• Systematics and biology of mites of agricultural and environmental significance, esp. Tetranychoidea, Eriophyoidea, Tarsonemidae, Raphignathoidea, Tydeidae and Phytoseiidae; ecology of Phytoseiidae and their use in biological control
• Systematics, evolution and biology of Parasitengona, esp. world fauna of the Trombidioidea and Erythraeoidea 
• Mite-insect interaction
• Mites of greenhouses

• Analysis of patterns of taxonomic diversity
• Biological and integrated pest control
• Behaviour and dynamics of predator-prey and parasite-host interaction
• Role of internet in science and e-publishing