel Award, G. & A. Claraz Foundation, Zurich, Switzerland.

External collaborations

Diversification of East Asian ferns with emphasize on the Himalaya: Xian-Chun Zhang (Institute of Botany, Chinese Academy of Science, Beijing, PR China).

Phylogeny of selected liverwort families: Jochen Heinrichs (Georg-August University Goettingen, Germany).

Amber fossils and land plant evolution: Alexander Schmidt (Courant Centre Geobiology, Georg-August University Goettingen, Germany).

TEMPO: Vincent Savolainen and Omar Fiz (Imperial College, London, UK).

Phylogeny and taxonomy of Polypodiacae and other fern families: Alan Smith (University of California, USA), Christopher Haufler (University of Kansas, USA), Peter Hovenkamp (Dutch National Herbarium Leiden, Netherlands), Michael Kessler (Institute of Systematic Botany, University of Zurich, Switzerland), Tom Ranker (University of Hawaii, USA), Xian-Chun Zhang (Academy of Sciences, Beijing, China).

Phylogeny of Aspleniacae: Leon Perrie and Pat Brownsey (Musem of New Zealand, Te Papa Tongarewa, Auckland, New Zealand), Wen-Liang Chiou (Taiwan Forest Institute, Taiwan), Klaus Mehltreter (Instituto de Ecologia, Xalapa, Mexico), Michael Kessler (Institute of Systematic Botany, University of Zurich, Switzerland), Xian-Chun Zhang (Institute of Botany, Chinese Academy of Science, Beijing, PR China).

Deep Phylgeny of Ferns: Kathleen Pryer and Eric Schuettpelz (Duke University, USA).

Ecologgy and fern diversity: David Coomes (University of Cambridge, UK), Michael Kessler (University of Zurich, Switzerland)

Cytology and diversification of Mascarene ferns: Mary Gibby, Stuart Lindsay (RBGE) and Sabine Hennequin (Paris)

British Bluebells: Pete Hollingsworth (RBGE) and NHM bluebell group

Evolution of the chloroplast genome of ferns: Paul Wolf (University of Utah USA), Dietmar Quandt (University of Bonn, Germany), Susann Wicke (University of Vienna, Austria), Chun-Neng Wang (National Taiwan University, Taiwan)

Genome evolution of ferns: Andrew Leitch (Queen Mary, University of London, UK), Ilia Leitch (Royal Botanic Gardens, Kew).

External committees

Several PhD committees including University of Reading, Royal Botanic Garden Edinburg, University of Turku, Marie & Pierre Curie University Paris.

Guest lectures and key-note lectures

More than 30 guest lectures in the last five years including lecture at the Chinese Academy of Science, Beijing (China), Sun Yatsen University Guangchou (China), Freie Universität Berlin (Germany), Ludwig-Maximilian University Munich (Germany), Martin-Luther Universität Halle-Wittenberg (Germany), Open University (UK), Purdue University (USA), University of Cambridge (UK), University of Florida (USA), University of Hamburg (Germany), University of Uppsala Sweden), Taiwan Forest Research Institute (Taiwan). Recent Key-lectures at Systematic 2008 at Goettingen (Germany, 2008), National Fern Meeting at Shenzhen (China, 2008), Niche Evolution at Zurich (Switzerland, 2009). 

Editorial boards

Systematic Biology (since 2005)

Taxon (since 2006)

Species, Phylogeny and Evolution (since 2006)

Editor of the “phylogenetic analyses” column of Taxon (since 2007)



General interest: Evolutionary Geobiology of Land Plants with focus on ferns and other seed-free land plants. My research incorporates various research approaches including phylogenetics, population genetics, evolutionary morphology and taxonomy.

Current Projects

Diversification of ferns in East Asia and South East Asia

East Asia and South East Asia include several of fern diversity hotspots.  These research projects are aiming to discover the processes of species assembling and maintenance in this region especially in responds to the rise of the Tibetan plateau and the collision of SE Asia and the Australian craton.  The research project involves collaborations with research groups in China (X-C Zhang, Institute of Botany, Chinese Academy of Science, Beijing, PR China).

Barcoding ferns and other land plants

Barcoding is a promising concept for fern diversity research.  My research groups is actively involved in the development of barcoding regions to obtain DNA based identification of gametophytes and young sporophytes of ferns.

Integrating climate/ distribution modeling, population genetics, and phylogenetics

The reconstruction of many evolutionary processes in the past make explicit statements concerning the distribution of taxa in the past and their evolutionary responds to climatic fluctuations.  This research aims to integrate analytical approaches developed in climate/ distribution modeling, population genetics and phylogenetics. Current studies address the question of climatic niche conservatism in major lineages of ferns. Distribution modeling depends on the access to reliable distribution data as conserved in the collections of the NHM. The project is carried out in collaboration with Nadia Bystriakova.

