The Echinoid Directory

Clitopygus Pomel, 1833, p.58

[=Clytopygus Poel, 1883, p. 59 (error)]

Diagnostic Features
  • Test subovate, of small to medium size, broad with greatest width posterior to centre; oral surface slightly sunken towards the peristome.
  • