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Seeking nightshades in South America

38 Posts tagged with the peru tag
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Field work began in earnest today – we headed from the coast up into the mountains, the destination was Cajamarca, by fast road 6 hours away – but we were taking the road less travelled. We took a tiny dirt track up a dry valley to a village called San Benito; our research in the herbarium in Trujillo told us this would be a good place to look for some special endemics. Wild tomatoes are most diverse in the dry western regions of Peru – so I was hoping to see some of the species I have not yet seen in the wild.

 

It took us a while to find the right road – road signs don’t really seem important in Peru, people generally know where they are going I guess! The area was fantastically dry, with rocky slopes and tall columnar cacti. This is the northern part of the Atacama, the desert created by a combination of the cold ocean current called the Humboldt Current coming from Antarctica and the rain shadow of the Andes to the east.

 

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The tall cactus peeking out from behind the hill is called Neoraimondia; Antonio Raimondi was a famous Peruvian botanist of the late 19th century and really began the exploration of the plant diversity of the country


The first plant we saw (well, the first one we were going to collect!) was a genus I have never seen in the field before – Exodeconus. It is an Atacama endemic, and this species is the only one to grow in northern Peru. Tiina collected a couple of other species last year in the southern part of the Peruvian coast.

 

The plants we saw in this valley were tremendously variable in size, from tiny with only a couple of leaves to large and fleshy and extending to a metre or more. It all depends on water, as is usual in a desert. Desert plants are masters at making do.

 

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Exodeconus maritimus growing in a shady  place under a rock – some leaves on other plants were the size of saucers! The flowers are beautiful, bright white with a deep purple centre


The tomatoes soon began to appear, like Exodeconus growing in slightly wetter microhabitats. The first species we saw was Solanum pennellii – the closest relative of the tomatoes proper. It doesn’t have the pointed anther cone of the rest of the group, so was placed in the genus Solanum, rather than Lycopersicon, as the tomatoes used to be known.

 

We now recognise all of the wild and cultivated tomatoes as members of the genus Solanum, based on the molecular studies done by my colleague David Spooner in the early 1990s. His results showed tomatoes are closely related to potatoes; many characteristics of the plant form also support this evolutionary relationship. So we now group the tomatoes as part of the large genus Solanum, reflecting their ancestry and evolution more accurately.

 

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Solanum pennellii has stubby anthers – here being buzzed by a small bee


All Solanum species are buzz pollinated by bees – the anthers open by tiny pores at the tips and female bees grasp them and set up a resonance inside using their flight muscles, pollen squirts out lands on the bee and she carries it to another flower – if you sit by flowering tomatoes long enough anywhere, bees will come and buzz, the sound is quite audible!

 

Tomatoes proper have a long beak on their anther cone, but there are pores inside – the beak is a shared evolutionarily derived character that tells all tomatoes are closely related to one another.

 

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Solanum arcanum – a northern Peru endemic species only recently described by my colleague Iris Peralta (with whom I was recently collecting in Argentina), has the elongate beak typical of wild tomato relatives. This species began to appear a bit further up the valley towards the mountains

 

The small fruits of wild tomatoes are usually green and hairy, but even from them you can tell the species apart. The sepals of Solanum pimpinellifolium – the progenitor of our cultivated tomato – are strongly turned back, while those of Solanum arcanum (below) are always held flat – easy!

 

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Solanum pimpinellifolium

 

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Solanum arcanum

 

We were elated with our success at finding the wild tomatoes, all in flower, but were becoming disappointed about one special species we were seeking – Solanum talarense, an endemic to the dry coastal valleys of northern Peru and rarely collected.

 

We had almost given up, the road was going up the valley into wetter habitats and higher elevations, but then we saw it – possibly the rattiest plant I have ever seen! Eaten by goats, despite its ferocious prickliness, there it was hanging on in rocks by the roadside.

 

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Solanum talarense amongst the rocks, completely eaten by goats (we think, but it certainly had been munched by something!)

 

Jumping out of the truck I felt prickles in my shoes – only to discover that several 2 cm long thorns had gone right the way through the bottom of my boots; it will be interesting to see how this affects them when we get to wetter places! Solanum talarense was most definitely THE plant of the day, totally weird and wonderful – a plant only a Solanum taxonomist could love.

 

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The 'spines' on the stems and leaves of Solanum talarense are technically prickles, outgrowths of the surface – true spines, like those that went through my boot, are bits of stem. Prickliness does not seem to have deterred the animals eating this plant at all!

 

Ascents from the coast to the Andes in Peru are amazing – in a single day you can go from sea level and a dry desert to 4000 metres elevation and dripping wet cloud forest. This time we went over a pass that was only 3500 metres elevation, to descend again into the dryer valley of Cajamarca (via a couple of other passes – the geography is incredibly complex in northern Peru).

 

DSC_7103_resized.jpgWet cloud forest beckons ahead in the mountains

 

In the village of San Benito, in the cloud forest – in the pouring rain – we found our last wild tomato of the day. Solanum habrochaites used to be called Lycopersicon hirsutum, but when the time came to change its name to put it into Solanum, there already was a Solanum hirsutum (a European species) – so we had to think of another species name for it. The name we chose – habrochaites – means softly hairy in Greek; we thought it described the plant exactly.

 

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The fruits of Solanum habrochaites are covered with long hairs each of which has a small sticky gland on top. These glands exude a substance that gives each wild tomato species its particular smell, and gives us that lovely smell of ripe tomatoes fresh from the vine

 

We reached Cajamarca at about 11 pm, a bit later than planned – maybe we spent too much time collecting in the desert, but none of us thought so! This first day collecting was a great success – lots of other wonderful northern Peru endemics and some real surprises and firsts for me (Leptoglossis schwenckioides, Browallia acutiloba and on and on).

 

Now for a day in the herbarium of the University of Cajamarca and a visit to the wonderful Peruvian botanist Isidoro Sánchez Vega, for whom I named a lovely species of Solanum a couple of years ago. I am hoping we find Solanum sanchez-vegae on this trip – maybe when we leave Cajamarca for points south. Can’t wait.

 


Posted on behalf of Sandy Knapp, Museum botanist on field work in Peru.

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Today we spent the day databasing specimens in the local Trujillo herbarium; all herbaria have 1-4 letter acronyms, standard codes so we all know what collection we are talking about – the one for the herbarium in Trujillo is HUT. It is a small collection and has been built up over the last few decades by a series of local botanists – the one I knew the best was Abundio Sagástegui, who sadly died a few years ago.

 

The collection is rich in material from this area of Peru – and has a lot of gems. For example – I described Solanum clivorum in 1992 and have only seen a few specimens. Here at HUT we saw eight new ones, thus expanding our knowledge about this species’ distribution hugely! 

 

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Space is at a premium at HUT – Tiina and my computer tried to share the desk – Tiina eventually won!

 

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Maria used the other computer to database specimens, balancing it all on a stool…..

 

Trujillo is a beautiful city, with old colonial architecture and wonderfully painted buildings. We managed to become part of a parade to celebrate the local team having won the national women’s volleyball championship – brass bands and all.

 

The Plaza de Armas is the heart of any Peruvian city – the one in Trujillo features brightly and freshly painted buildings, now mostly government offices (it is the capital of La Libertad Department) and a fantastic mustard coloured cathedral.

 

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The taxi count in Trujillo’s city centre is phenomenal – we counted about 10 taxis to every private car, and no buses – perhaps the streets are too narrow

 

Trujillo was named after the birthplace in Spain of the conquistador Francisco Pizarro, who led the expedition that defeated the Inca rulers of Peru. It has a long and distinguished history since that founding in 1536, and has been the site of many important events in Peruvian history; it was the first Peruvian city to declare independence from Spain.

 

But Trujillo's history began long before Europeans came onto the scene – it was at the centre of both the Moche and Chimu cultures, pre-Inca coastal peoples who constructed monumental pyramids of adobe (clay bricks) and mud. One thing I love about Peru is that wherever you are, the history stretches far back into extraordinary events and cultures, but those cultures are also alive and evident today.

 

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Many of the houses on the Plaza retain the original (or restored) Spanish colonial balconies made of wood from which women would watch festivities safely hidden from view

 

So after a day’s hard work in HUT, we re-identified about half of the Solanum collection, added new and exciting locality records to our field trip planning and made new friends.

 

Although many of the collections we entered into the database were already represented by duplicates from elsewhere (because plants are big, botanists usually collect several examples and share them with other collections), I still think it is important to have the specimens from small local herbaria like HUT in a main project database – it gives visibility to institutions that otherwise sometimes go un-noticed and lets other botanists know that there is treasure to be found in these small collections!

 

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Sandy, William and Paul outside the door to one of the rooms housing the HUT herbarium

 

Tomorrow collecting begins in earnest – our route has been planned using the information we have derived from the collections we saw today, plus those already in the database, and from the knowledge of where people haven’t been!

 

If all goes well we will go from the deserts of the coast to montane cloud forest near the town of Cajamarca, where we will find another small, but rich herbarium to work in the next day. What will be our most exciting find? It’s bound to be something we don’t expect……

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Heading north…

Posted by Sandy Knapp May 6, 2013

After a last push in the herbarium and a bit of shopping for containers for Erica to rear flies in (obtained by Maria in the Mercado Central in Lima, where you can quite literally buy anything!), we were ready to go…

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Maria Baden and Tiina Sarkinen with the fruits of an amazing shopping expedition to the Lima Central Market – the plastic containers will be used to rear fly larvae we find in fruits of Solanaceae

 

Leaving Lima at 6am (to beat the traffic) we drove through grey, gloomy fog – typical weather on the coast at this time of the year. The fogs come in off the cold Humboldt current – making it cold and damp, but never truly wet; the unique vegetation of the coast lives entirely off the moisture from these seasonal fogs. Several endemic species of Solanum live in the lomas – or fog forests – but we are a bit too early to catch them. The foggy season has only just started and plants are only just beginning to grow.

 

The coast of Peru always fascinates me – it is such a dry desert, yet people live wherever water presents itself. At intervals between desert sections, the Panamerican Highway on which we drove north crosses small rivers coming from the Andes to the east, and there agriculture flourishes.

 

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Sugar cane is a common sight in the river valleys; increasingly it is being grown under irrigation farther into the desert – surely an unsustainable practice. This is, after all, one of the driest deserts in the world!

 

Once we got further north, the skies cleared, the fog burned off and the sun came out – and on we went! When the Pacific came in view we all got out to have a look, even though the area was completely devoid of plants. The water is clear, blue and cold – we saw fishermen in tiny rowboats fishing what looked dangerously close to rocks just off the coast. This part of the coast is famous for its anchovy fishery – the fish are used for fishmeal, but also end up on pizzas worldwide.

 

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Our rental pickup parked near the coast – it really is a desert!

 

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Maria shooting video footage of the fishermen – the sea was an unreal colour of blue

 

One peculiar vegetation formation that occurs on the desert hills is the 'tillandsial' – patches of small bromeliads, plants related to Spanish moss or airplants, that are the only living things growing. Like all the other native plants of this coast, they get their water entirely from the fogs that come in off the sea. Their leaves are covered with scales that help trap the water from the air. In a way, they are like epiphytes, but on the ground!

 

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Tillandsial near the town of Casma – this species is probably Tillandsia purpurea

 

We didn’t stop for lunch until quite late – between towns in river valleys there is nothing at all. When we did stop we had our usual coffee rating - coffee can be wonderful (10) here, or dire (0)– this rated about a 2, pretty awful, but the Inca Kola sugar canister made up for it. Inca Kola is the local soda pop – fluorescent yellow and sticky, it is said to be flavoured with lemon grass; I don’t believe it!

 

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Tiina with the Inca Kola canister (we had to stop her asking to buy it, it is her favourite sort of enamel kitchenware!)

 

We made it to Trujillo and a wonderful colonial hotel; tomorrow we hit the herbarium on the search for more localities for Peruvian endemic Solanaceae. So – I’m sure we will find them, but the big question is – what will the coffee tomorrow morning be like?

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One of the things scientists do today that never happened in the past is to request official permission to conduct scientific research in another country. It might seem a bit bureaucratic and overly pernickety, but the issue of permits for collecting is an important way in which tropical countries rich in biodiversity manage their natural resources – probably the most important of which is biodiversity itself.

 

In Peru, all permissions for collection are managed through the Ministry of Agriculture, and there are clearly laid out rules for how to apply. I already have a permit for collecting Solanaceae – but this year I needed to sort out a permit for new work to be done under the Museum’s science initiatives.

 

The Museum's Natural Resources Initiative has three strands:

 

  • Critical Elements managed by Richard Herrington of Earth Sciences
  • Neglected and Emerging Diseases managed by Tim Littlewood of Life Sciences
  • Crop and Pest Wild Relatives (CPWR), managed by me – we jokingly call it the rocks, pox and crops initiative!

 

Crop and Pest Wild Relatives


Our main idea in the CPWR strand is to use the data from our and other collections to look at the distributions of crop wild relatives and the wild relatives of major crop pests, then use these data to model both plant and insect responses to the changing environment, taking into account the evolutionary relationships of each of the groups, a sort of orthogonal axis.

 

We have chosen to begin with the rich Solanaceae dataset I and collaborators have amassed over many years of databasing specimens in herbaria all over the world and manage through Solanaceae Source – it means the plant layer is done already! We will then begin to digitise (image, database and geolocate) all the Museum’s specimens of three major pest groups – beetles (relatives of the Colorado Potato Beetle, one of the worst pests of potato), leafhoppers or jumping plant lice (devastating pests of all kinds of crops), and fruit flies (big pests of tomato and aubergine). We also will do a new kind of collecting, where entomologists and botanists go in the field together – we will collect all the insects associated with particular Solanaceae species (well, really from any we see), thus compiling data on who lives where and on whom.

 

Hence the need to collect insects on Solanaceae in Peru – the centre of diversity for both wild potatoes and tomatoes. And the necessity of obtaining a legal permit to export the specimens so they can be compared with our collections and identified; I completed all the paperwork last night, and submitted it all at the Ministry today.

 

The importance of doing this now is that we are taking advantage of my current collecting trip to Peru for Tiina and my joint project on endemics, and two Museum entomologists are joining us in the middle of May – Erica McAlister (curator of flies and well known from her flygirl blog!) and Diana Percy (researcher on leafhoppers) will test out our collecting protocols and get the first field data for the initiative. It is exciting, as it feels like things are really starting!

 

Changes in Peru

 

Lima is a funny place – it is big, chaotic and has a very energetic, almost frenetic feel. It is in the dry coastal zone of Peru, so rain never (or very rarely) falls – the only moisture is fog from the sea. Getting to the Ministry involves wild taxi rides through crowded streets – dodging accidents and traffic jams. I lived in Peru in the 1980s, at a difficult time for the country; it was in the grip of the Sendero Luminoso (Shining Path) terrorist threat then.

 

Today Lima is a more open, vibrant place – and things are really happening. Even the huge multistory tower that is part of the Social Security complex next to the Peruvian National Natural History Museum looks like it is due for changes – the sign says 'Soon this tower will be at your service. After 30 years'. This building has stood empty since the early 1980s, towering over the museum gardens. So, let’s see if things really do change!

 

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The Seguro Social tower - ready for a long-delayed makeover!

 

We head to the north on Sunday – passing through the herbaria of Trujillo and Cajamarca to enter data from specimens of endemic species into Solanaceae Source. Then the fieldwork blog will really be about field work at last!

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Tiina Sarkinen, who until late March was working with me on South American Solanum, has now set out on her own with a new job at the Royal Botanic Garden, Edinburgh. We will continue to work together - the world of nightshade research is ever-expanding!

 

She left for our second big Peru trip at the beginning of April, and has just posted her first blog post through Edinburgh's website. The work is funded by National Geographic, so blogs will appear there too. Watch NaturePlus to see the work expand!

 

I go out to join Tiina on the 1st of May, after a brief stop-over to give a couple of lectures about Alfred Russel Wallace in the Amazon in the USA - our objective is the Cordillera of Huascaran and more exciting solanums!

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Making order out of chaos

Posted by Tiina Apr 10, 2012

29th March – Good bye to Freddy our car

Today we had to say goodbye to Freddy, our lovely and most faithful car. We are finally back in Lima!

 

The day passed sorting things out, and clearing and cleaning Freddy. Presses went back to the Museum’s store room, specimens into our preparation area, and so on. Most importantly, Paul re-united with his family – his 2 year old daughter Fabiana was full of smiles! In the evening we all went for a nice cake and ice cream to celebrate our safe homecoming.

 

 

30th March – Digitalising field notes

I have been typing our field notes bit by bit into our database. I started the job whilst we were in Arequipa. Yesterday I used the long drive to type some more in the car. Today I have spent all day doing this, and now as the typing has finished I am focusing on preparing herbarium labels. Our database software has a great interface for designing your labels – but you need to do some technical stuff to get the labels look perfect. It is pretty much like programming language, that is how it feels. Quite nice, very nice balance after all the field work.

 

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Once the label design is correct, the labels need to be proof read. It is good to make sure they look perfect – they will remain with the specimens for years to come!! Sandy sent us acid free paper from UK, so that we can print the labels properly. For long term storage, it is important to use acid free paper. These specimens are going to be five star top quality!

 

 

2nd April – Applying for an export permit

Our labels are ready now! We printed them out, and then cut them neatly.

 

label making (Mobile).JPG

 

I sorted the labels into numeric order and then off we went to the herbarium to put them with the specimens! It felt great, we were very excited seeing the specimens all ready to be mounted and processed into the museum.

 

We had a small party to celebrate us finishing. Lemon pie and passion fruit limonade, yum yum! Here is the crew of the San Marcos herbarium, you can see the beautiful garden just outside:

 

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Celebrations were also due as we handed in our export permit application this morning. We had prepared all the documents, and the museum’s secretary at the San Marcos herbarium had helped us to prepare another document necessary for the application. All that hard work paid off, and now we are just waiting to hear back from the Environmental Ministry.

 

 

3rd-4th April – Back in the herbarium with plants

Time to catch up with herbarium work. With all our experience now from our field trip, it is great to be back in the herbarium identifying specimens, and studying the material with more time in our hands.

 

Lima has two other herbaria in the Universidad Nacional Agraria Lamolina (MOL). We visited one of these herbaria today, the Weberbauer herbarium. The collection is rich in types, as many taxonomists used Weberbauer’s high quality specimens to describe new species. Unfortunately, the top duplicates used for describing these species were deposited in Berlin. These duplicates were destroyed during the Second World War, and now the remaining duplicates, many of which are in the Weberbauer herbarium in Lima, are extremely important. If they exist, they can be used to lectotypify the names for which types have been destroyed.

 

I needed to search through the material to find if some duplicates of Solanum sandianum or Solanum planifurcum might exist. I managed to find three specimens which are types of Solanum names, but none of them were the ones I was after. Good job done though, now these specimens can be scanned and put online for other taxonomists to use.

 

Here we are in the midst of annotating and databasing the material, with Professora Vilcapoma in the background.

 

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5th April – Beautiful Lima

It is officially Semana Santa! The museum is closed today, and we are taking some time off to explore the city.

 

Lima is going through a heat wave. Generally in late March Lima is foggy. The fog enters from the Pacific, and takes over the city. With fog, there is less sun and the atmosphere in the city is cold, humid and dim. This year, however, the fog has not arrived yet despite it being April already. Instead, it is +35 degrees celcius every day, full sun shine.

 

We took a walk on the beach to touch the Pacific. Emilio Perales, a junior lecturer from the forestry department of the Universidad Nacional Agraria Lamolina, joined us on our walk – and got into a water war with Andrew!

 

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Emilio explained to us how the local public transport system works. Lima’s famous micros are mindboggling to most tourists. Busses do not have numbers or routes, but instead, they are colour coded. The bus system is not that official even – it seems bus lines are just born out of necessity, rather than planned by the council. Anybody can become a bus driver as well, there is no licence involved.

 

Here are examples of the bus lines: the blue white line:

 

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These busses go from Avenida Arequipa through the Ovalo in Miraflores to Chorrillos via Barranco. It’s a great bus as it takes the more scenic route through Larcomar along the coast.