Adaptive radiation of land plants

Land plants underwent various diversification and extinction events since their successful colonisation of terrestrial habitats. The studies involved aim to discover general processes underlying these events such as the role of key innovations and to discover relationships among these diversification events such as evidence for parallel rapid diversification of several lineages of land plants in the Cretaceous. Sub-projects are focusing on the early diversification of land plants he diversification of land plants in the late Cretaceous, and Tertiary changes in plant diversity. See

Courant Research Centre Geobiology

Diversification of asplenioid ferns and the evolution of climate niche

The genus Asplenium is one of the most species rich genera of plants and the most diverse genus among ferns. Its evolution and taxonomy is poorly understood. The project has two main objectives. First, to uncover the phylogenetic relationships within the genus and the geographical and ecological aspects of the diversification of this highly diverse pantropical and pantemperate fern lineage. Second, to investigate the evolution of climate niches in plant lineages and relationships between diversification and extinction rate to ecological preferences.

Diversification of ferns in biodiversity hotspots and Oceanic islands 

This project is aiming to investigate major process of diversity assembly including neutral versus non-neutral assembling processes by studying the diversity of fern communities in biodiversity hotspots such as Madagascar and Oceanic islands such as Mascarene islands.

Taxonomic relationships of Polypodiaceae and other fern families

 This ongoing collaborative set of projects targets the diversity of the highly species rich and mainly epiphytic fern lineage of Polypodiaceae as well as other lineages of ferns such as scaly tree ferns (Cyatheacae) and Pteridaceae. Within this project the generic classification of these ferns are revised. Previous results include the description of one new genus and the redefinition of several other genera.

Diversification of liverworts through time

This is a collaborative study together with Jochen Heinrichs, Jörg Hentschel and Alexander Schmidt (all Georg-August University Goettingen). The major goals are the reconstruction of the diversification of leafy liverworts, especially Porelales, using DNA sequence variation and the fossil record of these plants.


Research Staff

Sabine Hennequin (Postdoctoral researcher) Marie-Curie Fellowship: Investigating the fern diversity of the Mascarne Islands. Sabine is now Maitre de Conference at the Universite Pierre et Marie Curie, Paris France.


Research Student Supervision


Wang Li (PhD student, Chinese Academy of Science, Beijing, P.R. China; co-supervised by Hong De-Yuan & Zhang Xian-Chun)

Robert Dyer (PhD student, NHM/ Imperial College; co-supervised by Vincent Savolainen)

Chang Yanfen (PhD student, NHM/ Xishuangbanna Tropical Garden, Chinese Academy of Sciences, Kunming, P.R. China; co-supervised by Li Jie & Lu Shu-Gang)

Lu Thi Ngan (PhD student, Vietnam National Museum of Nature, Hanoi, Vietnam)

Yu Ying (PhD student, East China Normal University, Shanghai, China; co-supervised by Zhu Rui-Liang)

Marie-Capucine Lepeigneux (DEA student, University Pierre et Marie Curie, Paris, France; co-supervisred by Sabine Hennequin)

Wita Wardani (MSc Student, University Reading, UK; co-supervised by Alastair Culham)



PhD students

Thomas Janssen (PhD student, Georg-August Univeristät Göttingen; now PostDoc in the SUN-Project at the Senckenberg Research Institute, Frankfurt a.M. Germany)

Hans-Peter Kreier (PhD student, Georg-August Univeristät Göttingen; now PostDoc in the research group of J. Heinrichs, Georg-August Univeristät Göttingen)

Jörg Hentschel (PhD student, Georg-August Universität Göttingen, Germany; now PostDoc at the University Jena)


Workstudents/ Masterstudents/ Diplomastudents/ 

Sally Fletcher (work experience student)

Fabienne Hennequin (work experience student)

Jennifer Huepeden (work experience student, Georg-August Universität Göttingen, Germany)

Rayko Jonas (Diploma student, Georg-August Universität Göttingen, Germany; co-supervised by Michael Kessler)

Jeannine Marquart (work experience student, Georg-August Universität Göttingen, Germany)

Peter Mundt (Staatsexamen, Georg-August Univeristät Göttingen)

Heiko Muth (Diploma student, Georg-August Univeristät Göttingen)

Adriana Navarro-Gomez (work experience student and Lab support)

Elisabeth Otto (Diploma student, Georg-August Univeristät Göttingen. now PhD student at the Freie Universität Berlin, Germany)

Janine Weix (Diploma student, Georg-August Univeristät Göttingen & Fachhochschule Bingen, now PhD student at the University of Bern, Switzerland)


Visitors to the "Schneider Research Team":

James Beck, Duke University Durham, NC, USA.

Nadia Bystriakova, Cambridge University, UK. 

Arjen de Groot, Institute of Environmental Biology, Utrecht University, Utrecht, Netherlands.

Wolf Eiserhardt, University of Hamburg, Germany (SYNTHESYS).

Sabine Hennequin, Pierre & Marie Curie University, Paris, France (SYNTHESYS).

Peter Hovenkamp, National Herbarium Netherlands, Leiden, Netherlands (SYNTHESYS).

Ares Jimenez Soria, Area de Biodiversidad y Conservacion, Departamento de Biologia y Geologia ESCET, Universidad Tey Juan Carlos, Madrid, Spain.

Marcus Lehnert, Staatliches Museum für Naturkunde, Stuttgart, Germany (SYNTHESYS).

Kathleen Pryer, Duke University Durham, NC, USA.

Dietmar Quandt, Ness Institut, Bonn, Germany.

Nick Rowe, Univeristy of Montepellier 2, France.

Thomas Speck, University of Freiburg, Germany.

Susan Sprunt, University of Miami, Ohio, OH, USA.

Hans Stenoien, Norwegian University of Science and Technology. Norway.

Susann Wicke, University of Vienna, Austria and Nees-Institut Bonn, Germany (SYNTHESYS).

Paul Wolf, Utah State University, Logan, UT, USA. 

Jasper Wubs, Institute of Environmental Biology, Utrecht University, Utrecht, Netherlands.

Wang Chun-Neng, Institute of Ecology and Evolutionary Biology, Department of Life Science, National Taiwan University, Taipei, Taiwan.


  • Ansell, S.W., H.K. Stenoien, M. Grundmann, H. Schneider, A. Hemp, ,N. Nauer, S.J. Russell & J.C. Vogel ( 2010 ) Population structure and historical biogeography of European Arabidopsis lyrata Heredidy . 10.1038/hdy.2010.10
  • Grundmann, M., F. Rumsey, S.W. Ansell, S.J. Russell, S.C.Darwin, J.C. Vogel, M. Spencer, J. Squirrel, P. Hollingsworth, S. Ortiz & H. Schneider ( 2010 ) Molecular phylogeny of bluebells, Hyacinthoides (Asparagaceae). Taxon 59 : 68-82. . Taxon
  • Hennequin, S., A. Ebihara, J.-Y. Dubuisson, H. Schneider ( 2010 ) Chromosome number evolution of Hymenophyllum (Hymenophyllaceae), with special reference to the subgenus Hymenophyllum Molecular Phylogenetics and Evolution . 10.1016/i.ynpev.2010.01.001
  • Stricklund-Constable, R., H. Schneider, S.W. Ansell, S.J. Russell & S. Knapp ( 2010 ) Species identity in the Solanum bahamense complex (Solanaceae, Solanum subgenus Leptostemum) Taxon 59 : 209-226 . Taxon
  • Wang, L., X-P. Qi, Q-P. Xiang, J. Heinrichs, H. Schneider, X-C. Zhang ( 2010 ) Phylogeny of the paleotropical fern genus Lepisorus (Polypodiaceae, Polypodiopsida) inferrred from four chloroplast regions Molecular Phylogenetics and Evolution 52 : 211-225 . 10.1016/jympev.2009.08.032
  • Wang, L., Z.-Q. Wu, Q.-P. Xiang, J. Heinrichs, H. Schneider & X.-C. Zhang ( 2010 ) A molecular phylogeny and a revised classification of tribe Lepisoreae (Polypodiaceae) based on an analysis of four plastid DNA regions. Botanical Journal Linnean Society 162 : 28-38. . 10.1111/j.1095-8339.2009.01018.x
  • Heinrichs, J., F. Klugmann, J. Hentschel & H. Schneider ( 2009 ) DNA taxonomy, cryptic speciation and diversification of the Neotropical-African liverwort, Marchesinia brachiata (Lejeuneaceae, Porellales). Molecular Phylogenetics and Evolution 53 : 1112-1121 . 10.1016/j.ympev.2009.05.032
  • CBOL Working Group: Hollingsoworth, P.M., L. L. Forrest, J. L. Spouge, M. Hajibabaei, S. Ratnasingham, M. van der Bank, M. W. Chase, R. S. Cowan, D. L. Erickson, A. J. Fazekas, S. W. Graham, K. E. James, K.-J. Kim, W. J. Kress, H. Schneider, J. van AlphenStahl, S. C. H. Barrett, C. van den Berg, D. Bogarin, K. S. Burgess, K. M. Cameron, M. Carine, J. Chacon, A. Clark, J. J. Clrkson, F. Conrad, D. S. Devey, C. S. Ford, T. A. H. Hederson, M. L. Hollingsworth, B. C. Husband, L. J. Kelley, P. R. Kesanakurti, J. S. Kim, Y.-D. Kim, R. Lahaye, H.-L. Lee, D. G. Long, S. Madrinan, O. Maurin, I. Meusnier, S. G. Newmaser, C.-W. Park, D. M. Percy, G. Petersen, J. E. Richardson, G. A. Salazar, V. Savolainen, O. Seberg, M. J. Wilkinson, D.-K. Y. & D. P. Little

    ( 2009 ) A DNA barcode for land plants. Proceedings of the National Academy of Science USA 106 : 12794-12797 . Proceedings of the National Academy of Science USA | doi:10.1073/pnas.0905845106
  • Dubuisson, J.-Y., H. Schneider & S. Hennequin ( 2009 ) Epiphytism in ferns: diversity and history. Comptes Rendus Biologies 332 : 120-128 . 10.1016/j.crvi.2008.08.018
  • Hentschel, J., M. J. von Konrat, T. Pocs, A. Schaefer-Verwimp, A. J. Shaw, H. Schneider & J. Heinrichs ( 2009 ) Molecular insights into the phylogeny and subgeneric classification of Frullania Raddi (Frullaniaceae, Porellales). Molecular Phylogenetics and Evolution 52 : . 10.1016/j.ympev.2008.12.021
  • Hunt, H.V., S.W. Ansell, S.J. Russell, H. Schneider & J.C. Vogel ( 2009 ) Genetic diversity and phylogeography in two diploid ferns, Asplenium fontanum subsp. fontanum and A. petrachae subsp. bivalens, in the western Mediterranean. Molecular Ecology 18 : 4940-4594 . 10.1111/j.1365-294X.2009.04402.x
  • Otto, E.  M., T. Janssen, H.-P. Kreier & H. Schneider ( 2009 ) New insights into the phylogeny of Pleopeltis and related Neotropical genera (Polypodiaceae, Polypodiopsida). Molecular Phylogenetics and Evolution . 10.1016/j.ympev.2009.05.001
  • Schneider, H., A.R. Smith & K.M. Pryer ( 2009 ) Is morphology really at odds with molecules in estimating fern phylogeny? Systematic Botany 34 : 455-475. . 10.1600/036364409789271209
  • Wang, L., X.-P. Qi, J. Heinrichs, H. Schneider & X-C Zhang ( 2009 ) Phylogeny of the paleotropical fern genus Lepisorus (Polypodiaceae, Polypodiopsida) inferred from four chloroplast DNA regoins. Molecular Phylogenetics and Evolution . 10.1016/j.ympev.2009.08.032
  • Ansell, S. W., M. Grundmann, S. J. Russell, H. Schneider & J. C. Vogel ( 2008 ) Genetic discontinuity, breeding systems, and population history of Arabis alpina (Brassicaceae) in Italy Molecular Ecology 17 : 2245-2257 . 10.1111/j.1365-294X.2008.03739.x
  • James, K., H. Schneider, S. Ansell, M. Evers, L. Robba, G. Uszynski, N. Pedersen, A. Newton, S. Russell, J. Vogel & A. Kilian ( 2008 ) DArT for pan-genomic evolutionary studies of non-model organisms PLOSone 3 : e1682 . 10.1371/journal.pone.0001682
  • Janssen, T., N. Bystriakova, F. Rakotondrainibe, D. Coomes, J.-N. Labat & H. Schneider ( 2008 ) Neoendemism in Madagascan tree ferns results from recent, coincident diversification bursts Evolution 62 : 1876-1889 . 10.1111/j.1558-5646.2008.00408.x
  • Kreier, H.-P., M. Rex, K. Weising. M. Kessler, A. R. Smith & H. Schneider ( 2008 ) Inferring the diversification of the epiphytic fern genus Serpocaulon (Polypodiaceae) in South America using chloroplast sequences and AFLPs Plant Systematics and Evolution 274 : 1-16 . 10.1007/s00606-008-0021-3
  • Kreier, H.-P., X.-R. Zhang, H. Muth, & H. Schneider ( 2008 ) The microsoroid ferns: inferring the relationships of a highly diverse lineage of Paleotropical epiphytic ferns (Polypodiacaee, Polypdiidae) Molecular Phylogenetics and Evolution 48 : 1155-1167 . 10.1016/j.ympev.2008.05.001
  • Salino, A., T. E. Almeida, A. R. Smith, A. Navarro-Gomez, H.-P. Kreier & H. Schneider ( 2008 ) A new Microgramma (Polypodiaceae) from Brazil and recircumscription of the genus based on phylogenetic evidence Systematic Botany 33 : 630-635 . 10.1600/036364408786500208
  • Schneider, H., H.-P. Kreier, T. Janssen & P. Hovenkamp ( 2008 ) Phylogenetic relationships of the fern genus Chrstiopteris shed new light onto the classification and biogeography of drynarioid ferns Botanical Journal Linnean Society 645-656 . 10.1111/j.1095-8339.2008.00825.x
  • Smith, A. R., K. M. Pryer, E. Schuettpelz, P. Korall,, H. Schneider & P. G. Wolf ( 2008 ) Fern Classification. In: Haufler CH, Ranker TA (eds.), In: Biology and Evolution of Ferns and Lycophytes . Cambridge University Press 417-467 .
