Dr Paul Eggleton

Dr Paul Eggleton

Merit Researcher, Entomology

Department: Life Sciences
Division: LS Insects
Contact: email

work: 02079425723
mobile: 07913019727


Termite ecology and systematics


I am interested in the ecology and systematics of soil and litter macrofauna, particularly termites, ants, earthworms and beetles. My research has mostly been in forest ecosystems, tropical and temperate, but I am becoming increasingly interested in drier habitats, particularly southern African savannahs. My work is on pattern and process, and is both descriptive and experimental. I am head of the Soil Biodiversity Group within the NHM.



PhD, Imperial College, United Kingdom, 1985 - 1989

BSc, University of Bristol, United Kingdom, 1982 - 1985

Employment history


Merit Researcher, the Natural History Museum, London, Life Sciences, United Kingdom, 2005 - ongoing


Ewers RM, Boyle MJW, Gleave RA, Plowman NS, Benedick S, Bernard H, Bishop TR, Bakhtiar EY, Chey VK, Chung AYC, Davies RG, Edwards DP, Eggleton P, Fayle TM, Hardwick SR, Homathevi R, Kitching RL, Khoo MS, Luke SH, March JJ, Nilus R, Pfeifer M, Rao SV, Sharp AC, Snaddon JL, Stork NE, Struebig MJ, Wearn OR, Yusah KM, Turner EC (2015) Logging cuts the functional importance of invertebrates in tropical rainforest. Nature Communications, 6 : 6836 - 6836. doi: 10.1038/ncomms7836

Fayle TM, Scholtz O, Dumbrell AJ, Russell S, Segar ST, Eggleton P (2015) Detection of Mitochondrial COII DNA Sequences in Ant Guts as a Method for Assessing Termite Predation by Ants. PLOS ONE, 10 (4)doi: 10.1371/journal.pone.0122533

Fayle TM, Eggleton P, Manica A, Yusah KM, Foster WA (2015) Experimentally testing and assessing the predictive power of species assembly rules for tropical canopy ants. Ecology Letters, 18 (3): 254 - 262. doi: 10.1111/ele.12403

Dahlsjö CAL, Parr CL, Malhi Y, Meir P, Rahman H, Eggleton P (2015) Density-body mass relationships: Inconsistent intercontinental patterns among termite feeding-groups. Acta Oecologica, 63 : 16 - 21. doi: 10.1016/j.actao.2015.01.003

Davies AB, Eggleton P, van Rensburg BJ, Parr CL (2015) Seasonal activity patterns of African savanna termites vary across a rainfall gradient. Insectes Sociaux, 62 (2): 157 - 165. doi: 10.1007/s00040-014-0386-y

Dahlsjö CAL, Parr CL, Malhi Y, Meir P, Eggleton P (2015) Describing termite assemblage structure in a Peruvian lowland tropical rain forest: a comparison of two alternative methods. Insectes Sociaux, 62 (2): 141 - 150. doi: 10.1007/s00040-014-0385-z

Dahlsjö CAL, Parr CL, Malhi Y, Meir P, Chevarria OVC, Eggleton P (2014) Termites promote soil carbon and nitrogen depletion: Results from an in situ macrofauna exclusion experiment, Peru. Soil Biology and Biochemistry, 77 : 109 - 111. doi: 10.1016/j.soilbio.2014.05.033

Luke SH, Fayle TM, Eggleton P, Turner EC, Davies RG (2014) Functional structure of ant and termite assemblages in old growth forest, logged forest and oil palm plantation in Malaysian Borneo. Biodiversity and Conservation, 23 (11): 2817 - 2832. doi: 10.1007/s10531-014-0750-2

Djernæs M, Klass K-D, Eggleton P (2015) Identifying possible sister groups of Cryptocercidae+Isoptera: A combined molecular and morphological phylogeny of Dictyoptera. Molecular Phylogenetics and Evolution, 84 : 284 - 303. doi: 10.1016/j.ympev.2014.08.019

Dahlsjö CAL, Parr CL, Malhi Y, Rahman H, Meir P, Jones DT, Eggleton P (2014) First comparison of quantitative estimates of termite biomass and abundance reveals strong intercontinental differences. Journal of Tropical Ecology, 30 (2): 143 - 152. doi: 10.1017/S0266467413000898

Carpenter D, Sherlock E, Sandhu R, Eggleton P (2013) Differences in nest structure influence the importance of Formica rufa group (Hymenoptera: Formicidae) wood ant nests as refugia for earthworms. Journal of Natural History, 47 : 2305 - 2309. doi: 10.1080/00222933.2012.763063

Beccaloni G, Eggleton P (2013) Order Blattodea. Zootaxa, 3703 (1): 46 - 48.

