Dr Ian Kitching

Dr Ian Kitching

Researcher, Entomology

Department: Life Sciences
Division: LS Insects
Contact: email

Specialisms

Lepidoptera, cladistics, morphology, nomenclature, phylogenetics, taxonomy

Summary

I am a systematic phylogenetic entomologist undertaking collections-based research to understand the evolution, ecology, biogeography and taxonomy of Lepidoptera with particular reference to the hawkmoths (Sphingidae) and their relatives. With colleagues in Europe and North America, I am investigating: the co-evolution between hawkmoths and their bat predators; the importance and influence of hybridization in the origin of species in the genus Hyles; and the utility of hawkmoths as a macroecological indicator group. In addition, I am developing the Sphingidae Taxonomic Inventory to make taxonomic data on this group of charismatic moths widely available, as well as converting the LepIndex, generic Names and HOSTS databases to a Species File, to produce a community editable, global taxonomic resource for all with an interest in Lepidoptera systematics. I am also part of the Mosquito Group, undertaking phylogenetic analyses of the highly economically important group of disease-vectoring insects.

Qualifications

Degrees

PhD, Imperial College, United Kingdom

Employment history

Academic

Junior Research Fellow to Band 3, NHM, United Kingdom, 1983 - ongoing

Publications

Rougerie R, Kitching IJ, Haxaire J, Miller SE, Hausmann A, Hebert PDN (2014) Australian Sphingidae – DNA Barcodes Challenge Current Species Boundaries and Distributions. PLoS ONE, 9 (7)doi: 10.1371/journal.pone.0101108

SIMONSEN THOMASJ, KITCHING IANJ (2014) Virtual dissections through micro-CT scanning: a method for non-destructive genitalia ‘dissections’ of valuable Lepidoptera material. Systematic Entomology, 39 (3): 606 - 618. doi: 10.1111/syen.12067

RAFI MUHAMMADATHER, SULTAN AMIR, KITCHING IANJ, PITTAWAY AR, MARKHASIOV MAXIM, KHAN MUHAMMADRAFIQUE, NAZ FALAK (2014) The Hawkmoth Fauna of Pakistan (Lepidoptera: Sphingidae). Zootaxa, 3794 (3): 393 - 393. doi: 10.11646/zootaxa.3794.3.4

Brechlin R, Kitching IJ (2014) Drei neue Arten der Gattung Cypa Walker, [1865] (Lepidoptera: Sphingidae). Entomo-Satsphingia, 7 (2): 5 - 14.

Ivshin N, Kitching IJ (2014) A new species of the genus Ambulyx Westwood, 1847 (Lepidoptera: Sphingidae) from northern Borneo and adjacent territories. Nachrichten des Entomologischen Vereins Apollo (N.F.), 35 (3): 109 - 119.

Kitching IJ, Culverwell CL, Harbach RE (2014) The phylogenetic conundrum of Lutzia (Diptera: Culicidae: Culicini): a cautionary account of conflict and support. Insect Systematics & Evolution, doi: 10.1163/1876312X-45032119

Kudo K, Nakao K, Kitching IJ (2014) A new species of the genus Acosmeryx Boisduval, [1875] from Yunnan and Guangxi, China (Lepidoptera, Sphingidae). Tinea, 23 (1): 35 - 40.

Moré M, Kitching IJ, Cocucci AA (2014) Sphingidae. , Roig -Juñent S, Claps LE, Morrone JJ, Biodiversidad de Artrópodos Argentinos., pp. 281 - 295. INSUE - UNTEediciones. San Miguel de Tucumán, Argentina.

Singh IJ, Kitching IJ (2014) A first record of Clanis hyperion Cadiou & Kitching, 1990 (Lepidoptera: Sphingidae) in Bhutan, and a preliminary checklist of the hawkmoths of Mendrelgang, Bhutan. Journal of Threatened Taxa, 6 (1): 5386 - 5388. doi: 10.11609/JoTT.o3399.5386-8

Kawahara AY, Breinholt JW, Ponce FV, Haxaire J, Xiao L, Lamarre GPA, Rubinoff D, Kitching IJ (2013) Evolution of Manduca sexta hornworms and relatives: Biogeographical analysis reveals an ancestral diversification in Central America. Molecular Phylogenetics and Evolution, 68 (3): 381 - 386. doi: 10.1016/j.ympev.2013.04.017

HARBACH RALPHE, KITCHING IANJ, CULVERWELL CLORNA, HOWARD THERESAM, LINTON YVONNE-MARIE (2013) Nyx pholeocola, a new genus and cavernicolous species of tribe Aedini (Diptera: Culicidae) from southern Thailand based on morphological and molecular data. Zootaxa, 3683 (2): 159 - 159. doi: 10.11646/zootaxa.3683.2.5

Ballesteros-Mejia L, Kitching IJ, Jetz W, Nagel P, Beck J (2013) Mapping the biodiversity of tropical insects: species richness and inventory completeness of African sphingid moths. Global Ecology and Biogeography, 22 (5): 586 - 595. doi: 10.1111/geb.12039

Beck J, Ballesteros-Mejia L, Nagel P, Kitching IJ (2013) Online solutions and the ‘Wallacean shortfall’: what does GBIF contribute to our knowledge of species' ranges?. Diversity and Distributions, 19 (8): 1043 - 1050. doi: 10.1111/ddi.12083

Kawahara AK, Breinholt J, Ponce F, Haxaire J, Xiao L, Lamarre GPA, Kitching IJ (2013) Phylogeny and evolution of Manduca sexta and relatives: ancestral biogeographical analysis reveals a Central american origin for the genus and species. Molecular Phylogenetics and Evolution,

Kitching IJ (2013) A new species of Cypa from New Guinea (Lepidoptera, Sphingidae, Smerinthinae). Suara Serangga Papua, 7 : 85 - 91.

Vaglia T, Kitching IJ (2013) Un nouveau Sphingidae du genre Acosmeryx Boisduval, [1875] décrit du Sabah (Bornéo), (Lepidoptera, Sphingidae). The European Entomologist, 4 : 161 - 168.

Zahiri R, Lafontaine JD, Holloway JD, Kitching IJ, Schmidt BC, Kaila L, Wahlberg N (2013) Major lineages of Nolidae (Lepidoptera, Noctuoidea) elucidated by molecular phylogenetics. Cladistics, 29 (4): 337 - 359. doi: 10.1111/cla.12001

Zahiri R, Lafontaine D, Schmidt C, Holloway JD, Kitching IJ, Mutanen M, Wahlberg N (2013) Relationships among the basal lineages of Noctuidae (Lepidoptera, Noctuoidea) based on eight gene regions. Zoologica Scripta, 42 (5): 488 - 507. doi: 10.1111/zsc.12022

Harbach RE, Kitching IJ, Culverwell CL, Dubois J, Linton Y-M (2012) Phylogeny of mosquitoes of tribe Culicini (Diptera: Culicidae) based on morphological diversity. Zoologica Scripta, 41 (5): 499 - 514. doi: 10.1111/j.1463-6409.2012.00546.x

Blagoderov V, Kitching I, Livermore L, Simonsen T, Smith V (2012) No specimen left behind: industrial scale digitization of natural history collections. ZooKeys, 209 (209): 133 - 146. doi: 10.3897/zookeys.209.3178

ARDITTI JOSEPH, ELLIOTT JOHN, KITCHING IANJ, WASSERTHAL LUTZT (2012) ‘Good Heavens what insect can suck it’- Charles Darwin, Angraecum sesquipedale and Xanthopan morganii praedicta. Botanical Journal of the Linnean Society, 169 (3): 403 - 432. doi: 10.1111/j.1095-8339.2012.01250.x

Beck J, Holloway JD, Khen CV, Kitching IJ (2012) Diversity Partitioning Confirms the Importance of Beta Components in Tropical Rainforest Lepidoptera. The American Naturalist, 180 (3)doi: 10.1086/666982

Brechlin R, Kitching IJ (2012) Ein neue Art der Gattung Callambulyx Rothschild & Jordan, 1903 (Lepidoptera: Sphingidae). Entomo-Satsphingia, 5 (3): 56 - 60.

Rougerie R, Haxaire J, Kitching IJ, Hebert PDN (2012) DNA barcodes and morphology reveal a hybrid hawkmoth in Tahiti (Lepidoptera : Sphingidae). Invertebrate Systematics, 26 (6): 445 - 445. doi: 10.1071/IS12029

Smetacek P, Kitching IJ (2012) The hitherto undescribed male of Orthozona quadrilineata (Moore, 1862) (Lepidoptera: Erebidae). Journal of Threatened Taxa, 4 (14): 3366 - 3368.

Smetacek P, Kitching IJ (2012) The hawkmoths of Ladakh, Jammu & Kashmir, India (Lepidoptera: Sphingidae). Nachrichten des Entomologischen Vereins Apollo (N.F.), 32 : 113 - 115.

ZAHIRI REZA, HOLLOWAY JEREMYD, KITCHING IANJ, LAFONTAINE JDONALD, MUTANEN MARKO, WAHLBERG NIKLAS (2012) Molecular phylogenetics of Erebidae (Lepidoptera, Noctuoidea). Systematic Entomology, 37 (1): 102 - 124. doi: 10.1111/j.1365-3113.2011.00607.x

Ballesteros-Mejia L, Kitching IJ, Beck J (2011) Projecting the potential invasion of the Pink Spotted Hawkmoth ( Agrius cingulata ) across Africa. International Journal of Pest Management, 57 (2): 153 - 159. doi: 10.1080/09670874.2010.546439

Hundsdoerfer A, Pittaway A, Mende M, Kitching I, Harbich H (2011) Larval pattern morphotypes in the Western Palaearctic Hyles euphorbiae complex (Lepidoptera: Sphingidae: Macroglossinae). Insect Systematics & Evolution, 42 (1): 41 - 86. doi: 10.1163/187631211X555302

Hundsdoerfer AK, Mende MB, Kitching IJ, Cordellier M (2011) Taxonomy, phylogeography and climate relations of the Western Palaearctic spurge hawkmoth (Lepidoptera, Sphingidae, Macroglossinae). Zoologica Scripta, 40 (4): 403 - 417. doi: 10.1111/j.1463-6409.2011.00477.x

Ignatov II, Janovec JP, Centeno P, Tobler MW, Grados J, Lamas G, Kitching IJ (2011) Patterns of Richness, Composition, and Distribution of Sphingid Moths Along an Elevational Gradient in the Andes-Amazon Region of Southeastern Peru. Annals of the Entomological Society of America, 104 (1): 68 - 76. doi: 10.1603/AN09083

Kitching IJ (2011) Reconciling the zoological and botanical approaches to taxonomy and nomenclature: lessons from the CATE experience. Nature Precedings,

Kitching IJ (2011) A new species of Leptoclanis Rothschild & Jordan, 1903 from Upemba National Park, Democratic Republic of Congo (Lepidoptera: Sphingidae). Entomologia Africana, 16 : 7 - 12.

Kitching I, Sadler S (2011) Lepidoptera, Insecta. , Harrison T, Paleontology and geology of Laetoli: human evolution in context., pp. 549 - 554. Springer Science + Business Media B.V.. Dordrecht.

Van Nieukerken E, Kaila L, Kitching IJ, Kristensen NP, Lees DC, Minet J, Mitter C, Mutanen M, Regier JC, Simonsen TJ, Wahlberg N, Yen S-H, Zahiri R, Adamski D, Baixeras J, Bartsch D, Bengtsson B, Brown JW, Bucheli SR, Davis DR, De Prins J, De Prins W, Epstein ME, Gentili-Poole P, Gielis C, Hättenschwiler P, Hausmann A, HOLLOWAY JD, Kallies A, Karsholt O, Kawahara AY, Koster JC, Kozlov MV, Lafontaine JD, Lamas G, Landry J-F, Lee S, Nuss M, Park K-T, Penz CM, Rota J, Schintlmeister A, Schmidt BC, Sohn J-C, Solis MA, Tarmann GM, Warren AD, Weller S, Yakovlev RV, Zolotuhin VV, Zwick A (2011) Order Lepidoptera. In: Zhang, Z.-Q. (ed) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa, 3148 : 212 - 221.

Wilson JJ, Rougerie R, Schonfeld J, Janzen DH, Hallwachs W, Hajibabaei M, Kitching IJ, Haxaire J, Hebert PDN (2011) When species matches are unavailable are DNA barcodes correctly assigned to higher taxa? An assessment using sphingid moths. BMC Ecology, 11

Attie M, Kitching IJ, Veslot J (2010) PATTERNS OF LARVAL HOSTPLANT USAGE AMONG HAWKMOTHS (LEPIDOPTERA, SPHINGIDAE) FROM LA REUNION, WITH A COMPARISON OF THE MASCARENES WITH OTHER REGIONS OF THE WORLD. REVUE D ECOLOGIE-LA TERRE ET LA VIE, 65 (1): 3 - 44.

Brechlin R, Kitching IJ (2010) Drei neue Taxa der miranda-Gruppe der Gattung Loepa Moore, 1859 (Lepidoptera, Saturniidae). Entomo-Satsphingia, 3 (1): 12 - 18.

Brechlin R, Kitching IJ (2010) Eine neue Art der Gattung Salassa Moore, 1859 (Lepidoptera: Saturniidae, Salassinae). Entomo-Satsphingia, 3 (1): 9 - 11.

Brechlin R, Kitching IJ (2010) Vier neue Arten der Gattung Psilogramma Rothschild & Jordan, 1903 (Lepidoptera: Sphingidae). Entomo-Satsphingia, 3 (3): 66 - 74.

Brechlin R, Kitching IJ (2010) Einige Anmergunken zur moorei-Gruppe der Gattung Ambulyx Westwood, 1847 mit Beschreibung neur taxa (Lepidoptera, Sphingidae). Entomo- Satsphingia, 3 (3): 20 - 26.

Hundsdoerfer AK, Kitching IJ (2010) A Method for Improving DNA Yield from Century-plus Old Specimens of Large Lepidoptera while Minimizing Damage to External and Internal Abdominal Characters. ARTHROPOD SYSTEMATICS & PHYLOGENY, 68 (1): 151 - 155.

Nässig WA, Kitching IJ, Peigler RS, Treadaway CG (2010) The group of Cricula elaezia: comments on synonyms and priority questions, with illustrations of barcode similarity trees, distribution maps, a revised checklist and a formerly unknown female (Lepidoptera: Saturniidae). Nachrichten des Entomologischen Vereins Apollo (N.F.), 11 : 145 - 165.

Vaglia T, Haxaire J, Kitching I, Liyous M (2010) Contribution à la connaissance des Theretra Hübner, 1819, des complexes clotho (Drury, 1773), boisduvalii (Bugnion, 1839) et rhesus (Boisduval, [1875]) d’Asie continentale et du Sud-est (Lepidoptera, Sphingidae). The European Entomologist 3: 41-77. The European Entomologist, 3 : 41 - 77.

Zahiri R, Kitching IJ, Lafontaine JD, Mutanen M, Kaila L, Holloway JD, Wahlberg N (2010) A new molecular phylogeny offers hope for a stable family level classification of the Noctuoidea (Lepidoptera). Zoologica Scripta, 40 (2)doi: 10.1111/j.1463-6409.2010.00459.x

REINERT JOHNF, HARBACH RALPHE, KITCHING IANJ (2009) Phylogeny and classification of tribe Aedini (Diptera: Culicidae). Zoological Journal of the Linnean Society, 157 (4): 700 - 794. doi: 10.1111/j.1096-3642.2009.00570.x

Mauricio Bini L, Diniz-Filho JAF, Rangel TFLVB, Akre TSB, Albaladejo RG, Albuquerque FS, Aparicio A, Araújo MB, Baselga A, Beck J, Isabel Bellocq M, Böhning-Gaese K, Borges PAV, Castro-Parga I, Khen Chey V, Chown SL, de Marco, Jr P, Dobkin DS, Ferrer-Castán D, Field R, Filloy J, Fleishman E, Gómez JF, Hortal J, Iverson JB, Kerr JT, Daniel Kissling W, Kitching IJ, León-Cortés JL, Lobo JM, Montoya D, Morales-Castilla I, Moreno JC, Oberdorff T, Olalla-Tárraga MÁ, Pausas JG, Qian H, Rahbek C, Rodríguez MÁ, Rueda M, Ruggiero A, Sackmann P, Sanders NJ, Carina Terribile L, Vetaas OR, Hawkins BA (2009) Coefficient shifts in geographical ecology: an empirical evaluation of spatial and non-spatial regression. Ecography, 32 (2): 193 - 204. doi: 10.1111/j.1600-0587.2009.05717.x

Beccaloni G, Holloway J, Kitching IJ, Sattler K, Scoble MJ, Tuck K, Vane-Wright RI (2009) A tribute to Dr Gaden S. Robinson (1949-2009). Systematics and Biodiversity, 7 : 351 - 354.

Beck J, Kitching IJ (2009) Drivers of moth species richness on tropical altitudinal gradients: a cross-regional comparison. Global Ecology and Biogeography, 18 (3): 361 - 371. doi: 10.1111/j.1466-8238.2009.00447.x

Brechlin R, Kitching IJ (2009) Ein Überblick über die Gattung Sataspes Moore, [1858] mit Beschreibung zweier neuer Arten von den Philippinen (Lepidoptera, Sphingidae). Entomo-Satsphingia, 2 (1): 13 - 22.

Brechlin R, Kitching IJ (2009) Degmaptera cadioui n. sp., eine neue Sphingide (Lepidoptera) von Palawan, Philippinen. Entomo-Satsphingia, 2 (1): 5 - 7.

Clark BR, Godfray HCJ, Kitching IJ, Mayo SJ, Scoble MJ (2009) Taxonomy as an eScience. Philosophical Transactions of the Royal Society A: Mathematical, Physical and Engineering Sciences, 367 (1890): 953 - 966. doi: 10.1098/rsta.2008.0190

Hundsdoerfer AK, Rubinoff D, Attié M, Wink M, Kitching IJ (2009) A revised molecular phylogeny of the globally distributed hawkmoth genus Hyles (Lepidoptera: Sphingidae), based on mitochondrial and nuclear DNA sequences. Molecular Phylogenetics and Evolution, 52 (3): 852 - 865. doi: 10.1016/j.ympev.2009.05.023

JANZEN DANIELH, HALLWACHS WINNIE, BLANDIN PATRICK, BURNS JOHNM, CADIOU JEAN-MARIE, CHACON ISIDRO, DAPKEY TANYA, DEANS ANDREWR, EPSTEIN MARCE, ESPINOZA BERNARDO, FRANCLEMONT JOHNG, HABER WILLIAMA, HAJIBABAEI MEHRDAD, HALL JASONPW, HEBERT PAULDN, GAULD IAND, HARVEY DONALDJ, HAUSMANN AXEL, KITCHING IANJ, LAFONTAINE DON, LANDRY J-F, LEMAIRE CLAUDE, MILLER JACQUELINEY, MILLER JAMESS, MILLER LEE, MILLER SCOTTE, MONTERO JOSE, MUNROE EUGENE, GREEN SUZANNERAB, RATNASINGHAM SUJEEVAN, RAWLINS JOHNE, ROBBINS ROBERTK, RODRIGUEZ JOSEPHINEJ, ROUGERIE RODOLPHE, SHARKEY MICHAELJ, SMITH MALEX, SOLIS MALMA, SULLIVAN JBOLLING, THIAUCOURT PAUL, WAHL DAVIDB, WELLER SUSANJ, WHITFIELD JAMESB, WILLMOTT KEITHR, WOOD DMONTY, WOODLEY NORMANE, WILSON JOHNJ (2009) Integration of DNA barcoding into an ongoing inventory of complex tropical biodiversity. Molecular Ecology Resources, 9 : 1 - 26. doi: 10.1111/j.1755-0998.2009.02628.x

Kawahara AY, Mignault AA, Regier JC, Kitching IJ, Mitter C (2009) Kawahara, A.Y., Mignault, A.A., Regier, J.C., Kitching, I.J. & Mitter, C. Phylogeny and biogeography of hawkmoths (Lepidoptera: Sphingidae): evidence from five nuclear genes. PLoS One, 4 (5)

Regier JC, Zwick A, Cummings MP, Kawahara AY, Cho S, Weller S, Roe A, Baixeras J, Brown JW, Parr C, Davis DR, Epstein M, Hallwachs W, Hausmann A, Janzen DH, Kitching IJ, Solis MA, Yen S-H, Bazinet AL, Mitter C (2009) Toward reconstructing the evolution of advanced moths and butterflies (Lepidoptera: Ditrysia): an initial molecular study. BMC Evolutionary Biology, 9 (1): 280 - 280. doi: 10.1186/1471-2148-9-280

Mayo S, Allkin R, Baker W, Blagoderov V, Brake I, Clark B, Govaerts R, Godfray HCJ, Haigh A, Hand R, Harman K, Jackson M, Kilian N, Kirkup DW, Kitching IJ, Knapp S, Lewis GP, Malcolm-Tompkins P, von Raab-Straube E, Roberts DM, Scoble MJ, Simpson D, Smith CR, Smith V, Villalba S, Walley L, Wilkin P (2008) Alpha e-taxonomy: responses from the systematics community to the biodiversity crisis. Kew Bulletin, 63 : 1 - 16.

REINERT JOHNF, HARBACH RALPHE, KITCHING IANJ (2008) Phylogeny and classification of Ochlerotatus and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society, 153 (1): 29 - 114. doi: 10.1111/j.1096-3642.2008.00382.x

Smith CR, Godfray HC, Scoble MJ, Clark BR, Kitching IJ, Mayo SJ, Blagoderov V, Haigh AL, Jackson M, Sadler S, Lay LN, Young RPW (2008) Introducing CATE - a model for moving taxonomy to the web. , Yata O, The 2nd Report on Insect Inventory Project in Tropical Asia (TAIIV) "The development of insect inventory project in Tropical Asia (TAIIV)"., pp. 137 - 144. Kyushu University. Fukuoka, Japan.

Vaglia T, Haxaire J, Kitching IJ, Meusnier I, Rougerie R (2008) Morphology and DNA barcoding reveal three cryptic species within the Xylophanes neoptolemus and loelia species-groups (Lepidoptera: Sphingidae). ZOOTAXA, 1923 (1923): 18 - 36.

Harbach RE, Rattanarithikul R, Howard TM, Linton Y-M, Kitching IJ (2007) Systematics of a new genus and cavernicolous species of the mosquito tribe Aedini (Diptera : Culicidae) from Thailand. PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON, 109 (2): 469 - 488.

Harbach RE, Vu Duc Huong , Kitching IJ (2007) Systematics of Kimia, a new genus of Sabethini (Diptera: Culicidae) in the Oriental Region. Proceedings of the Entomological Society of Washington, 109 (1): 102 - 120.

BECK JAN, KITCHING IANJ (2007) Estimating regional species richness of tropical insects from museum data: a comparison of a geography-based and sample-based methods. Journal of Applied Ecology, 44 (3): 672 - 681. doi: 10.1111/j.1365-2664.2007.01291.x

Beck J, Kitching IJ (2007) Correlates of range size and dispersal ability: a comparative analysis of sphingid moths from the Indo-Australian tropics. Global Ecology and Biogeography, 16 (3): 341 - 349. doi: 10.1111/j.1466-8238.2007.00289.x

Beck J, Kitching IJ (2007) The latitudinal distribution of sphingid species richness in continental southeast Asia: What causes the biodiversity 'hot spot' in northern Thailand?. RAFFLES BULLETIN OF ZOOLOGY, 55 (1): 179 - 185.

Brechlin R, Kitching IJ (2007) Zwei neue Taxa aus der Gattung Acosmeryx Boisduval, [1875] (Lepidoptera: Sphingidae). Nachrichten des Entomologischen Vereins Apollo (N.F.), 28 : 5 - 10.

Godfray HCJ, Clark BR, Kitching IJ, Mayo SJ, Scoble MJ (2007) The Web and the Structure of Taxonomy. Systematic Biology, 56 (6): 943 - 955. doi: 10.1080/10635150701777521

Hawkins BA, Albuquerque FS, Araújo MB, Beck J, Bini LM, Cabrero-Sañudo FJ, Castro-Parga I, Diniz-Filho JAF, Ferrer-Castán D, Field R, Gómez JF, Hortal J, Kerr JT, Kitching IJ, León-Cortés JL, Lobo JM, Montoya D, Moreno JC, Olalla-Tárraga MÁ, Pausas JG, Qian H, Rahbek C, Rodríguez MÁ, Sanders NJ, Williams P (2007) A GLOBAL EVALUATION OF METABOLIC THEORY AS AN EXPLANATION FOR TERRESTRIAL SPECIES RICHNESS GRADIENTS. Ecology, 88 (8): 1877 - 1888. doi: 10.1890/06-1444.1

Haxaire J, Kitching IJ, Melichar T (2007) Hommage à Jean-Marie Cadiou. The European Entomologist, 1 : 24 - 28.

Kitching IJ, Zahiri R (2007) Taxonomic notes and new records of the genus Sphingonaepiopsis Wallengren, 1858 (Lepidoptera : Sphingidae) in Iran. ZOOTAXA, 1610 (1610): 41 - 52.

Scoble MJ, Clark BR, Godfray HCJ, Kitching IJ, Mayo SJ (2007) Revisionary taxonomy in a changing e-landscape. Tijdschrift voor Entomologie, 150 : 305 - 317.

Beck J, KITCHING IJ, Linsenmair KE (2006) Extending the study of range-abundance relations to tropical insects: sphingid moths in Southeast Asia. Evolutionary Ecology Research, 8 (4): 677 - 690.

Beck J, Kitching IJ (2006) Hawkmoths (Lepidoptera, Sphingidae) of Southeast Asia: online distribution maps and island checklists. Zoological Society of Wallacea Newsletter, 4 : 3 - 14.

Beck J, Kitching IJ, Linsenmair KE (2006) Measuring range sizes of South-East Asian hawkmoths (Lepidoptera: Sphingidae): effects of scale, resolution and phylogeny. Global Ecology and Biogeography, 15 (4): 339 - 348. doi: 10.1111/j.1466-822X.2006.00230.x

Beck J, Kitching IJ, Linsenmair KE (2006) Diet breadth and host plant relationships of Southeast-Asian sphingid caterpillars. Ecotropica, 12 : 1 - 13.

Beck J, Kitching IJ, Eduard Linsenmair K (2006) Determinants of regional species richness: an empirical analysis of the number of hawkmoth species (Lepidoptera: Sphingidae) on the Malesian archipelago. Journal of Biogeography, 33 (4): 694 - 706. doi: 10.1111/j.1365-2699.2005.01418.x

BECK JAN, KITCHING IANJ, LINSENMAIR KEDUARD (2006) Wallace’s line revisited: has vicariance or dispersal shaped the distribution of Malesian hawkmoths (Lepidoptera: Sphingidae)?. Biological Journal of the Linnean Society, 89 (3): 455 - 468. doi: 10.1111/j.1095-8312.2006.00686.x

Beck J, Kitching IJ, Linsenmair KE (2006) Effects of Habitat Disturbance can be Subtle Yet Significant: Biodiversity of Hawkmoth-Assemblages (Lepidoptera: Sphingidae) in Southeast-Asia. Biodiversity and Conservation, 15 (1): 465 - 486. doi: 10.1007/s10531-005-0306-6

Kitching IJ. , (2006) The biology of Death’s Head Hawkmoths, lepidopteran kleptoparasites of honey bees. The Central Association of Bee-Keepers. 20

León-Cortés JL, Kitching IJ, Molina-Martínez A (2006) Distribution patterns of Sphingidae in Mexico. , Llorente-Bousquets J, Morrone JJ, Componentes bióticos principales de la entomofauna mexicana., pp. 629 - 647. Facultad de Ciencias, Universidad Nacional Autónoma de México. México, D.F..

REINERT JOHNF, HARBACH RALPHE, KITCHING IANJ (2006) Phylogeny and classification of Finlaya and allied taxa (Diptera: Culicidae: Aedini) based on morphological data from all life stages. Zoological Journal of the Linnean Society, 148 (1): 1 - 101. doi: 10.1111/j.1096-3642.2006.00254.x

Harbach RE, Kitching IJ (2005) Reconsideration of anopheline mosquito phylogeny (Diptera: Culicidae: Anophelinae) based on morphological data. Systematics and Biodiversity, 3 (4): 345 - 374. doi: 10.1017/S147720000500174X

Hundsdoerfer AK, Kitching IJ, Wink M (2005) The phylogeny of the Hyles euphorbiae complex (Lepidoptera : Sphingidae): Molecular evidence from sequence data and ISSR-PCR fingerprints. ORGANISMS DIVERSITY & EVOLUTION, 5 (3): 173 - 198. doi: 10.1016/j.ode.2004.11.012

Hundsdoerfer AK, Kitching IJ, Wink M (2005) A molecular phylogeny of the hawkmoth genus Hyles (Lepidoptera: Sphingidae, Macroglossinae). Molecular Phylogenetics and Evolution, 35 (2): 442 - 458. doi: 10.1016/j.ympev.2005.02.004

Hundsdoerfer AK, Kitching IJ, Wink M (2005) Morphological variability of Hyles t. tithymali (Boisduval, 1832) caterpillars on the Canary Islands (Lepidoptera: Sphingidae). Entomologische Zeitschrift, 115 : 29 - 33.

Moré M, Kitching IJ, Cocucci AA. , (2005) Sphingidae: Esfíngidos de Argentina. Hawkmoths of Argentina. Literature of Latin America. Buenos Aires. 166

REINERT JOHNF, HARBACH RALPHE, KITCHING IANJ (2004) Phylogeny and classification of Aedini (Diptera: Culicidae), based on morphological characters of all life stages. Zoological Journal of the Linnean Society, 142 (3): 289 - 368. doi: 10.1111/j.1096-3642.2004.00144.x

Hundsdoerfer A, Kitching IJ, Wink M (2004) Biogeography and phylogeny of the genus Hyles (Lepidoptera: Sphingidae) based on mt-DNA sequences. Organisms, Diversity & Evolution,

Kitching IJ, Haxaire J (2004) Clanis phalaris cottoni ssp. n., nouveau sphinx du nord de la Thaïlande (Lepidoptera, Sphingidae). Lépidoptères, 2 : 35 - 45.

Watson AT, O’Neill MA, Kitching IJ (2004) Automated identification of live moths (Macrolepidoptera) using digital Automated Identification SYstem (DAISY). Systematics and Biodiversity, 1 : 287 - 300.

Yen SH, Kitching IJ, Tzen CS (2003) A new subspecies of hawkmoth from Lanyu, Taiwan, with a revised and annotated checklist of the Taiwanese Sphingidae (Lepidoptera). ZOOLOGICAL STUDIES, 42 (2): 292 - 306.

Hundsdörfer A, Kitching IJ, Wink M (2003) The molecular phylogeny of the Hyles euphorbiae complex (Lepidoptera: Sphingidae). Organisms, Diversity & Evolution, 3

Kitching IJ (2003) Phylogeny of the death's head hawkmoths, Acherontia[Laspeyres], and related genera (Lepidoptera: Sphingidae: Sphinginae: Acherontiini). Systematic Entomology, 28 (1): 71 - 88. doi: 10.1046/j.1365-3113.2003.00199.x

Kitching IJ, Haxaire J (2003) On the identity of Cechenena aegrota (Butler, 1875) and related species (Lepidoptera, Sphingidae). Bulletin de la Société entomologique de France, 108 : 355 - 360.

KITCHING IANJ (2002) The phylogenetic relationships of Morgan's Sphinx, Xanthopan morganii (Walker), the tribe Acherontiini, and allied long-tongued hawkmoths (Lepidoptera: Sphingidae, Sphinginae). Zoological Journal of the Linnean Society, 135 (4): 471 - 527. doi: 10.1046/j.1096-3642.2002.00021.x

Robinson GS, Ackery PR, Kitching IJ, Beccaloni GW, Hernández LM (2002) Hostplants of the moth and butterfly caterpillars of America north of Mexico. Memoirs of the American Entomological Institute, 69 : 1 - 824.

Ando T (2001) Reproductive Isolation in a Native Population of Petunia sensu Jussieu (Solanaceae). Annals of Botany, 88 (3): 403 - 413. doi: 10.1006/anbo.2001.1485

Brechlin R, Kitching IJ (2001) Neue Informationen zu einigen Arten der Gattung Antheraea Hübner 1819 ["1816"] von den Andamanen, Indien (Lepidoptera: Saturniidae). Nachrichten des Entomologischen Vereins Apollo (N.F.), 22 : 119 - 123.

Brechlin R, Kitching IJ, Cadiou J-M (2001) Description of a new species of the genus Polyptychus Hübner, [1819] "1816" (Lepidoptera: Sphingidae) from the Lesser Sunda Islands, Indonesia, with notes on several species of the subtribe Choerocampina. Nachrichten des Entomologischen Vereins Apollo (N.F.), 22 : 31 - 35.

Cadiou J-M, Kitching IJ (2001) Polyptychus affinis Rothschild & Jordan, 1903, revisited (Lepidoptera: Sphingidae). Entomologica Africana, 6 : 35 - 40.

Kitching IJ, Ledezma J, Baixeras J (2001) An annotated checklist of the Sphingidae of Bolivia (Insecta: Lepidoptera). Gayana, 65 : 79 - 111.

Lachlan RB, Kitching IJ (2001) A new species of hawk moth (Lepidoptera: Sphingidae) from Papua New Guinea. Australian Entomologist, 28 : 117 - 122.