  • Ansell, S. W., H. Schneider, N. Pederson, M. Grundmann, S. J. Russell & J. C. Vogel ( 2007 ) Molecular evolution of a tandemly repeated trnF pseudogene in the chloroplast genome of Arabidopsis lyrata Journal of Evolutionary Biology 20 : 2400-2411 . 10.1111/j.1420-9101.2007.01397.x
  • Gradstein, R.S., M. Schwertfeger, M. Kessler, V. Wissemann & H. schneider ( 2007 ) The Old Botanical Garden of the University of Göttingen – recent developments and scientific research Museum Aktuell 136 : 11-15 .
  • Heinrichs, J., J. Hentschel, R. Wilson, K. Feldberg & H. schneider ( 2007 ) Evolution of leafy liverworts (Jungermanniidae, Marchantiophyta): estimating divergence times from chloroplast DNA sequences using penalized likelihood with integrated fossil evidence Taxon 56 : 31-44 . Taxon
  • Hentschel, J., J. Paton, G. H. Schneider & J. Heinrichs ( 2007 ) Acceptance of Liochlaena Nees and Solenostema Mitt., the systematic position of Eromonotus Pearson and notes on Jungermannia L. s.l. (Jungermanniidae) based on chloroplast DNA sequence data Plant Systematics and Evolution 268 : 147-157 .
  • Hentschel, J., K. Feldberg, H.-J. Zündorf, F. H. Hellwig,. H. Schneider & J. Heinrichs ( 2007 ) The systematic position of Pachyglossa and Clasmatocolea (Jungermanniopsida: Lophocoleaceae) inferred from nrDNA ITS sequences and morphology Taxon 56 : 1136-1142 . Taxon
  • Hentschel, J., R.-L. Zhu, D. G. Long, P. G. Davison, H. Schneider, R. Gradstein & J. Heinrichs ( 2007 ) Phylogeny of Porella (Porellaceae, Jungermanniopsida) based on nuclear and chloroplast DNA sequences Molecular Phylogenetics and Evolution 45 : 693-705 . 10.1016/j.ympev.2007.05.005
  • Janssen, T., H.-P. Kreier & H. Schneider ( 2007 ) Origin and diversification of African ferns with special emphasis on Polypodiaceae Brittonia 59 : 159-181 . 10.1663/0007-196X(2007)59[159:OADOAF]2.0.CO;2
  • Korall, P., J. S. Metzgar, D. S. Conant, H. Schneider & K. M. Pryer ( 2007 ) Phylogeny of scaly tree ferns (Cyatheaceae) as revealed by five plastid loci American Journal of Botany 94 : 873-886 . American Journal of Botany
  • Kreier, H.-P., A. F. Rojas-Alvarado, A. R. Smith & H. Schneider ( 2007 ) Hyalotrichopteris is indeed a Campyloneurum (Polypodiaceae) American Fern Journal 97 : 127-135 . 10.1640/0002-8444(2007)97[127:HIIACP]2.0.CO;2
  • Metzgar, J., H. Schneider & K. M. Pryer ( 2007 ) Phylogeny and divergence time estimates for the fern genus Azolla (Salviniaceae) International Journal of Plant Sciences 168 : 1045-1053 .
  • Nagalingum, N. S., H. Schneider & K. M. Pryer ( 2007 ) Molecular phylogenetic relationships and morphological evolution in the heterosporous fern genus Marsilea Systematic Botany 32 : 16-25 . 10.1600/036364407780360256
  • Schneider, H. ( 2007 ) Plant morphology as the cornerstone to the integration of fossil sand extant taxa in phylogenetic analyses Species, Phylogeny and Evolution 1 : 65-71 .
  • Schuettplez, E., H. Schneider, L. Huiet, M. D. Windham & K. M. Pryer ( 2007 ) A molecular phylogeny of the fern family Pteridaceae: assessing overall relationships and the affinities of previously unsampled genera Molecular Phylogeny and Evolution 44 : 1172-1185 . 10.1016/j.ympev.2007.04.011
  • Wilson, R., J. Heinrichs, S. R. Gradstein & H. Schneider ( 2007 ) Steady diversification of derived liverworts under Tertiary climatic fluctuations Biology Letters 3 : 556-569 . 10.1098/rsbl.2007.0287
  • Wilson, R., S. R. Gradstein, H. Schneider & J. Heinrichs ( 2007 ) Unravelling the phylogeny of Lejeuneaceae (Jungermanniopsida): evidence for four main lineages Molecular Phylogenetics and Evolution 43 : 270-282 . 10.1016/j.ympev.2006.10.017
  • Grundmann, M., H. Schneider, S. J. Russell & J.C. Vogel ( 2006 ) Phylogenetic relationships of the moss genus Pleurochaete Lindb. (Bryales: Pottiaceae) based on chloroplast and nuclear genomic markers Organisms, Diversity, Evolution 6 : 33-45 . 10.1016/j.ode.2005.04.005