 

Another one is the red white bus that comes to Chorillos too, but it takes a different route throught Avenida Tacna.

 

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Then there are the more complex bus lines, like the green bus with black roof, two green stripes, one white stripe, and red at the bottom. Who knows where it goes …

 

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The yellow white bus comes to Chorrillos through Abancay

 

yellow white (Mobile).JPG

 

I wish there would be a guide to these busses, but there isn’t. You just have to learn them as you go. It is quite an anarchistic public transport system. The bus drivers are not licensed either. I suppose you wake up one morning, decide to become a bus driver, hackle for a bus, buy it cheap, and paint it (this is the important part!). The next most important thing you need to become a successful bus line is a fierce assistant. We call these assistants bus pimps. The assistant’s job is to shout the route aloud from the moving bus, by hanging from the open bus door and by making as much noice and hand waving as you possibly can. If you accidentally make an eye contact with the bus pimps, you will quickly find yourself inside a bus on a route to somewhere – they are very keen on getting more customers!

 

Once you’ve got your head around the general system, you are off. The busses drive extremely arrogantly, which means you get to your destination fast.

 

People don’t believe me about the colour code when I try to explain it to them, so I hope the blog will help to de-mystify the system to any confused tourists in Lima.

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24th March – Night in the mountains

 

The night was cold in the car. Time passed slowly. At five thirty, suns first rays become visible, and by six the rays warmed us enough through the glass to tempt us out of Freddy.

 

By 7 am we had set up our field office. We started with our fruits first, as there were lots of them to do:

 

7 am (Mobile).JPG

 

Then we prepared all the specimens we had collected before yesterday’s accident, ready to be dried under our gas fire. Once that was done, we continued with the seeds. Sandy has a clever technique for drying seeds: you squeez fruits onto newspaper, then fold the newspaper and let the seeds dry onto the newspaper naturally in air over a few nights before bagging them into proper envelopes for long term storage.

 

By 9 am we were back at the fruits, there were still few to be done.

 

9 am (Mobile).JPG

 

Then we started to clean the car, and to re-organise our things. Whilst clearing our things, we realised we could ring Paul to ask how things were. We were so excited about this! We took our british mobile, charged it with my laptop which had nearly full charge, and rang Paul. He gave us great news: he had two tires with him, one new and one fixed, and he was on his way to us!

 

By 12 am we had eaten our lunch, and had nothing left to do except play with stones.

 

12 am (Mobile).JPG

 

By 2.40 pm we saw a bus – this was it! Paul was on it!!! Happy re-union, and before too long, Freddy had new shoes!

 

new tyres (Mobile).JPG

 

That night we were happy to be in a hotel, sleeping in nice beds, and eating lovely food in Arequipa. Ahhhh.

 

 

25th March – One man down

Paul has been conquered by bacteria – he has tonsillitis!! Lucky that we are in a big city with great doctors. Paul is now resting and all is well. We are feeling bad that all this drama with the tyres might have caused him to fall ill …

 

 

26th March – Herbarium visit

Arequipa is big – it’s one of the biggest cities in Peru. There is a great university, with a great local herbarium. Paul and I had planned a visit, and despite Paul’s poor health, he demanded to come along to the herbarium today with me.

 

Going to a new herbarium feels like opening a treasure chest: what will the folders contain? Will there be something weird but wondeful, perhaps some potential new species hiding away? Here we are looking:

 

herbarium (Mobile).JPG

 

Indeed we found something curious – a specimen similar of the tiniest of all solanums, Solanum chamaesarachidium, which we had seen earlier in our trip in Puno as well as in Argentina. But it wasn’t quite the same, there was something different – different leaf shape, larger calyces where lobes are more fused, and larger inflorescences with longer pedicels and peduncles. The specimen was without flowers though, leaving more to the imagination!

 

mystery (Mobile).JPG

 

We decided that the next day we would go and see if we could hunt it down – locality: on the way to Canyon de Colca from Chivay!

 

 

27th March – Hunting for the rare one

We were so hopeful yesterday! The trip to Canyon de Colca was a long one, and by the time we got to our destination, we had only little time to explore the area by foot. Temptation took over, and we spent until 5pm walking around the hills, trying to find a minute species of Solanum hiding behind rocks and shrubs.

 

We found other stuff, like my favorite Solanum excisirhombeum, the usual suspect in high elevations:

 

excisirhombeum (Mobile).JPG

 

In the lower elevations, in dry sand dunes, we collected Exodeconus, genus of Solanaceae that is adapted to extremely dry conditions:

 

exodeconus (Mobile).JPG

 

See how the anthers are light blue, just poking out from the flower in the above picture, and more clearly visible in the lower one:

 

exodeconus (2) (Mobile).JPG

 

But we did not find the rare specimen we had seen in the herbarium yesterday. What was most fustrating was that the specimen came with exact latitude and longitude position, and with the help of our GPS, we should have been able to find it! But it turned out that the position given on the label was 5 km from the road, and we did not have time to walk that far. It was difficult turning back…

 

 

28th March – Sea, finally!!

This morning it was time to head home – it has been 27 days traveling through the south of Peru now, and we are all missing our families and friends. It feels good to look back. We have found great things, many which require more work in the herbarium to figure out if they are perhaps something new.

 

We have also done some silly mistakes from which we have learnt a lot. The big tour we did after leaving Sandy in Cusco was not, afterall, all that exciting. The road did not pass great Solanum habitats, and hence, we did not collect as much as we expected. With hindsight, we should have taken another road. But then again, if we would have done that, we would not have found the isolated population of the white flowered Solanum chamaesarachidium near Marcapata!

 

When we saw the sea this afternoon around 3pm, it was a great moment. The Panamerican highway descents from Arequipa slowly to the coast at Camana – what a view. We have made it to over so many mountain passes, it felt that seeing the sea symbolised safe homecoming.

 

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21st March – Neotypifying, and yet again, neotypifying

 

Panic over! We found cash and didn't have to wash any dishes. It was the well known classic "going to the bank" method that saved us . Lucky we had some US dollars to change into Peruvian nuevo soles. Still running strong, fuel in the car, food in the stomachs, and money burning in the pockets .

 

Yesterday we drove all day for nearly 300 km on small mountain roads just to get to Sandia. The reason we so desperately want to collect around here is that there are two species described from around Sandia that need re-typification. This is because the material that was used to describe these species, i.e. their type collections, were destroyed during the Second World War in the Berlin herbarium. To replace the lost types, new types need to be made, and this process is called neotypification.

 

The two types we wanted to recollect were for Solanum planifurcum and Solanum sandianum. Both species were described by Bitter, S. planifurcum from the outskits of Sandia from 2100 – 2500m elevation, and confusingly, S. sandianum from higher up above Cuyocuyo, a town c. 20 km from Sandia, at 3800 m elevation. Some claim these species are synonyms, and represent just extreme variation of a single species. Today we shall see!