Robinson GS, Ackery PR, Kitching IJ, Beccaloni GW, Hernández LM. , (2001) Hostplants of the moth and butterfly caterpillars of the Oriental Region. The Natural History Museum & Southdene Sdn Bhd. London & Kuala Lumpur. 744

Forey PL, Kitching IJ (2000) Experiments in coding multistate characters. , Scotland R, Pennington RT, Homology and Systematics. Coding characters for phylogenetic analysis., pp. 54 - 80. Taylor & Francis. London.

Kitching IJ, Cadiou J-M. , (2000) Hawkmoths of the world: an annotated and illustrated revisionary checklist (Lepidoptera: Sphingidae). The Natural History Museum & Cornell University Press. London & Ithaca, NY. 226

Kitching IJ, Forey P, Williams DM (2000) Cladistics. , Levin S, Encyclopedia of Biodiversity., pp. 677 - 692. Academic Press. San Diego.

Kitching IJ, Treadaway CG, Hogenes W (2000) Psilogramma villani n. sp., a new hawkmoth from the Philippines and Sulawesi (Lepidoptera: Sphingidae). Nachrichten des Entomologischen Vereins Apollo (N.F.), 21 : 57 - 60.

Pittaway AR, Kitching IJ (2000) Notes on selected species of hawkmoths (Lepidoptera: Sphingidae) from China, Mongolia and the Korean Peninsula. Tinea, 16 : 170 - 211.

Anthony TG, and REH, Kitching IJ (1999) Phylogeny of the Pyretophorus Series of Anopheles subgenus Cellia (Diptera: Culicidae). Systematic Entomology, 24 (2): 193 - 205. doi: 10.1046/j.1365-3113.1999.00092.x

Yela JL, Kitching IJ (1999) La filogenia de Noctuidos, revisada (Insecta: Lepidoptera: Noctuidae). Noctuid phylogeny revisited (Insecta: Lepidoptera: Noctuidae). , Melic A, de Haro JJ, Méndez M, Ribera I, Evolución y filogenia de Arthropoda (Evolution and phylogeny of Arthropoda)., pp. 485 - 520. Sociedad Entomológica Aragonesa.

HARBACH RALPHE, KITCHING IANJ (1998) Phylogeny and classification of the Culicidae (Diptera). Systematic Entomology, 23 (4): 327 - 370. doi: 10.1046/j.1365-3113.1998.00072.x

Cadiou J-M, Kitching IJ (1998) Corrigendum [to the colour plate for "A new species of Ampelophaga from Nepal and northeastern India (Lepidoptera: Sphingidae)"]. Lambillionea, 98

Cadiou J-M, Kitching IJ (1998) A new species of Ampelophaga from Nepal and northeastern India (Lepidoptera: Sphingidae). Lambillionea, 98 : 353 - 358.

Kitching IJ, Forey PL, Humphries CJ, Williams DM. , (1998) Cladistics. Oxford University Press. Oxford. 228

Kitching IJ, Jin XB (1998) A new species of Sphinx (Lepidoptera: Sphingidae) from Sichuan province, China. Tinea, 15 : 275 - 278.

Kitching IJ, Rawlins JE (1998) The Noctuoidea. , Kristensen NP, Handbuch der Zoologie 4 (Arthropoda), (2) (Insecta), (35) Lepidoptera. Moths and butterflies 1 (Evolution, systematics and biogeography)., pp. 355 - 401. Walter de Gruyter. Berlin.

Tennent WJ, Kitching IJ (1998) A reappraisal of two endemic hawkmoths (Lepidoptera: Sphingidae) from the Solomons Archipelago. Nachrichten des Entomologischen Vereins Apollo (N.F.), 19 : 1 - 21.

Kitching IJ, Owada M, Brechlin R (1997) A revision of the genus Pentateucha (Lepidoptera, Sphingidae), with the description of two new species from eastern China and Taiwan. Tinea, 15 : 79 - 93.

Inoue H, Kennett RD, Kitching IJ. , Pinratana A (1996) Sphingidae. Chok Chai Press. Bangkok. 149

Kitching IJ (1996) Identifying complementary areas for conservation in Thailand: an example using owls, hawkmoths and tiger beetles. Biodiversity and Conservation, 5 (7): 841 - 858. doi: 10.1007/BF00054738

Kitching IJ, Brechlin R (1996) New species of the genera Rhodoprasina Rothschild & Jordan and Acosmeryx Boisduval from Thailand and Vietnam, with a redescription of Rhodoprasina corolla Cadiou & Kitching (Lepidoptera: Sphingidae). Nachrichten des Entomologischen Vereins Apollo (N.F.), 17 : 51 - 66.

Kitching IJ, Spitzer K (1995) An annotated checklist of the Sphingidae of Vietnam. Tinea, 14 : 171 - 195.

Tuck KR, Kitching IJ (1995) Microwaving moths to relax them. Antenna, 19 : 105 - 106.

Vane-Wright RI, Smith CR, Kitching IJ (1994) Systematic assessment of taxic diversity by summation. , Forey PL, Humphries CJ, Vane-Wright RI, Systematics and conservation evaluation., pp. 309 - 326. Clarendon Press. Oxford.

Hasan SA, Kitching IJ (1993) A cladistic analysis of the tribes of the Pentatomidae (Heteroptera). Kontyu, 61 : 651 - 669.

Kitching IJ, Rudge SR (1993) First record of Darapsa myron (Sphingidae) from Thailand. Journal of the Lepidopterists' Society, 79 : 240 - 242.

Forey PL, Humphries CJ, Kitching IJ, Scotland RW, Siebert DJ, Williams DM. , (1992) Cladistics: a practical course in systematics. Oxford University Press. Oxford. 191

Kitching IJ (1992) Tree building techniques. , Forey PL, Humphries CJ, Kitching IJ, Scotland RW, Siebert DJ, Williams DM, Cladistics: a practical course in systematics., pp. 44 - 71. Oxford University Press. Oxford.