Davies AB, EGGLETON P, Rensburg BJ, Parr CL (2013) Assessing the Relative Efficiency of Termite Sampling Methods along a Rainfall Gradient in African Savannas. Biotropica, 45 (4): 474 - 479. doi: 10.1111/btp.12030

Davies AB, Rensburg BJ, EGGLETON P, Parr CL (2013) Interactive Effects of Fire, Rainfall, and Litter Quality on Decomposition in Savannas: Frequent Fire Leads to Contrasting Effects. Ecosystems, 16 (5): 866 - 880. doi: 10.1007/s10021-013-9657-0

CARPENTER D, HAMMOND PM, SHERLOCK E, LIDGETT A, LEIGH K, EGGLETON P (2012) Biodiversity of soil macrofauna in the New Forest: a benchmark study across a national park landscape. Biodiversity and Conservation, 21 (13): 3385 - 3410. doi: 10.1007/s10531-012-0369-0

CARPENTER D, SHERLOCK E, JONES DT, CHIMINOIDES J, WRITER T, Neilson R, Boag B, Keith AM, EGGLETON P (2012) Mapping of earthworm distribution for the British Isles and Eire highlights the under-recording of an ecologically important group. Biodiversity and Conservation, 21 (2): 475 - 485. doi: 10.1007/s10531-011-0194-x

Bone J, Archer M, Barraclough D, EGGLETON P, Flight D, Head M, JONES DT, Scheib C, Voulvoulis N (2012) Public Participation in Soil Surveys: Lessons from a Pilot Study in England. Environmental Science & Technology, 46 (7)doi: 10.1021/es203880p

Bone J, Barraclough D, EGGLETON P, Head M, JONES DT, Voulvoulis N (2012) Prioritising soil quality assessment through the screening of sites: the use of publicly collected data. Land Degradation and Development, 25 (3)doi: 10.1002/ldr.2138

Carpenter D, Hammond PM, Sherlock E, Lidgett A, Leigh K, Eggleton P (2012) Biodiversity of soil macrofauna in the New Forest: a benchmark study across a national park landscape. Biodiversity and Conservation, : 1 - 26.

Davies AB, EGGLETON P, van Rensburg BJ, Parr CL (2012) The pyrodiversity-biodiversity hypothesis: a test with savanna termite assemblages. Journal of Applied Ecology, 49 (2): 422 - 430. doi: 10.1111/j.1365-2664.2012.02107.x

Fayle TM, Edwards DP, Turner EC, Dumbrell AJ, EGGLETON P, Foster WA (2012) Public goods, public services and by-product mutualism in an ant-fern symbiosis. Oikos, 121 (8): 1279 - 1286. doi: 10.1111/j.1600-0706.2011.20062.x

ROSSER N, EGGLETON P (2012) Can higher taxa be used as a surrogate for species-level data in biodiversity surveys of litter/soil insects?. Journal of Insect Conservation, 16 (1): 87 - 92. doi: 10.1007/s10841-011-9395-6

SNADDON JL, Turner EC, FAYLE TM, Khen CV, EGGLETON P, Foster WA (2012) Biodiversity hanging by a thread: the importance of fungal litter-trapping systems in tropical rainforests. Biology Letters, 8 (3)doi: 10.1098/rsbl.2011.1115

Beccaloni GW, Eggleton P (2011) Higher Classification of Blattodea. Zootaxa, 198

BECCALONI GW, EGGLETON P (2011) Order Blattodea Brunner von Wattenwyl, 1882. In: Zhang, Z.-Q. (ed.). Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa, 3148 : 199 - 200.

Bone J, Head M, JONES DT, Barraclough D, Archer M, Scheib C, Flight D, EGGLETON P, Voulvoulis N (2011) From chemical risk assessment to environmental quality management: the challenge for soil protection. Environmental Science & Technology, 45 : 104 - 110.

EGGLETON P (2011) An introduction to termites: biology, taxonomy and functional morphology.. , Bignell, DE, Roisin, Y & Lo, N , Biology of Termites: A Modern Synthesis., pp. 1 - 26. Springer, Dordrecht.

Foster WA, Snaddon JL, Turner EC, FAYLE TM, COCKERILL TD, Ellwood MDF, BROAD GR, Chung AYC, EGGLETON P, Khen CV, Yusah KM (2011) Establishing the evidence base for maintaining biodiversity and ecosystem function in the oil palm landscapes of South East Asia. Philosophical Transactions of the Royal Society B-Biological Sciences, 366 (1582): 3277 - 3291. doi: 10.1098/rstb.2011.0041

JONES DT, EGGLETON P (2011) Global biogeography of termites: a compilation of sources. , Bignell, DE, Roisin, Y & Lo, N , Biology of Termites: A Modern Synthesis., pp. 477 - 498. Springer, Dordrecht.

Lo N, EGGLETON P (-1) Termite phylogenetics and co-cladogenesis with symbionts. , Bignell, DE, Roisin, Y & Lo, N , Biology of Termites: A Modern Synthesis., pp. 27 - 50. Springer, Dordrecht,.

Palin OF, EGGLETON P, Malhi Y, Girardin CAJ, Rozas-Dávila , Parr CL (2011) Termite diversity along an Amazon-Andes elevation gradient, Peru. Biotropica, 43 (1): 100 - 107. doi: 10.1111/j.1744-7429.2010.00650.x

Six DL, Poulsen M, Hansen AK, Wingfield MJ, Roux J, EGGLETON P, Slippers B, Paine TD (2011) Anthropogenic effects on interaction outcomes: examples from insect-microbial symbioses in forest and savanna ecosystems. Symbiosis, 53 (3): 101 - 121. doi: 10.1007/s13199-011-0119-1

Snaddon JL, Turner EC, Fayle TM, Khen CV, Eggleton P, Foster WA (2011) Biodiversity hanging by a thread: the importance of fungal litter-trapping systems in tropical rainforests. Biology letters,

FAYLE TM, Turner EC, Snaddon JL, Chey VK, Chung AYC, EGGLETON P, Foster WA (2010) Oil palm expansion into rain forest greatly reduces ant biodiversity in canopy, epiphytes and leaf-litter. Basic and Applied Ecology, 11 (4): 337 - 345. doi: 10.1016/j.baae.2009.12.009

Krell FK, Ross E, GIUSTI A, Krell-Westerwalbesloh S, EGGLETON P (2010) Comparison of leaf litter and litter plus upper soil extractions in assessing forest floor macroinvetebrate fauna. Entomologist’s Monthly Magazine, 146 : 71 - 82.

Nobre T, Eggleton P, Aanen DK (-1) Supplemental data Vertical transmission as the key to the colonisation of Madagascar by fungus-growing termites?. null,

EGGLETON P, INWARD K, SMITH J, JONES DT, SHERLOCK E (2009) A six year study of earthworm (Lumbricidae) populations in pasture woodland in southern England shows their responses to soil temperature and soil moisture. Soil Biology and Biochemistry, 41 (9): 1857 - 1865. doi: 10.1016/j.soilbio.2009.06.007

Monaghan MT, Wild R, Elliot M, FUJISAWA T, Balke M, INWARD DJG, LEES DC, Ranaivosolo R, EGGLETON P, BARROCLOUGH TG, VOGLER AP (2009) Accelerated species inventory on Madagascar using coalescent-based models of species delineation. Systematic Biology, 58 (3): 298 - 311. doi: 10.1093/sysbio/syp027

Cornwell WK, Cornelissen JHC, Allison SD, Bauhus J, EGGLETON P, Preston CM, Scarff F, Weedon JT, Wirth C, Zann AE (2009) Plant traits and wood fates across the globe: rotted, burned, or consumed?. Global Change Biology, 15 (10): 2431 - 2449. doi: 10.1111/j.1365-2486.2009.01916.x

FAYLE TM, Chung AYC, Dumbrell AJ, EGGLETON P, Foster WA (2009) The effect of rain forest canopy architecture on the distribution of epiphytic ferns (Asplenium spp.) in Sabah, Malaysia.. Biotropica, 41 (6): 676 - 681. doi: 10.1111/j.1744-7429.2009.00523.x

Monaghan MT, Wild R, Elliot M, Fujisawa T, Balke M, Inward DJG, Lees DC, Ranaivosolo R, Eggleton P, Barraclough TG, others (2009) Accelerated species inventory on Madagascar using coalescent-based models of species delineation. Systematic biology,

Nobre T, Eggleton P, Aanen DK (2009) Vertical transmission as the key to the colonization of Madagascar by fungus-growing termites?. Proceedings of the Royal Society B: Biological Sciences,

Nobre T, EGGLETON P, Aanen DK (2009) Vertical transmission as the key to the colonization of Madagascar by fungus-growing termites?. Proceedings of the Royal Society of London, Series B, 277 (1680)doi: 10.1098/rspb.2009.1373

RAHEEM DC, NAGGS F, CHIMONIDES PD, Preece RC, EGGLETON P (2009) Fragmentation and pre-existing species turnover determine land-snail assemblages of tropical rain forest. Journal of Biogeography, 36 : 1923 - 1938.

Smith J, Potts SG, Woodcock BA, Eggleton P (2009) The impact of two arable field margin management schemes on litter decomposition. applied soil ecology, 41 : 90 - 97.

SCHOLTZ OI, MACLEOD N, EGGLETON P (2008) Termite soldier defence strategies: a reassessment of Prestwich's classification and an examination of the evolution of defence morphology using extended eigenshape analyses of head morphology. Zoological Journal of the Linnean Society, 153 (4): 631 - 650. doi: 10.1111/j.1096-3642.2008.00396.x

Carpenter D, Hodson ME, Eggleton P, Kirk C (2008) The role of earthworm communities in soil mineral weathering: a field experiment. Mineralogical Magazine, 72 : 33 - 36.

EGGLETON P (2008) Methods for sampling termites. Insect Sampling in Forest Ecosystems, : 221 - 221.

RAHEEM DC, NAGGS F, Preece RC, Mapatuna Y, Kariyawasam L, Eggleton P (2008) Structure and conservation of Sri Lankan land-snail assemblages in fragmented lowland rainforest and village home gardens. Journal of Applied Ecology, 45 : 1019 - 1028.

SMITH J, Potts S, EGGLETON P (2008) Evaluating the efficiency of sampling methods in assessing soil macrofauna communities in arable systems. European Journal of Soil Biology, 44 : 271 - 276.

SMITH J, Potts S, EGGLETON P (2008) The value of sown grass margins for enhancing soil macrofaunal biodiversity in arable systems. Agriculture, Ecosystems and Environment, 127 : 119 - 125.

SMITH J, Potts SG, Woodcock BA, EGGLETON P (2008) Can arable field margins be managed to enhance their biodiversity, conservation and functional value for soil macrofauna?. Journal of Applied Ecology, 45

INWARD DJG, VOGLER AP, EGGLETON P (2007) A comprehensive phylogenetic analysis of termites (Isoptera) illuminated key aspects of their evolutionary biology. Molecular Phylogenetics and Evolution, 44 (3): 953 - 967. doi: 10.1016/j.ympev.2007.05.014

Vanbergen AJ, Watt AD, Mitchell R, Truscott AM, Palmer SCF, Ivits E, EGGLETON P, Jones TH, Sousa JP (2007) Scale-specific correlations between habitat heterogeneity and soil fauna diversity along a landscape structure gradient. Oecologia, 153 (3): 713 - 725. doi: 10.1007/s00442-007-0766-3

Inward D, Beccaloni G, Eggleton P (2007) Death of an order: a comprehensive molecular phylogenetic study confirms that termites are eusocial cockroaches. Biology Letters, 3 (3): 331 - 335. doi: 10.1098/rsbl.2007.0102

Carpenter D, Hodson ME, EGGLETON P, Kirk C (2007) Earthworm induced mineral weathering: preliminary results. European Journal of Soil Biology, 43

EGGLETON P, BECCALONI G, Inward D (2007) Invited reply: Responses to Lo et al. Biology Letters, 3 (5)doi: 10.1098/rsbl.2007.0367

Eggleton P (2006) 16 The and Gut Habitat: Its Evolution Termite. Intestinal Microorganisms of Termites and Other Invertebrates, 6 : 373 - 373.

Eggleton PP (2006) spatio-temporal variability and diversity of water column prokaryotic communities in the eastern north atlantic. FEMS Microbiology Ecology,

Eggleton P, Inward D, Jones D (2006) One more novelty: tropical termite diversity came out of Africa. null,

Fayle T, Foster W, Eggleton P, Chung A, Ellwood M, Turner E (2006) Islands in the sky: the ant fauna of epiphytic Bird’s nest ferns (Asplenium nidus complex) in lowland rain forest, Sabah, Malaysia. null,

JONES DT, Davis RG, EGGLETON P (2006) Sampling termites in forest habitats: a reply to Roisin and Leponce. Austral Ecology, 31 : 429 - 431.

Nobre T, Nunes L, EGGLETON P, Bignell DE (2006) Distribution and genetic variation of Reticulitermes (Isoptera: Rhinotermitidae) in Portugal. Heredity, 96 : 403 - 409.

Smith J, Chapman A, Eggleton P (2006) Baseline biodiversity surveys of the soil macrofauna of London’s green spaces. Urban Ecosystems, 9 : 337 - 349.

EGGLETON P, Vanbergen AJ, JONES DT, Lambert MC, Rockett C, HAMMOND PM, BECCALONI J, Marriott D, Ross E, Giusti A (2005) Assemblages of soil macrofauna across a Scottish land use intensification gradient: influences of habitat quality, heterogeneity and area. Journal of Applied Ecology, 42 (6): 1153 - 1164. doi: 10.1111/j.1365-2664.2005.01090.x

Duur AK, EGGLETON P (2005) Fungus-growing termites originated in African rain forests. Current Biology, 15 : 851 - 855.

Eggleton P (2006) The termite gut habitat: its evolution and co-evolution. , , Intestinal microorganisms of termites and other invertebrates., pp. 373 - 404. Springer Berlin Heidelberg.

JONES DT, Verkert RHJ, EGGLETON P (2005) Methods for sampling termites. , Leather S, Insect Sampling in Forest Ecosystems., pp. 221 - 253. Blackwell. Oxford.

KRELL F, Chung AYC, DE BOISE E, EGGLETON P, GIUSTI A, INWARD K, KRELL-WESTERWALBESLOH S (2005) Quantitative extraction of macro-invertebrates from temperate and tropical leaf-litter and soil: efficiency and time-dependent taxonomic biases of the Winkler extraction.. Pedobiologia, 49 (2): 175 - 186. doi: 10.1016/j.pedobi.2004.10.005

Primanda A, Eggleton P, MacLeod N (2005) Worker mandible shape and feeding groups in termites. Seminar Nasional MIPA2005, FMIPA Universitas Indonesia Depok, : 24 - 26.

DONOVAN SE, Purdy KJ, Kane MD, EGGLETON P (2004) Comparison of Euryarchaea strains in the guts and food-soil of the soil-feeding termite Cubitermes fungifaber across different soil-types. Applied and Environmental Microbiology, 70 : 3884 - 3892.

Davies RG, Hernández LM, Eggleton P, Didham RK, Fagan LL, Winchester NN (2003) Environmental and spatial influences upon species composition of a termite assemblage across neotropical forest islands. Journal of Tropical Ecology, 19 : 509 - 524.

DAVIES RG, EGGLETON P, JONES DT, GATHORNE-HARDY FJ, HERNÁNDEZ LM (2003) Evolution of termite functional diversity: analysis and synthesis of local ecological and regional influences on local species richness. Journal of Biogeography, 30 : 847 - 877.

DAVIES RG, HERNÁNDEZ L, EGGLETON P, Didham RK, Fagan L, Winchester N (2003) Assemblage composition responses of termites to fragmentation at a land-bridge island site in French Guiana. Journal of Tropical Ecology, 19 : 509 - 524.

Eggleton P, Davies RG (2003) Isoptera, termites. The natural history of Madagascar (eds Goodman SM, Benstead JP), : 654 - 660.

EGGLETON P, DAVIES RG (2003) Termites (Isoptera). , Goodman SM, Benstead JP, The Natural History of Madagascar., pp. 654 - 660. Chicago University Press. Chicago.

Jones DT, Susilo FX, Bignell DE, Hardiwinoto S, Gillison AN, Eggleton P (2003) Termite assemblage collapse along a land-use intensification gradient in lowland central Sumatra, Indonesia. Journal of Applied Ecology, 40 : 380 - 391.

Krell F-T, Krell-Westerwalbesloh S, Weiß I, Eggleton P, Linsenmair KE (2003) Spatial separation of Afrotropical dung beetle guilds: a trade-off between competitive superiority and energetic constraints (Coleoptera: Scarabaeidae). Ecography, 26 : 210 - 222.

Donovan SE, Eggleton P, Martin A (2002) Species composition of termites of the Nyika plateau forests, northern Malawi, over an altitudinal gradient. African Journal of Ecology, 40 (4): 379 - 385. doi: 10.1046/j.1365-2028.2002.00397.x

Aanen DK, Eggleton P, Rouland-Lefevre C, Guldberg-Frøslev T, Rosendahl S, Boomsma JJ (2002) The evolution of fungus-growing termites and their mutualistic fungal symbionts. Proceedings of the National Academy of Sciences, 99 : 14887 - 14892.

Chung AYC, Chey VK, Speight MR, Eggleton P, Hammond PM, others (2002) A survey on defoliation and phytophagous insects in four habitat types in Sabah, Malaysia.. Journal of Tropical Forest Science, 14 : 116 - 130.

Eggleton P, Bignell DE, Hauser S, Dibog L, Norgrove L, Madong B (2002) Termite diversity across an anthropogenic disturbance gradient in the humid forest zone of West Africa. Agriculture, ecosystems & environment, 90 : 189 - 202.

Eggleton P, Davies RG, Connetable S, Bignell DE, Rouland C (2002) The termites of the Mayombe Forest Reserve, Congo (Brazzaville): transect sampling reveals an extremely high diversity of ground-nesting soil feeders. Journal of Natural History, 36 : 1239 - 1246.

GATHORNE-HARDY F, Davies RG, Eggleton P, Jones DT, others (2002) Quaternary rainforest refugia in south-east Asia: using termites (Isoptera) as indicators. Biological Journal of the Linnean Society, 75 : 453 - 466.

Brauman A, Doré J, Eggleton P, Bignell D, Breznak JA, Kane MD (2001) Molecular phylogenetic profiling of prokaryotic communities in guts of termites with different feeding habits. FEMS Microbiology Ecology, 35 : 27 - 36.

Chung AYC, Chey VK, Eggleton P, Hammond PM, Speight MR (2001) Variation in beetle (Coleoptera) diversity at different heights of tree canopy in a native forest and forest plantation in Sabah, Malaysia. Journal of Tropical Forest Science, 13 : 369 - 385.

Donovan SE, Eggleton P, Bignell DE (2001) Gut content analysis and a new feeding group classification of termites. Ecological Entomology, 26 : 356 - 366.

Donovan SE, Eggleton P, Dubbin WE, Batchelder M, Dibog L (2001) The effect of a soil-feeding termite, Cubitermes fungifaber, (Isoptera: Termitidae) on soil properties: termites may be an important source of soil microhabitat heterogeneity in tropical forests. Pedobiologia, 45 : 1 - 11.

Eggleton P (2001) Termites and trees: a review of recent advances in termite phylogenetics. Insectes Sociaux, 48 : 187 - 193.

Eggleton P, Tayasu I (2001) Feeding groups, lifetypes and the global ecology of termites. Ecological Research, 16 : 941 - 960.

GATHORNE-HARDY F, Eggleton P, others (2001) The effects of altitude and rainfall on the composition of the termites (Isoptera) of the Leuser Ecosystem (Sumatra, Indonesia). Journal of Tropical Ecology, 17 : 379 - 393.

Han Q, Kambhampati S, Eggleton P (2001) Molecular phylogeny of termites based on mitochondrial DNA sequence of COII and ND5. null,

Han Q, Kambhampati S, Eggleton P (2001) Phylogenetic analysis and mandible evolution of the Termitidae (Isoptera). null,

Bignell D, Eggleton P (2000) Termites in Ecosystems. , , DE Bignell - Termites: evolution, sociality, symbioses, ecology, 2000.. Kluwer Academic Publishers.

Chung AYC, Eggleton P, Speight MR, Hammond PM, Chey VK (2000) The diversity of beetle assemblages in different habitat types in Sabah, Malaysia. Bulletin of entomological research, 90 : 475 - 496.

Chung AYC, Hammond PM, Eggleton P, Speight MR, Chey VK, others (2000) A general survey of the Staphylinidae (Insecta: Coleoptera) assemblage in Sabah, Malaysia.. Malayan Nature Journal, 54 : 355 - 367.

Donovan SE, Jones DT, Sands WA, Eggleton P (2000) Morphological phylogenetics of termites (Isoptera). Biological Journal of the Linnean Society, 70 : 467 - 513.

Eggleton P (2000) Global patterns of termite diversity. , , Termites: evolution, sociality, symbioses, ecology., pp. 25 - 51. Springer.

Gathorne-Hardy FJ, Collins NM, Buxton RD, Eggleton P, others (2000) A faunistic review of the termites (Insecta: Isoptera) of Sulawesi, including an updated checklist of the species.. Malayan Nature Journal, 54 : 347 - 353.

Jones DT, Eggleton P (2000) Sampling termite assemblages in tropical forests: testing a rapid biodiversity assessment protocol. Journal of Applied Ecology, 37 : 191 - 203.

Kambhampati S, Eggleton P (2000) Taxonomy and phylogeny of termites. , , Termites: evolution, sociality, symbioses, ecology., pp. 1 - 23. Springer.

