The Echinoid Directory

• Periproct displaced laterally from the anterior-posterior axis towards ocular I; ocular plate I insert.
• Apical disc small and raised above the corona; with indented sutures and prominent sutural pits which house tubercles.
• Distinct ring of small tubercles along the inner margins of genital plates bounding the periproct.
• Ambulacral plating trigeminate adorally and trigeminate or unigeminate aborally.
• Ambital and adapical interambulacral tubercles perforate and crenulate, oral interambulacral tubercles and all ambulacral tubercles imperforate.
• Single suranal plate incorporated into disc.

Middle Jurassic (Bathonian-Callovian) to Upper Cretaceous (Maastrichtian) of Europe, Jamaica, Saudi Arabia and the Former Soviet Union.

Heterosalenia martini Cotteau, 1861, p. 96, by monotypy.

• H. martini Cotteau, 1861; Late Cenomanian of France.
• H. occidentalis Hawkins, 1923; Early Maastrichtian of Jamaica.
• H. corallina Dames, 1872; Late Oxfordian of Germany.
• H. suatensis Weber, 1934; Late Oxfordian of the Crimea.
• H. brocki Kier, 1972; Bathonian-Callovian of Saudi Arabia.
• H. ornata Kier, 1972; Bathonian-Callovian of Saudi Arabia.
• H. dhrumaensis Kier, 1972; Bathonian-Callovian of Saudi Arabia.

Carinacea, Calycina, Salenioida; stem group Saleniidae.

Presumed paraphyletic by exclusion of Polysalenia and Novasalenia.

Can be separated into two species groups, a Jurassic complex characterized by having ambulacral plating simple (unigeminate) adapically and a Cretaceous group which has trigeminate plating throughout. The Jurassic species should probably be separated into a new genus.

Cotteau, G. 1861 in 1861-1867. Paléontologie Française. Terrain Crétacé, Tome Septième. Échinides.  G. Masson et fils, Paris

Donovan, S.K. 1990. Jamaican Cretaceous Echinoidea. 2. Goniopygus supremus Hawkins, 1924, Heterosalenia occidentalis Hawkins, 1923, and a comment on Trochalosoma chondra ((Arnold & Clark, 1927). Mesozoic Research 2, 205-217.