 

We started our morning from Sandia near S. planifurcum type locality, and luckily found a population just outside Sandia in 2100 m elevation.

 

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We kept collecting populations until 2500 m, and observed variation along the elevational gradient. Further up, all the way to 3200 m we could observe populations of S. planifurcum. Then gradually, things started to change. At 3400 we found what Bitter would have called S. sandianum, just outside Cuyocuyo, in the local rubbish dump. Not a pretty collection locality, I admit, but there is was.

 

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Solanum sandianum seems to have narrower leaves, which are less hairy and shiny above. Flowers vary in colour, but in general they are dark purple rather than pale lilac as Solanum planifurcum. Calyx shape and size, corolla, stamens and style characters seem to vary less. But are the differences in leaf shape, size and indumentum enough to justify recognising two species? We discussed this with Paul and decided that this is a perfect case where molecular sequence data can help us to decide. If molecular data gives evidence that these taxa are not sister to each other, then we will look for morphological differences that could be used to distinguish them. If, on the other hand, molecular data shows these individuals from different elevations along the road from Sandia to Cuyocuyo to be all mixed within a single clade, I think the case is clear for sunking these names for synonomy!

 

Having done a good days work in collecting along the gradient from Sandia to Cuyocuyo, we were heading back towards Juliaca. As we got closer to the city, we opted to drive all the way to Puno for the night as the city is famous for its beautiful location on the shore of Lake Titicaca.

 

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The roads weren’t great, and again, we arrived to Puno very late at night. We managed to find a hostel without trouble this time. It was time for some relaxation, and winding down after a long drive. Lucky for the boys, there was a footfall game on in the evening, Peru was playing against Chile! Tensions were high, it was 0-1 for Chile for long time, and although Peru managed to score before too long, Chile eventually won. The boys were sad, some tears were shed, but I consoled them by reminding them that there was the beautiful view of Lake Titicaca to see in the morning.

 

 

22nd March – Views over Lake Titicaca

 

I always thought as a child that Lake Titicaca was a hot place with sandy beaches and tropical fruits all over. Just to clarify, this was not the fault of the Finnish education system at all. I just had manage to form this image in my head that Lake Titicaca was a place for sunbathing.

 

The truth is very different. Lake Titicaca is the highest elevation lake in the world, and it is not that warm.

 

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Again we had a long drive ahead of us, we had to make some miles to make it across the Andes for the final time. This time we were crossing the Cordillera Occidental to the coastal deserts near Moquegua. It’s a long 7 hour drive from Puno to Moquegua, and this is without any collecting or taking side roads. Of course we couldn’t avoid the temptation to take few sideroads, but just enough to collect few Solanum fragile specimens near Puno.

 

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Solanum fragile is the panda of the Solanum world – it’s flowers are cute as buttons! The calyx lobes are shy but showy, just a little bit recurved as you can see in the picture, and then the stigma! See how long it is, it’s excerted more than the length of the stamens! The species grows high up around 4000 m elevation, in rock crevases – despite this it has all the elegance of a high society lady with light blue petals, large corollas, and the showy appearance!

 

And of course there was Salpichroa hiding amongst the rocks as well! Salpichroa glandulosa is distinct amongst the genus having very densely hairy leaves.

 

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We dissected a flower in the field, and discovered that the nectary disk at the base of the corolla tube is orange!

 

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The slopes on the western side of the Andes near Moquegua are extremely dry and sandy. At 4400 m elevation there was sand everywhere. We kept looking for our sand loving friend Solanum chamaesarachidium, but couldn’t find it. I doubt it occurs this far west, the only populations known from Peru are in Puno.

 

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The western slopes lower down are home for the Regmandra and the tomato clade of Solanum. We collected Solanum peruvianum along the road at around 2500 m elevation, a wild species related to tomato.

 

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Today finding a hotel was easy – we hailed one from the car!!!! We got to Moquegua late, and we were tired. On a tight street just near the main Plaza of the city, whilst waiting for a traffic jam to clear, we manage to find a hotel on our left with their carrage door just a meter from our car, and that was it! We hailed the owner to open the doors before causing too much of a chaos on the street, and checked ourselves in. Sometimes it is easy, sometimes not…

 

 

23rd March – Flowers of the dry hills

 

Having slept peacefully, we headed out early to hunt for a collection I had seen in the herbarium in Lima. It was a specimen near Torata near Moquegua, that resembled Solanum arequipense, a species that has remained mysterious since its original description by Bitter. As for some other species of Bitter, the type of Solanum arequipense was destroyed in the Berlin fire during the Second World War. Without the type, the species has remained elusive and despite the name having been used in various floristic accounts, nobody really knows what the species is really like and what entity the name really refers to.

 

But there it was, just next to Quebrada Torata. This taxon is very similar to Solanum aloysiifolium, species that is found in northern Argentina and Bolivia. My colleague Gloria (see Argentina blog) will find these photos very exciting! Despite the similarity, this species is definetely something different. The leaves vary from entire to serrate, stamens are c. 3 m long, style clearly excerted, with a very capitate stigma.

 

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The calyx lobes are larger than in Solanum aloysiifolium, and most importantly, the fruits remain green when mature. The fruits are speckled with these white small dots, which I first thought to be nothing special. But every fruit, in every individual we have seen, seems to have them.


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We also collected another wild tomato species, Solanum chilense. It grows lower down in elevation.

 

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Another new species for the day was Solanum corymbosum, a member of the Solanum section Parasolanum group. These species are related to Solanum section Solanum. They all have small flowers with tiny anthers, but with large stigmas. Solanum corymbosum occurs in low elevation dry habitats, whilst the others occur at slightly higher elevation. Solanum corymbosum has cute red fruits that resemble mini-tomatoes.

 

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In the extremely dry habitats we passed near Omate we collected Exodeconus

 

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and Nolana, both genera of Solanaceae specialised in desert habitats.

 

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The day was getting late, and we were still in the middle of the desert, slowly ascending to the mountains on our way to Arequipa. It was getting dark, and we were aware that we would be arriving late to Arequipa that night. Except that suddenly we realised we were not going to arrive to Arequipa at all: we had a punctured front left tire! We also had a slow puncture in our back left tyre, and although this tyre looked still OK, air was coming out more and more rapidly.

 

So punctures in both of our left hand side tyres, what could we do? We took our spare out, thinking that changing the flat tyre from the front could get us as far as Arequipa, three hours away. Andrew and Paul took the spare down from its hiding place, and started changing the tyre. In less than 30 minutes, they were putting bolts on and tightening the spare – we were nearly ready to go again! Except that the spare turned out to be flat as well…

 

There was nothing else left to do except to take the spare and walk to the next village to get pumping. I stayed in the car, guarding our poor Freddy as we call our handsome 4by4. The spot were we had had to stop was a dangerous one – there was not adequate space to easily get pass our Freddy if busses or trucks would turn up. I was equiped with a powerful head torch, and hazards triangle which I put on the road.