Davies RG, Eggleton P, Dibog L, Lawton JH, Bignell DE, Brauman A, Hartmann C, Nunes L, Holt J, Rouland C (1999) Successional response of a tropical forest termite assemblage to experimental habitat perturbation. Journal of Applied Ecology, 36 : 946 - 962.

Dibog L, Eggleton P, Norgrove L, Bignell DE, Hauser S (1999) Impacts of canopy cover on soil termite assemblages in an agrisilvicultural system in southern Cameroon. Bulletin of Entomological Research, 89 : 125 - 132.

Eggleton P (1999) Termite species description rates and the state of termite taxonomy. Insectes sociaux, 46 : 1 - 5.

Eggleton P, Davies R (1999) checklist of termites (Isoptera) from Kaieteur National Park, Guyana. Proceedings of the Entomological Society of Washington,

Eggleton P, Homathevi R, Jones DT, MacDonald JA, Jeeva D, Bignell DE, Davies RG, Maryati M (1999) Termite assemblages, forest disturbance and greenhouse gas fluxes in Sabah, East Malaysia. Philosophical Transactions of the Royal Society B: Biological Sciences, 354 : 1791 - 1802.

Jeeva D, Bignell DE, Eggleton P, Maryati M (1999) Respiratory gas exchanges of termites from the Sabah (Borneo) assemblage. Physiological entomology, 24 : 11 - 17.

MacDonald JA, Jeeva D, Eggleton P, Davies R, Bignell DE, Fowler D, Lawton J, Maryati M (1999) The effect of termite biomass and anthropogenic disturbance on the CH4 budgets of tropical forests in Cameroon and Borneo. Global Change Biology, 5 : 869 - 879.

Dibog L, Eggleton P, Forzi F (1998) Seasonality of soil termites in a humid tropical forest, Mbalmayo, southern Cameroon. Journal of Tropical Ecology, 14 : 841 - 850.

Didham RK, Hammond PM, Lawton JH, Eggleton P, Stork NE (1998) Beetle species responses to tropical forest fragmentation. Ecological Monographs, 68 : 295 - 323.

Didham RK, Lawton JH, Hammond PM, Eggleton P (1998) Trophic structure stability and extinction dynamics of beetles (Coleoptera) in tropical forest fragments. Philosophical Transactions of the Royal Society of London. Series B: Biological Sciences, 353 (1367): 437 - 451. doi: 10.1098/rstb.1998.0221

Eggleton P, Davies RG, Bignell DE (1998) Body size and energy use in termites (Isoptera): the responses of soil feeders and wood feeders differ in a tropical forest assemblage. Oikos, : 525 - 530.

Lawton JH, Bignell DE, Bolton B, Bloemers GF, Eggleton P, Hammond PM, Hodda M, Holt RD, Larsen TB, Mawdsley NA, others (1998) Biodiversity inventories, indicator taxa and effects of habitat modification in tropical forest. Nature, 391 : 72 - 76.

Macdonald JA, Eggleton P, Bignell DE, Forzi F, Fowler D (1998) Methane emission by termites and oxidation by soils, across a forest disturbance gradient in the Mbalmayo Forest Reserve, Cameroon. Global Change Biology, 4 : 409 - 418.

Sleaford F, BIGNELL DE, Eggleton P (1998) A pilot analysis of gut contents in termites from the Mbalmayo Forest Reserve, Cameroon. Ecological Entomology, 21 : 279 - 288.

Bignell DE, Eggleton P, Nunes L, Thomas KL (1997) Termites as mediators of carbon fluxes in tropical forest: budgets for carbon dioxide and methane emissions. Forests and insects, : 109 - 134.

Eggleton P, Bignell DE (1997) Secondary occupation of epigeal termite (Isoptera) mounds by other termites in the Mbalmayo Forest Reserve, southern Cameroon, and its biological significance. Journal of African Zoology (Belgium),

Eggleton P, Homathevi R, Jeeva D, Jones DT, Davies RG, Maryati M (1997) The species richness and composition of termites (Isoptera) in primary and regenerating lowland dipterocarp forest in Sabah, East Malaysia. Ecotropica, 3 : 119 - 128.

Nunes L, Bignell DE, Lo N, Eggleton P (1997) On the respiratory quotient (RQ) of termites (Insecta: Isoptera). Journal of insect physiology, 43 : 749 - 758.

