The Echinoid Directory

• Test small with moderately sharp margin; ovate in outline.
• Apical disc central with 4 gonopores.
• Internal partitions present; details unrecorded.
• Aboral pores rudimentary; petals absent.
• Interambulacral zones much narrower than ambulacra at ambitus.
• Interambulacrum disjunct on oral but not on aboral surfaces. Two pairs of ambulacral plates separate basicoronals from subsequent interambulacral plates. Interambulacral plating continuous from ambitus aborally.
• Basicoronal and first paired post-basicoronal interambulacral plates small and enclosed by ambulacral basicoronal plates.
• Periproct aboral, towards posterior margin.
• Food grooves absent.
• Pores and tubercles arranged into combed areas towards outer margin of oral surface.

Late Oligocene to Early Miocene, South Australia.

Prowillungaster major Wang, 1994, by original designation.

• P. major Wang, 1994; Oligocene to early Miocene, southern Australia.
• P. minor Wang, 1994; Late Oligocene, Mount Gambier, Australia.

Clypeasteroida; Clypeasterina; Arachnoididae; Arachnoidinae.

?Paraphyletic by exclusion of Willungaster.

Distinguished from Willungaster only in having interambulacral plating continuous from ambitus to apical disc and in lacking sexual dimorphism. Distinguished from Scutellinoides in having small interambulacral plates continuing on the aboral surface to the apex.

Wang, C.-C. 1994. Phylogenetic analysis of the fossulasterid echinoids (Order Clypeasteroida) from Australia. Journal of the Geological Society of China 37, 225-246.