 

By the time boys headed to the village with the spare tyre, it was pitch black. There I was, finally having an opportunity to breath, enjoy an evening by myself and watch the starts on the clear Andean sky. Beautiful! This is how it looked before going totally dark. I stopped taking photos in the dark as star skies never turn up nice but trust me it was beautiful.

 

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Temperatures rocket down during nights in the Andes, so I stayed inside the car, keeping myself warm, waiting for any passersby. Nothing came for nearly an hour, but then I finally saw lights in the night sky, coming from behind the nearby curve. It was a truck!!! I took my headtorch, shone to indicate our poor positioning in the middle of two curves on narrow part of the road, and hoped that they would slow down before passing Freddy which was missing its both left handside tyres.

 

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The truck barely slowed down. It took a milly second to observe the situation and the narrow space before taking its decision to go for it. The truck passed well, which was good news. This meant buses might be able to pass as well, and we knew to expect a bus soon.

 

Boys returned with bad news: the spare tyre was not only flat but with a massive hole in it.

 

So with three flat tyres and two OK ones, we had no option than to split our team. Paul with his peruvian fluency offered to head to Arequipa with two tires, so that he could return by next morning with repairs. Luckily a car passed by to give Paul a lift. Andrew and I stayed with the car. How would our night in Freddy go – and would the busses get through? We were nervous, but the stars were giving us comfort.

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Back to the "real" world?

Posted by Sandy Knapp Mar 23, 2012

I left Tiina, Andy and Paul in Cusco and began my slow re-entry into the world outside plant collecting – culture shock for sure! Flying into Lima the world looked very different – big cargo ships, anchovy fishing fleets, dry desert – something I hadn’t seen for what seemed like years, but really was only a couple of weeks.

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I tried really hard not to be envious of their journey on the Interoceanica – failed of course, but I will hear about it eventually! They are sure to find great things, can’t wait to see them.

I spent a day and a half in Lima, working in the herbarium again trying to sort out a few mysteries, catching up with friends and generally getting things set up for when the others return. Some of the great old friends from previous times in Peru were in Lima (Blanca Leon and Ken Young from the University of Texas) – we all had a great Sunday walking along the seafront near their apartment in Miraflores.

Although I have now come back to the Museum and am getting to grips with the “real” world, the field trip work does not end yet. All those lovely solanums we collected need to have their data typed up into the database so the labels can be printed out. In order to export our part of the specimens from Peru we must provide complete data labels, too often people come and collect, leave the plants but never send the labels, making the collections a burden for the staff of our sister institution in Lima. Field work needs to be collaborative from start to finish, and the finish is long after one leaves what is considered the “field”!

The Museum hasn’t changed utterly since I left, so picking up the threads of what I was doing before is pretty straightforward. The best bit is that I feel a new excitement for what I am doing in Solanum taxonomy, a new appreciation for the collections we have and the ones we have just made, and have come back altogether rejuvenated – full of new ideas and plans. Seeing plants in their native habitat, doing what they just do, is without doubt once of the most important ways to increase understanding of the diversity and scope of nature. The collections we hold at the Museum are important, of course, but it is the combination of knowledge from the collections and field that really makes for good science - an integrative whole that is more than the sum of its parts.

This is definitely the best job in the world!! Keep tuned in for Tiina's posts from Peru, and once she is back here in London, we will keep you up to date with what we do with the wonderful nightshades we have collected in South America.

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Goodbye western side!

Posted by Sandy Knapp Mar 18, 2012

from Tiina:

Today was a day of goodbyes. First we had to say goodbye to Sandy who is flew to Lima to catch her flight back to London. She has to teach in Spain in a week’s time, so it’s time to get things ready for the next trip. We have had great weeks together collecting Solanum and talking Solanaceae research with our colleagues in Peru and Argentina. Now the challenge is to continue our field work successfully with just three of us left!

 

We drove to Cusco to drop Sandy off at the airport. Cusco is a large city in a small valley, and by now the city has spread to the surrounding hills. As we navigated through the old and curvy streets of Cusco to get to the airport, I took a photo of the beautiful scenery over Cusco taken from the northern hills.

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We continued our day after sad goodbyes to collect in the surrounding hills of Cusco. It was wonderful collecting in such a historic place – we were basically collecting around old ruins! The whole area of Cusco surroundings is full of ruins, where ever you look. Each stone seems to have a carving or human made shape to it. One large rock that I went around turned out to have stairs carved into it! Here is a great sight of the caves in the ruins of Qenqo with Andrew and Paul.

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Amongst these wonderful areas we found a new Salpichroa species we haven’t seen on this trip yet, Salpichroa gayi. Unlike other species of Salpichroa, it has very unusual yellow-purple flowers. The corolla lobes curve very unusually at their tips, which we only spotted once we looked at our close-up photos in the evening!

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Another species we collected near Cusco was Solanum probolospermum, the usual suspect. This species is very common all through the department of Cusco west of the Eastern Cordillera. It’s a climbing species that scrambles on top of shrubs and roadside vegetation. It has attractive large purple flowers, and usually softly pubescent leaves. During our collections we have observed that the species varies enormously in habit and leaf shape, as well as corolla colour, shape and size. You might ask if there is nothing it doesn’t vary in… We are asking the same question by now. Despite all this trouble the species is giving us, here is a nice shot of the individuals we found growing in sunny hillside in Cusco. Sandy will be happy to see Paul is learning how to use our wonderful camera increasingly well!

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After a nice stroll around Cusco, it was time for second goodbyes. This time we are saying goodbye to the western side of the Andes. We are moving on to our next grand tour – a triangle route to the city of Puno through Puerto Inambari. This trip will take us through the Eastern Cordillera via the highway Interoceanica that goes through Peru all the way to Brazil. The road crosses the Andes at a pass at 4750 m elevation – can’t wait to see the views! After the high pass the road descents to the Amazon basin to 500 m elevation or so. This route will allow us to collect in the humid eastern slopes of the Andes where many interesting species of solanums live. We will take the Interoceanica highway all the way to Puerto Inambari which is way into the Amazon. You can see the pencil pointing to the city on the map above. Here we will turn south-east, where we turn south east towards Puno. The road from Puerto Inambari to Puno will take us over the Andes once again back to the western side.

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So goodbyes it was to the western side! We started off on the Interoceanica at four in the afternoon, with km0 saying 4700 km to São Paulo. It’s very tempting to think we could drive to São   Paulo for a lovely dinner, sushi perhaps as they have famously good sushi bars there, but perhaps we have to leave that for another day L. No need to be sad for too long though, there will be great plants to see J!

