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The Echinoid Directory

Goniopygus Agassiz, 1838, p. 19

[=Cyphopygus Pomel, 1883, p. 89, type species Goniopygus major Agassiz, 1838; =Polygoniopygus Valette, 1906, p. 11, type species Goniopygus pillati Cotteau, 1883; =Tetragoniopygus Fell & Pawson, 1966, p. U412, type species Goniopygus supremus Hawkins, 1924; =Heteropodia de Loriol, in White, 1887, p. 254, type species Heteropodia whitei de Loriol, 1887 ]

Diagnostic Features
  • Test flattened below, gently domed above to flat apical disc.
  • Apical disc thick forming raised platform; dicyclic; plates firmly bound to corona. Genital plates pentagonal; gonopore at outer interradial point bounded by interambulacral zones; inner edge sunken, forming a periproctal ledge. Ocular plates smaller and projecting. Smooth other than for a single perianal tubercle developed on the inner edge of genital plates which is sunken. Periproct relatively small and subpentagonal.
  • Ambulacra straight; pore-pairs small, uniserial; slightly larger on apical surface, the two pores separated by a raised interporal knob; ambulacra expanded orally into phyllodes.
  • Ambulacral plating in triads throughout, with goniopygid-style compounding; two unequal elements extending to the perradial suture and united by the primary tubercle, plus a third small demiplate in each triad.
  • Interambulacral plates a little wider than tall; with a large primary tubercle on each; secondary tubercles and miliary granulation generally sparse and no scrobicular ring. Primary tubercles largest just above ambitus.
  • Tubercles imperforate and non-crenulate, with no surrounding platform.
  • Basicoronal interambulacral absent; no sphaeridial pits.
  • Peristome subcircular, slightly sunken; buccal notches sharp with raised rim; no tag.
  • Spines relatively short (less than test diameter); stout shaft, with cortical layer in outer part which is often finely ribbed; angular in cross section.
Distribution Upper Jurassic (Tithonian) to Palaeocene, Europe, North Africa, North and South America.
Name gender masculine
Type
Goniopygus peltatus Agassiz, 1838, p. 19, by original designation.
Species Included
  • G. arabicus Smith, 1995; Maastrichtian, United Arab Emirates.
  • G. arnaudi Cotteau, 1883; Turonian, France.
  • G. brossardi Coquand in Cotteau, 1865; Cenomanian, Tunisia.
  • G. conicus Peron & Gauthier; Cenomanian, Algeria.
  • G. coquandi Cotteau, 1865; Cenomanian, Algeria.
  • G. coutini Lambert, 1931; Turonian, Algeria.
  • G. decoratus Desor, 1855; Valanginian, France.
  • G. delphinensis Gras, 1848; Barremian-Aptian, France, England, Spain [includes G. noguesi Cotteau, 1863].
  • G. durandi Peron & Gauthier, 1881; Santonian, Algeria.
  • G. heberti Cotteau, 1866; Danian, Belgium and The Netherlands.
  • G. impressus Peron & Gauthier, 1879; Cenomanian, Algeria.
  • G. innesi Gauthier, 1901; Santonian, Egypt.
  • G. intricatus Agassiz, 1838; Valanginian-Hauterivian, France.
  • G. loryi Cotteau, 1865; Lower Aptian, France.
  • G. major Agassiz, 1838; Cenomanian, France.
  • G. marticensis Cotteau, 1865; Santonian, France.
  • G. menardi Desmarest, 1825; Upper Albian-Cenomanian, western Europe.
  • G. meslei Peron & Gauthier, 1879; Cenomanian, Algeria.
  • G. messauod Peron & Gauthier, 1879; Cenomanian, Algeria.
  • G. minor Sorignet, 1850; Palaeocene, western Europe.
  • G. peltatus (Agassiz, 1836); Barremian-Aptian, France.
  • G. peroni Thomas & Gauthier, 1889; Upper Cenomanian, Tunisia.
  • G. pilleti Cotteau, 1883; Tithonian, France.
  • G. royanus d\'Archiac, 1851; Campanian, France.
  • G. superbus Cotteau & Gauthier, 1895; Campanian-Maastrichtian, Iraq, Saudi Arabia. [see Smith 1995]
  • G. tetraphyma Lambert, 1907: Maastrichtian, Spain.
  • G. whitei (de Loriol, in White 1887); Palaeocene, Brasil.
  • G. whitneyi Smith & Rader, 2009; Albian, Texas, USA
  • G. sp. Smith & Rader, 2009; Lower Albian, USA, Europe
  • plus others....
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    Classification and/or Status

    Echinacea, Arbacioida, Acropeltidae

    Presumed monophyletic.
    Remarks

    The sunken periapical tubercles and smooth unornamented apical disc plates distinguish this from all other Acropeltidae.

    Agassiz, L. 1838. Monographies d'Échinodermes vivants et fossiles. Première monographie: Des Salénies. Petitpierre, Neuchâtel, 32 pp., 5 pls.

    Jagt, J. W. M. 2000. Late Cretaceous-Early Palaeogene echinoderms and the K/T boundary in the southeast Netherlands and northeast Belgium. Part 4:Echinoids. Scripta Geologica 121, 181-375.

    Mortensen, T. 1935. A monograph of the Echinoidea II. Bothriocidaroida, Melonechinoida, Lepidocentroida and Stirodonta. C. A. Reitzel, Copenhagen.

    Smith, A. B. 1995. Late Campanian- Maastrichtian echinoids from the United Arab Emirates-Oman border region. Bulletin of the Natural History Museum, London (Geology Series) 51, 121-240.