  • Hartmann, F. A., R. Wilson, H. Schneider & J. Heinrichs ( 2006 ) Testing hypotheses on species delimitations and disjunctions in the liverwort Bryopteris (Jungermanniopsida: Lejeuneaceae) International Journal of Plant Sciences 167 : 1205-1214 . 10.1086/508023
  • Heinrichs, J., M. Linder, H. Groth, J. Hentschel, K. Feldberg, C. Renker, J. J. Engel, M. von Konrat, D. G. Long & H. Schneider ( 2006 ) Goodbye or welcome Gondwana – insights into the phylogenetic biogeography of the leafy liverwort Plagiochila with description of Proskaurea, gen. nov. (Plagiochilaceae, Jungermanniales) Plant Systematics and Evolution 258 : 227-250 . 10.1007/s00606-006-0411-3
  • Hentschel, J., R. Wilson, M. Burghardt, H.-J. Zündorf, H. Schneider & J. Heinrichs ( 2006 ) Reinstatement of Lophocoleaceae (Jungermanniopsida) based on chloroplast gene rbcL data: exploring the importance of female involucres for the systematics of Jungermanniales Plant Systematics and Evolution 258 : 211-226 . 10.1007/s00606-006-0408-y
  • Hu, S., D. L. Dilcher, H. Schneider & D. M. Jarzen ( 2006 ) Eusporangiate ferns from the Dakota formation, Minnesota, USA International Journal of Plant Sciences 167 : 579-589 . 10.1086/501235
  • Korall, P., D. S. Conant, H. Schneider, K. Udea, H. Nishida & K. M. Pryer ( 2006 ) On the phylogenetic position of Cystodium: it’s not a tree ferns – it’s a polypod! American Fern Journal 96 : 45-53 . 10.1640/0002-8444(2006)96[45:OTPPOC]2.0.CO;2
  • Korall, P., K. M. Pryer, J. Metzgar, H. Schneider & D. S. Conant ( 2006 ) Tree ferns: monophyletic groups and their relationships as revealed by four protein-coding plastic loci Molecular Phylogenetics and Evolution 39 : 830-845 .
  • Kreier, H.-P. & H. Schneider ( 2006 ) Phylogeny and biogeography of the staghorn fern genus Platycerium (Polypodiaceae, Polypodiidae) American Journal Botany 93 : 217-225 . American Journal Botany
  • Kreier, H.-P. & H. Schneider ( 2006 ) Reinstatement of Loxogramme dictyopteris, based on phylogenetic evidence, for New Zealand endemic fern, Anarthropteris lanceoltata Australian Systematic Botany 19 : 309-314 .
  • Nagalingum, N. S., H. Schneider, M. Sakuj & K. M. Pryer ( 2006 ) Comparative morphology of reproductive structures in heterosporous water ferns and a reevaluation of the sporocarp International Journal of Plant Sciences 167 : 805-815 . 10.1086/503848
  • Schneider, H. ( 2006 ) Aleuritopteris. In: Johns, R. J., .P J. Edwards, T. M. A. Utteridge & H. C. F. Hopkins. (eds.), In: Guide to the alpine and subalpine Flora of Mount Jaya . Royal Botanic Gardens, Kew 43-47 .
  • Schneider, H. & E. Schuettpelz ( 2006 ) Identifing fern gametophytes using DNA sequences Molecular Ecology Notes 6 : 989-991 . 10.1111/j.1471-8286.2006.01424.x
  • Schneider, H., H.-P. Kreier, L. R. Perrie & P. J. Brownsey ( 2006 ) The relationships of Microsorum (Polypodiaceae) species occurring in New Zealand New Zealand Journal of Botany 44 : 121-127 .
  • Schneider, H., H.-P. Kreier, R. Wilson & A. R. Smith ( 2006 ) The Synammia enigma: evidence for a temperate lineage of polygrammoid ferns (Polypodiaceae, Polypodiidae) in southern South America Systematic Botany 31 : 30-40 .
  • Smith, A. R., H.-P. Kreier, C. H. Haufler, T. A. Ranker & H. Schneider ( 2006 ) Serpocaulon (Polypodiaceae), a new genus segregated from Polypodium Taxon 55 : 919-930 .
  • Smith, A. R., K. M. Pryer, E. Schuettpelz, P. Korall, H. Schneider & P. G. Wolf ( 2006 ) A classification for extant ferns Taxon 55 : 705-731 .
  • Heinrichs, J., S. R. Gradstein, R. Wilson & H. Schneider ( 2005 ) Towards a natural classification of liverworts (Marchiantiophyta) based on the chloroplast gene rbcL Cryptogamie Bryology 26 : 131-150 .
  • Janssen, T. & H. Schneider ( 2005 ) Exploring the evolution of humus collecting leaves in drynarioid ferns (Polypodiaceae, Polypodiidae) Plant Systematics and Evolution 252 : 175-197 . 10.1007/s00606-004-0264-6