Kitching IJ (1992) The determination of character polarity. , Forey PL, Humphries CJ, Kitching IJ, Scotland RW, Siebert DJ, Williams DM, Cladistics: a practical course in systematics., pp. 22 - 43. Oxford University Press. Oxford.

Walker AK, Kitching IJ, Austin AD (1990) A REASSESSMENT OF THE PHYLOGENETIC RELATIONSHIPS WITHIN THE MICROGASTRINAE (HYMENOPTERA: BRACONIDAE). Cladistics, 6 (3): 291 - 306. doi: 10.1111/j.1096-0031.1990.tb00546.x

Cadiou J-M, Kitching IJ (1990) New Sphingidae from Thailand (Lepidoptera). Lambillionea, 98 (4): 3 - 34.

Carter DJ, Kitching IJ, Scoble MJ (1988) The adaptable caterpillar. The Entomologist, 107 : 68 - 79.

Kitching IJ (1988) A new species of Ocalaria (Noctuidae: Catocalinae) and analysis of some morphological characters useful for elucidating noctuid phylogeny. Journal of the Lepidopterists' Society, 42 : 218 - 230.

KITCHING IANJ (1987) Genitalic heterogeneity in Cornutiplusia circumflexa (Lepidoptera: Noctuidae: Plusiinae). Systematic Entomology, 12 (4): 449 - 457. doi: 10.1111/j.1365-3113.1987.tb00217.x

Kitching IJ (1987) Spectacles and Silver Ys: a synthesis of the systematics, cladistics and biology of the Plusiinae (Lepidoptera: Noctuidae). Bulletin of the British Museum (Natural History) (Entomology), 54 : 76 - 262.

Kitching IJ, Vane-Wright RI, Ackery PR (1987) The cladistics of Ideas (Lepidoptera, Danainae). Cladistics, 3 : 14 - 34.

KITCHING IANJ (1986) Allozyme variation in the milkweed butterflies (Lepidoptera: Danainae). Zoological Journal of the Linnean Society, 86 (4): 367 - 389. doi: 10.1111/j.1096-3642.1986.tb01816.x

DeVries PJ, Harvey DJ, Kitching IJ (1986) The ant associated epidermal organs on the larva of Curetis regula Evans (Lycaenidae). Journal of Natural History, 20 : 621 - 633.

DeVRIES PHILIPJ, KITCHING IANJ, VANE-WRIGHT RI (1985) The systematic position of Antirrhea and Caerois, with comments on the classification of the Nymphalidae (Lepidoptera). Systematic Entomology, 10 (1): 11 - 32. doi: 10.1111/j.1365-3113.1985.tb00561.x

KITCHING IJ (1985) Early stages and the classification of the milkweed butterflies (Lepidoptera: Danainae). Zoological Journal of the Linnean Society, 85 (1): 1 - 97. doi: 10.1111/j.1096-3642.1985.tb01516.x

Kitching IJ (1984) An historical review of the higher classification of the Noctuidae (Lepidoptera). Bulletin of the British Museum (Natural History) (Entomology), 49 : 153 - 234.

Kitching IJ (1984) Enzyme variation in the Danainae. , Vane-Wright RI, Ackery PR, The Biology of Butterflies., pp. 191 - 192.

KITCHING IJ (1984) The use of larval chaetotaxy in butterfly systematics, with special reference to the Danaini (Lepidoptera: Nymphalidae). Systematic Entomology, 9 (1): 49 - 61. doi: 10.1111/j.1365-3113.1984.tb00500.x

Kitching IJ (1983) Classifications: the static products of taxonomists?. Antenna, 7 : 49 - 50.

Highlighted publications

SIMONSEN THOMASJ, KITCHING IANJ (2014) Virtual dissections through micro-CT scanning: a method for non-destructive genitalia ‘dissections’ of valuable Lepidoptera material. Systematic Entomology, 39 (3): 606 - 618. doi: 10.1111/syen.12067

ARDITTI JOSEPH, ELLIOTT JOHN, KITCHING IANJ, WASSERTHAL LUTZT (2012) ‘Good Heavens what insect can suck it’- Charles Darwin, Angraecum sesquipedale and Xanthopan morganii praedicta. Botanical Journal of the Linnean Society, 169 (3): 403 - 432. doi: 10.1111/j.1095-8339.2012.01250.x

Zahiri R, Kitching IJ, Lafontaine JD, Mutanen M, Kaila L, et al (2010) A new molecular phylogeny offers hope for a stable family level classification of the Noctuoidea (Lepidoptera). Zoologica Scripta, 40 (2)doi: 10.1111/j.1463-6409.2010.00459.x

REINERT JOHNF, HARBACH RALPHE, KITCHING IANJ (2009) Phylogeny and classification of tribe Aedini (Diptera: Culicidae). Zoological Journal of the Linnean Society, 157 (4): 700 - 794. doi: 10.1111/j.1096-3642.2009.00570.x

Hundsdoerfer AK, Rubinoff D, Attié M, Wink M, Kitching IJ (2009) A revised molecular phylogeny of the globally distributed hawkmoth genus Hyles (Lepidoptera: Sphingidae), based on mitochondrial and nuclear DNA sequences. Molecular Phylogenetics and Evolution, 52 (3): 852 - 865. doi: 10.1016/j.ympev.2009.05.023

Kawahara AY, Mignault AA, Regier JC, Kitching IJ, Mitter C (2009) Kawahara, A.Y., Mignault, A.A., Regier, J.C., Kitching, I.J. & Mitter, C. Phylogeny and biogeography of hawkmoths (Lepidoptera: Sphingidae): evidence from five nuclear genes. PLoS One, 4 (5)