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We were aiming to stay the first night of our tour in a city called Ocongate, which is roughly three hours into the highway. The area we passed is full of small Andean villages, where most people speak Quechua, language very different from Spanish or English. Some of the local villages we passed had wonderful names – but we just didn’t know how to pronounce them! To give an example, here is a picture of a road sign to one of the villages:

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In case you are still trying to cough the name out, here is how it goes: cc is pronounced with a dry throat sound of “kha”. As there happen to be two double cc in the village name, it becomes a little problematic. Definitely not a word to pronounce whilst eating your dinner! So once you know how to pronounce double c, then it all goes smoothly, or does it?! We are still awake and trying!

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from Sandy:

For my last day of collecting with Team Solanum we decided to go down another of the roads out of the Sacred Valley up over the mountains and to the Amazon slope. This one made a loop, so we thought it would be a good idea! Well, it was, but it made for a VERY long day. First we ascended up a shale scree slope on a road (yes a road) to 4578m elevation over the Abra de Amparaes….

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We then descended through a wonderful glacial valley, classically shaped as a U – it was full of tiny communities with herds of llamas and alpacas. These iconic Andean animals are related to camels and are hardy enough to withstand the altitude and temperatures – although they are set out to graze every day and brought back to stone corrals every night by the local people. You can see the camel-like faces….

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Our first exciting solanum of the day was the enigmatic Solanum “Cusco-branched” – a plant sort of like Solanum probolospermum of the Abra de Malaga, but with dense branched hairs all over its stems and leaves. Is it the same, is it different? We needed to really collect it intensely over the range of elevations to be sure… it seems that this branched hair thing is on the Amazon side (E slopes), with Solanum probolospermum (with simple hairs) on the western slopes. We collected this plant as often as we could over the elevational gradient to be able to see if this idea holds once we get back and compare our collections to those made by others earlier and in different places.

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Down and down the valley we went, encountering more and more solanums as we got lower. We also saw lovely waterfalls and fantastic rock faces…..  this is truly a spectacular place. Driving along a one-lane road cut into the rock wall can be heart-stopping of course, but we are getting used to that!

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We encountered the junction to begin our ascent up the other branch of the road at about 2500m elevation – and just a few hundred metres up that road, a real prize revealed itself – Solanum sinuatiexcisum, the relative of Solanum fiebrigii we had collected in Argentina. This was a huge herb more than 2 m tall though – quite a different beast.

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By this time we were getting a bit nervous about climbing over the pass again (different road though this time) in the dark, so kept our collecting to a minimum…. this of course meant stopping a lot, and getting excited over all the new things we were seeing. One mystery was a small tree with shiny leaves and green fruits that I think might be a new species – I will need to check in the herbarium in Lima once I am back to be sure though, I think I have been calling this Solanum maturecalvans for ages, but it is really different seen alive – this is why field work is so important…. For now it is called Solanum “not-maturecalvans”!

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In a small village just before Lares we saw the beautiful solanaceous species Brugmansia sanguinea close up and personal – this is a magic plant, you can just tell can’t you by the lovely flowers!

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On and up we drove, racing to get onto the paved road before dark – we saw the locals bringing in their llamas, single file along a ridge…

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We made it back to Pisac just as the entire two was closing down at 8pm – a good final meal, a hot shower (the best we have had in Peru yet, in the Hotel Pisac Ayllu) and plant dryer organising finished the day. I will be really sad to leave the team here for the next set of adventures – they are hoping to take a route I have long wanted to go on, but life elsewhere is calling. I wonder what they will find, and I wonder if I will be able to resurface into the NHM again?

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From Tiina:

Today we started early again – destination was Paucartambo and beyond. Paucartambo is on the western side of the Andes west of Cusco, and beyond Paucartambo in the Amazon lowlands is Pilcopata. We wanted to collect on the montane forests again on the western side (Solanum heaven!), but in order to do this we needed to drive over a pass to cross the mountains to the other side.

 

The slope to the east from Pisac to Paucartambo is not as steep as in Abra de Malaga. The pass this time is ONLY 4000m high (everything is relative!).

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The road from the pass down to Paucartambo descents slowly, and past Paucartambo the forest is still relatively dry. Paucartambo itself is a lovely village with a really organised looking market – labelled as to types of produce – here is the “tuber section” – where they sell potatoes and other Andean tuber crops – Sandy went in and had a long conversation with a woman about her potatoes – so many different types!

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It only gets really humid once you get to the point where the road forks to Tres Cruces, where the Manu National Park begins. We got as far as the forking point and had to turn back in order to get home in time. We did find Solanum “morel-malaga” again – a sign that we had reached the humid eastern slopes again.  This individual of “morel-malaga” was a giant: the stem was 2.5 cm in diameter at the base. Again it was growing in a landslide site in very disturbed rocky soil. Everything was the same as in the previous collection we had made in Abra de Malaga, but just bigger!

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Manu  National Park is wonderful. It’s massive, stretching from the high elevation moist pre-puna to the lowland Amazonian rain forest. We couldn’t collect in Manu despite how wonderful it looked – collecting in Peruvian national parks requires a special collecting permit, which we don’t have, so instead of collecting we chatted to the park rangers that were at the main reception were we stopped to turn around. They recognised Sandy by her name, as she has collected along the Paucartambo – Pilcopata road before. What a temptation to be that close to the park and not to be able to go in and collect! Maybe we’ll get to see it one day… we promised the rangers we would be back…..

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After the excitement of our collections along the road to Quillabamba, we began today with a very prosaic collection  - in the garden of our hotel! They had a species of Tiina’s Morelloid clade growing right in their window (one reason we chose this particular hostal) – Solanum polytrichostylum. It was also growing along the stream, so it is defintely native here. It is great that people (not just us!) like solanums enough to cultivate them for ornament – this one is called “suyttu ccaya” in Quechua.

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We packed all our stuff (including the plants that had arrived via the encomienda from Andahuaylas) and set off up the Sacred Valley, as the valley of the Río Urubamba is called. The valley is the site of many Inca and pre-Inca ruins and settlements – many of these are famous, but many more are not excavated and are just part of everyday life. We decided to look for one of Tiina’s mysteries up a small dirt road out of the town of Urubamba; the collection locality was a bit vague and was from the 1960s so we didn’t really know what we would find.

 

Well, we didn’t find the mystery, but did find one of my favourite species, Solanum maturecalvans, at the end of a logging road. Its waxy white flowers seem not quite real, and the crinkled leaf base gave rise to the name of one of its synonyms – Solanum crotalobasis – the rattlesnake tail base! 

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Synonymy is one of those things that seems like a mistake – naming the same thing twice, but is really good evidence of how science really works. In the case of Solanum maturecalvans the German botanist Georg Bitter named two plants he thought were different Solanum maturecalvans and Solanum crotalobasis – they were from different places, looked a bit different, and he really didn’t have many specimens to compare in the early part of the 20th century.  Now I come along, some decades later, and decide that those two type specimens belong to the same species – therefore one (the later one described) becomes a synonym. Not a mistake, just a re-interpretation of the evidence, along with decades more collecting and specimens to look at.

