The Echinoid Directory

Collyrites Desmoulins, 1835, p. 212

[=Smithiaster Barras, 2007, p. 158 ]

Diagnostic Features	Test ovate in outline; no anterior sulcus. Apical disc disjunct with posterior ocular plates widely separated and also a short distance above the periproct. Ocular plates II and IV meeting behind genital plate 2. Genital plates 1 and 4 small. Interambulacral series 1 and 4 separated by single or double irregular row of elongate catenal plates linking anterior and posterior parts of apical disc. Posterior oculars and associated ambulacra separated from periproct by at least one pair of interambulacral plates. Ambulacra all similar; subpetaloid; pore-pairs small and circumflexed; extending to ambitus before reducing in size. Aboral plates relatively low, wide and closely packed. Short phyllodes developed (5-6 pore-pairs enlarged towards peristome); one of each pair of basicoronal ambulacral plates with two pore-pairs. Peristome anterior of centre, small, subcircular; facing downwards. Plastron plating protosternous. Peripoct small, tear-drop-shaped and just supramarginal; not sunken and no subanal indentation. Catinal plates usually present between posterior oculars and start of periproct. Tuberculation fine scattered tubercles set in groundmass of granules.
Distribution	Middle to Upper Jurassic (Bathonian-Oxfordian), Europe, North Africa, India.
Name gender	masculine
Type	Ananchytes elliptica Lamarck, 1816, p. 318. Syntype: Museum National d\'Histoire Naturelle, Paris B49317.
Species Included	C. ellipticus (Lamarck, 1816); Bathonian-Callovian, Middle Jurassic, France [includes C. dorsalis (Agassiz, in Agassiz & Desor, 1847); Callovian, Europe, India]. C. cheveti Vadet, Nicolleau & Pineau 1998; Callovian, France. C. desoriana Cotteau, 1855; Oxfordian, France. C. lorsrteuri? ; Oxfordian, Bologne: not checked. C. pinguis Desor, 1858, C. acuta (Desor, 1842): not checked. C. tuarkyrensis Poretzkaja, 1962: not checked.
Classification and/or Status	Irregularia; Stem group Atelostomata; Collyritidae. Probably paraphyletic.
Remarks	Distinguished from the rather similar Pygomalus and Pygorhytis by having the posterior ambulacra and their oculars separated from the periproct by at least one pair of interambulacral plates. Distinguished from Cyclolampas by the lack of well-developed phyllodes. Cardiopelta and Proholaster are very similar in test architecture but both have a frontal groove. Barras, C. 2006. British Jurassic irregular echinoids. Monograph of the Palaeontographical Society, 1-168, pls 1-14 (publication number 625). Des Moulins, C. 1835-1837. Études sur les Échinides. Actes de la Société Linnéenne de Bordeaux, 7 (1835), 9 (1837), 45-364. Jesionek-Szymanska, W. 1963. Echinides irreguliers du Dogger de Pologne. Actes Palaeontologica Polonica 8(3), 293-396, pls 1-7. Thierry, J. 1985. Evolutionary history of Jurassic Collyritidae (Echinoidea, Disasteroidea) in the Bassin de Paris (France). Proceedings of the Fifth International Echinoderms Conference, Galway 1984. A. A. Balkema, Rotterdam. Vadet, A. Nicolleau, P. & Pineau, J. P. 1998. Echinides du Callovien de la Sarthe et de l\'Orne. II-B: Les echinides irreguliers. Memoires de la Societe Academique du Boulonnais Serie Histoire Naturelle 19, 1-128, pls 1-40.