  • Schneider, H., T. A. Ranker, S. J. Russell, R. Cranfill, J. Geiger, R. Aguraijua, K. R. Wood, M. Grundmann & J. C. Vogel ( 2005 ) Origin and diversification of the Hawaiian fern genus Diellia coincides with the renewal of Hawaiian terrestrial life in the Miocene Proceedings of the Royal Society London, Series B 272 : 455-460 . doi: 10.1098/rspb.2004.2965
  • Groth-Malonek M., J. Heinrichs, H. Schneider & S. R. Gradstein ( 2004 ) Phylogenetic relationships in the Lejeuneaceae (Hepaticae) inferred using ITS sequences of nuclear ribosomal DNA Organisms, Diversity & Evolution 4 : 51-58 .
  • Pryer, K. M., E. Schuettpelz, P. G. Wolf, H. Schneider, A. R. Smith & R. Cranfill ( 2004 ) Phylogeny and evolution of monilophytes: a new perspective on ferns, with a focus on early-diverging leptosporangiate lineages American Journal of Botany 91 : 1582-1598 . American Journal of Botany
  • Pryer, K. M., H. Schneider & S. Magallón ( 2004 ) The radiation of ferns, horsetails and seed plants. In Cracraft, J. & Donoghue, M., (eds.), In: Assembling the tree of life . Oxford University Press : New York , 138-153 .
  • Ranker, T. A., A. R. Smith, B. S. Parris, J. M. O. Geiger, C. H. Haufler, S .C. K. Straub & H. Schneider ( 2004 ) Phylogeny and evolution of grammitid ferns (Grammitidaceae): a case of rampant morphological homoplasy Taxon 53 : 415-428 . Taxon
  • Schneider, H. ( 2004 ) Palmer D.D. Hawai’i’s ferns and fern allies. Plant Systematics and Evolution. Plant Systematics and Evolution 244: 121-130. Book Review. .
  • Schneider, H., A. R. Smith, R. Cranfill, T. J. Hildebrand, C. H. Haufler & T. A. Ranker ( 2004 ) Unraveling the phylogeny of polygrammoid ferns (Polypodiaceae & Grammitidaceae): exploring aspects of the diversification of epiphytic plants Molecular Phylogenetics and Evolution 31 : 1041-1063 . Molecular Phylogenetics and Evolution
  • Schneider, H., E. Schuettpelz, K. M. Pryer, R. Cranfill, S. Magallón & R. Lupia ( 2004 ) Ferns diversified in the shadow of angiosperms Nature 428 : 553-557 . Nature
  • Schneider, H., S. J. Russell, C. J. Cox, F. Bakker, S. Henderson, M. Gibby & J. C. Vogel ( 2004 ) Chloroplast phylogeny of asplenioid ferns based on rbcL and trnL-F spacer sequences (Polypodiidae, Aspleniaceae) and its implications for the biogeography of these ferns Systematic Botany 29 : 260-274 . 10.1600/036364404774195476
  • Schneider, H., T. Janssen, P. Hovenkamp, A. R. Smith, R. Canfill, C. H. Haufler & T. A. Ranker ( 2004 ) Phylogenetic relationships of the enigmatic Malesian fern Thylacopteris (Polypodiaceae, Polypodiidae) International Journal of Plant Sciences 165 : 1077-1087 . 10.1086/423882
  • Dubuisson, J-Y., S. Henequin, F. Rokatondrainibe & H. Schneider ( 2003 ) Ecological diversity and adaptive tendencies in the tropical fern genus Trichomanes L. (Hymenophyllaceae) with special reference to climbing and epiphytic habitats Botanical Journal of the Linnean Society 143 : 41-63 . Botanical Journal of the Linnean Society
  • Gradstein, S. R., E. Reiner-Drehwald & H. Schneider ( 2003 ) A phylogenetic analysis of the genera of Lejeuneaceae (Hepaticae) Botanical Journal of the Linnean Society 143 : 391-410 . Botanical Journal of the Linnean Society
  • Haufler, C. H., W. A. Grammer, E. Hennipman, T. A. Ranker, A. R. Smith & H. Schneider ( 2003 ) Systematics of the Ant-fern genus Lecanopteris: testing phylogenetic hypotheses with DNA sequences. Systematic Botany 28 : 217-227 . Systematic Botany
  • Schneider, H. & Cheksum Supiah Tawan ( 2003 ) Taenitis blechnoides. In de Winter, W. P., Amoroso, V. B., (eds.), In: PROSEA 15 (2) Ferns and fern allies . Backhuys Publishers : Leiden , 188-190 .
  • Schneider, H. & Go Rusea ( 2003 ) Pteris. In de Winter, W. P., Amoroso, V. B., (eds.), In: PROSEA 15 (2) Ferns and fern allies . Backhuys Publishers : Leiden , 166-170 .