 

In the same bit of forest we found Solanum maturecalvans we also saw a lovely Jaltomata with blood red nectar – how cool is that!

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Coming out of the mountains down through the town we saw Streptosolen jamesonii growing by the side of the road – it is really native to northern Peru and Ecuador, but we collected it anyway. It is a very interesting plant with a twisted corolla – essentially it has an upside down flower.

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We then headed up the Urubamba valley to the town of Pisac, the site of another massive Inca fortress.

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Here the excavations and restorations are still on-going – it was amazing to see the terracing being restored using ancient techniques, down to mixing the mud used as mortar by foot power!

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We got caught in a torrential rainshower, but afterwards explored the hills behind the town and found the wonderful Salpichroa weberbaueri – named for Augusto Weberbauer, the author of the wonderful book The Vegetable World of the Peruvian Andes, who had collected the type specimen.See an earlier post from Lima for more about him.....

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A bit of culture, a bit of botany – what a great day!

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from Tiina:

Staying at an Inca ruin has been a great experience, and today we made it even more special by heading over the clouds to the Amazonian side of the Andes.

 

Early in the morning we got to the pick-up, ready packed with lunch to drive along the road from Ollantaytambo towards Quillabamba. The road crosses the eastern cordillera at c. 4400 m elevation at Abra de Malaga, and the road then turns down towards the Amazon basin. The Amazon basin side is much more humid than the inter-Andean slopes we have seen thus far, but Paul, Andy and I had never seen these humid montane forests before so we were very keen!

 

Heading off on the road from Ollantaytambo towards the pass we were looking out for Solanum sumacaspi. This is a species of the Geminata clade of Solanum, a tree with remarkably glabrous leaves. Solanum sumacaspi only occurs in the Urumbamba valley, and only few collections of it are known. Luckily we did find it in flower AND fruit, which was fantastic. Sandy has described the species but never seen it in the wild, so this was a moment to cherish!

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Close to Solanum sumacaspi we also found Solanum probolospermum, one of my morelloid solanums. This species is nearly like Solanum pallidum, except that Solanum probolospermum has only simple unicellular hairs, whilst Solanum pallidum has branched hairs. We are yet to find out what the exact limits of these species are – more on that later when we get to see Solanum pallidum itself in Puno hopefully!

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We found more and more Solanum probolospermum when we went up the road. It was great to observe the large variation present in the species in terms of growth form. In some populations, we also observed some fantastic mutations – flowers with six corolla lobes, fasciated corollas, and super-numerous inflorescences. It was all a bit too much for Paul, who started doubting our taxonomic expertise when we said it was the same species. Observing natural variation can be mind boggling at times!

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Once we crossed the pass and started going down the eastern slope of the Andes, the air became more humid and clouds surrounded us. The humidity from the Amazon basin hits the Andean mountains on the eastern slopes, which creates moist montane forest habitats as well as special pre-montane rain forest habitats lower down. This time did not have time to go as low as the pre-montane forests unfortunately. Just as well, as many solanums are found growing in the higher elevation montane forest habitats. First, we found a fantastic species of Morelloid solanums, which we have named as “morel-malaga” for now. This species shows very unusual characters for Morelloids: it has enlarged pedicel apices, calyx with reduced calyx tube and tiny lobes, and most unusually, oval-shaped fruits. We observed this species growing along the eastern side from 3600 to 3450 m elevation, at which point we had to turn around to get home in time. As usual for solanums, this species shows great variation in habit and size, but it clearly likes disturbed microhabitats such as landslides. On our return to herbarium in Lima, we’ll keep our eyes open for this species, as some of the field characters such as pedicel size and fruit shape are not obvious on dried herbarium specimens.

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Second find of the day was Salpichroa didieranum. Generally Salpichroa flowers are c. 1-2 cm long, but this species has long yellow tubular flowers up to 12 cm long! Paul was well pleased, this genus is his favourite!

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Best thing of the day was a bunch of mad cyclists we saw. They passed us when we were collecting our final “morel-malaga” populations in the humid montane forest. It was raining at this point, the moisture was engulfing us in a thick fog, and we could barely see more than 30 meters ahead. First we saw one mad cyclist passing by. He was ecstatic with joy, rolling down the slope, and before we knew it, he had disappeared in the clouds. Second cyclist followed, this time he greeted us with joyful “hello” with a hint of Irish accent. At this point we thought these were just eccentric tourists. But then followed a whole crowed of them – cyclists coming down the hill, some silent some screaming for joy rolling down the hill in rain! What a bunch of mad but happy people, I wish I could do a similar downhill cycling route one day! I bet they just kept rolling down until they got to the Amazon lowlands!

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We decided to have a day off from driving madly across the Peruvian countryside and to explore the amazing ruins of Ollantaytambo. This is a small town with a huge ruin attached that is on the banks of the Río Urubamba.

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It was here that the Inca Manco Capac defeated the Spanish army, only to have them come back in more force later and over-run the fortress – he then retreated to Machu Pichu and beyond….  Today trains run from Cusco to Ollantaytambo to Machu Pichu several times daily (last time I was here it was once a day, starting at the crack of dawn! – how times change).

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Part of the site of Ollantaytambo is a temple to the sun (or so the guidebooks say) hewn of gigantic stones that were transported from across the river and high in the mountains. This temple was never finished, and one can see gigantic stones left along the trail from the quarry across the Urubamba. Like all Inca stonework, you can’t put a piece of paper between these stones – what is amazing about Ollantaytambo is the sheer size of these blocks (see Paul for scale in the picture). How did they do it?

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We climbed high on trails around the entire site, and managed to see some really amazing scenery, take some silly pictures of all of us, and see some interesting plants.

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To collect, we went across the river on an old Inca bridge (recontructed of course) and walked amongst terraced fields of maize and other crops.

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We of course found some solanums – the msot exciting of which was what we called Solanum “pseudoexcisirhombeum” – like our friend from the high puna, but a bit different.  For a start it has smaller flowers….

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Hot off the presses (several days later) – on downloading the original description from the Biodiversity Heritage Library (BHL), we find that it matches this Urubamba plant better than the puna plants from Ayacucho – so we will need to figure out what those are now! Being able to look at the literature in the field is truly amazing…

 

Landslides are a fact of life in Peru (as you will have seen from earlier posts!) and the Ollantaytambo area is no exception – this river of mud came down the mountain last October or so (we were told be a resident) at night with a huge whoosh. It hadn’t even been raining apparently, a piece of mountain just fell off. Wow.

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Ollantaytambo of course is more than just ruins, it is also a lovely village, and prides itself on retaining its Inca heritage. It also has some lovely Spanish touches as well, like these bulls on the rooftop for good luck, along with a bottle of champagne for celebration!

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Having a day off from driving was good for all of us – we have recharged our batteries and are ready to go again…  what will tomorrow bring? More solanums we are sure!

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