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'I set out to show a salmon's-eye view of a swimming bear swiping at a fish with its huge paws,' says Paul.
The location he chose was Katmai National Park in Alaska, and he arrived in time for the late-summer runs of red and chum salmon heading upstream to spawn. Grizzly bears cluster around good fishing spots on the rivers to gorge on salmon, in preparation for the long winter. At shallow points, they wade into the water to try to grab the fish. 'I found an accessible pool,' says Paul, 'just below a steep set of river rapids, where the salmon rested before the difficult climb upstream.' He submerged his camera on a ballasted tripod, and with a radio-controlled remote trigger sat across the river and waited - for four days. Bears turned up with different hunting strategies - sprinters and splashers, stalkers and swimmers. This adult female was one of the best hunters, using all the techniques and eventually providing the action Paul was after.
Discover the incredible stories of life on our planet through powerful photography and expert insight.
Tickets on sale now.
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