Tayasu I, Abe T, Eggleton P, Bignell DE (1997) Nitrogen and carbon isotope ratios in termites: An indicator of trophic habit along the gradient from wood-feeding to soil-feeding. Ecological Entomology, 22 : 343 - 351.

Watt AD, Stork NE, Eggleton P, Srivastava D, Bolton B, Larsen TB, Brendell MJD, Bignell DE (1997) Impact of forest loss and regeneration on insect abundance and diversity. Forests and insects, : 273 - 286.

Dharmarajah J, Mohamed M, Eggleton P, Bignell D, Sabah UM (1996) FLUKS KARBON OLEH ANAI-ANAIDI HUT AN rf1 JJANTROPIKA LEMBAH DANUM, SAB AH. Ke arah pengeksploitasian dan pengurusan sumber alam yang mapan, : 119 - 119.

Eggleton P, Bignell DE, Sands WA, Mawdsley NA, Lawton JH, Wood TG, Bignell NC (1996) The diversity, abundance and biomass of termites under differing levels of disturbance in the Mbalmayo Forest Reserve, southern Cameroon. Philosophical Transactions of the Royal Society B: Biological Sciences, 351 : 51 - 68.

Lawton JH, Bignell DE, Bloemers GF, Eggleton P, Hodda ME (1996) Carbon flux and diversity of nematodes and termites in Cameroon forest soils. Biodiversity & Conservation, 5 : 261 - 273.

Gaston KJ, Williams PH, Eggleton P, Humphries CJ (1995) Large Scale Patterns of Biodiversity: Spatial Variation in Family Richness. Proceedings of the Royal Society B: Biological Sciences, 260 (1358): 149 - 154. doi: 10.1098/rspb.1995.0072

Bignell DE, Eggleton P (1995) On the elevated intestinal pH of higher termites (Isoptera: Termitidae). Insectes Sociaux, 42 : 57 - 69.

Eggleton P, Bignell DE (1995) Monitoring the response of tropical insects to changes in the environment: troubles with termites. Insects in a changing environment, : 473 - 497.

Eggleton P, Bignell DE, Sands WA, Waite B, Wood TG, Lawton JH (1995) The species richness of termites (Isoptera) under differing levels of forest disturbance in the Mbalmayo Forest Reserve, southern Cameroon. Journal of Tropical Ecology, 11 : 85 - 98.

Eggleton P, Williams PH, Gaston KJ (1994) Explaining global termite diversity: productivity or history?. Biodiversity and Conservation, 3 (4): 318 - 330. doi: 10.1007/BF00056505

Eggleton P (1994) Termites live in a pear-shaped world: a response to Platnick. null,

Eggleton P, Vane-Wright RI (1994) Some principles of phylogenetics and their implications for comparative biology. Phylogenetics and ecology, : 345 - 366.

Eggleton P, Vane-Wright RI (1994) Phylogenetics and ecology. Linnean Society symposium series,

Wiseman S, Eggleton P. , (1994) The termiticide Market. PJB Publications.

Eggleton P, Belshaw R (1993) Comparisons of dipteran, hymenopteran and coleopteran parasitoids: provisional phylogenetic explanations. Biological Journal of the Linnean Society, 48 : 213 - 226.

Eggleton P, Belshaw R (1992) Insect parasitoids: an evolutionary overview. Philosophical Transactions of the Royal Society of London. Series B: Biological Sciences, 337 : 1 - 20.

Stork NE, Eggleton P (1992) Invertebrates as determinants and indicators of soil quality. American journal of alternative agriculture, 7 : 38 - 47.

Eggleton P (1991) Patterns in male mating strategies of the Rhyssini: a holophyletic group of parasitoid wasps (Hymenoptera: Ichneumonidae). Animal behaviour, 41 : 829 - 837.

Eggleton P, Gaston KJ (1990) " Parasitoid" Species and Assemblages: Convenient Definitions or Misleading Compromises?. Oikos, : 417 - 421.

EGGLETON P (1990) Male reproductive behaviour of the parasitoid wasp Lytarmes maculipennis (Hymenoptera: Ichneumonidae). Ecological entomology, 15 : 357 - 360.

Eggleton P (1989) The phylogeny and evolutionary biology of the Pimplinae (Hymenoptera: Ichneumonidae). Unpublished PhD thesis, University of London,

Watt A, Argent G, Bibby C, Lengeler JC, Eggleton P, Garwood N, Gillison A, Hawthorne W, Healey J, Hall J, others (1989) Evaluation and development of methods of rapid biodiversity assessment in relation to the conservation of biodiversity in tropical moist forest: final report. null,