  • Pinter, I., F. Bakker, J. Barrett, C. Cox, M. Gibby, S. Henderson, M. Morgan-Richards, F. Rumsey, S. Russell, S. Terwick, H. Schneider & J. C. Vogel ( 2002 ) Phylogenetic and biosystematic relationships in four highly disjunct polyploid complexes in the subgenera Ceterach and Phyllitis (Aspleniaceae) Organisms, Diversity & Evolution 2 : 299-311 . 10.1078/1439-6092-00050
  • Pryer, K. M., H. Schneider, E. A. Zimmer & J. A. Banks ( 2002 ) Deciding among green plants for whole genome studies Trends in Plant Sciences 12 : 550-554 . 10.1016/S1360-1385(02)02375-0
  • Schneider, H ( 2002 ) Nooteboom, H.P. Davalliaceae: a family of Old World (sub-) tropical ferns. Plant Systematics and Evolution 234: 239-241. CD Review. .
  • Schneider, H. & K. M. Pryer ( 2002 ) Structure and function of spores in the aquatic heterosporous fern family Marsileaceae International Journal of Plant Sciences 163 : 485-505 . 10.1086/340736
  • Schneider, H., A. R. Smith, R. Cranfiill, C. H. Haufler, T. A. Ranker & T. Hildebrand ( 2002 ) Gymnogrammitis dareiformis is a polygrammoid fern (Polypodiaceae) – resolving an apparent conflict between morphological and molecular data Systematics and Evolution 234 : 121-136 . 10.1007/s00606-002-0207-z
  • Schneider, H., K. M. Pryer, R. Cranfill, A. R. Smith & P. G. Wolf ( 2002 ) The evolution of vascular plant body plans – a phylogenetic perspective. In Cronk, Q. C. B., Bateman, R., Hawkins, J. A., (eds.), In: Developmental Genetics and Plant Evolution . Taylor & Francis : London , 330-364 .
  • Biogeography, ecology and phylogeny of Pteridaceae (Filicatae) in Malesia - Taenitidoideae and Cheilanthoideae. In ( 2001 ) Biogeography, ecology and phylogeny of Pteridaceae (Filicatae) in Malesia - Taenitidoideae and Cheilanthoideae. In Saw, L. G., Chua, L.S.L., Khoo, K.C., (eds.), In: Taxonomy: The Cornerstone of Biodiversity, Proccedings of the Fourth International Flora Malesia Symposium . Ampang Press : Kuala Lumpur , 211-224 .
  • Pryer, K. M., H. Schneider, A. R. Smith, R. Cranfill, P. G. Wolf, J. S. Hunt & S. D. Sipes ( 2001 ) Horsetails and ferns are a monophyletic group and the closest living relatives to seed plants Nature 409 : 618-621 . 10.1038/35054555
  • Schneider, H. & P. Kenrick ( 2001 ) The first record of lindsaeoid ferns (Lindsaeaceae, Polypodiidae) from the Lower Cretaceous of Wyoming (Aspen Shale, Albian). Review of Palaeobotany and Palynology 115: 33-41 .
  • Wolf, P.G., H. Schneider & T.A. Ranker ( 2001 ) Geographic distributions of homosporous fern taxa: does dispersal obscure evidence of vicariance? Journal of Biogeography 18 : 263-270 . 10.1046/j.1365-2699.2001.00531.x
  • Lupia, R., H. Schneider, G.M. Moeser, K.M. Pryer & P.R. Crane ( 2000 ) Marsileaceae sporocarps and spores from the late Cretaceous of Georgia, U.S.A. International Journal of Plant Sciences 161 : 957-988 .
  • Schneider, H. ( 2000 ) Morphology and anatomy of roots in the filmy fern tribe Trichomaneae H. Schneider (Hymenophyllaceae, Filicatae) and the evolution of rootless taxa Botanical Journal of the Linnean Society 132 : 29-46 . 10.1111/j.1095-8339.2000.tb01853.x
  • Wollenweber, E. & H. Schneider ( 2000 ) Lipophylic exudates of Pteridaceae – chemistry and chemotaxonomy Biochemical Systematics and Ecology 28 : 751-777 . 0.1016/S0305-1978(99)00118-0
  • Schneider, H. ( 1999 ) Yet another fern with storage roots – Cheilanthes bolborrhiza Mickel & Beitel (Pteridaceae: Pteridophyta) from Mexico and El Salvador Fern Gazette 15 : 269-274 .
  • Schneider, H. ( 1996 ) Root anatomy of Aspleniaceae and the implications for systematics of the fern family Fern Gazette 12 : 160-168 .
  • Schneider, H. ( 1996 ) The root anatomy of ferns: a comparative study. In Camus, J. M., Gibby, M. & Johns, R. J., (eds.), In: Pteridology in Perspective . Royal Botanic Gardens, Kew 271-283 .
  • Schneider, H. ( 1995 ) Farne in Ecuador - Anmerkungen zur Biologie ausgewählter Arten. Farnblätter 26/27 : 42-57 .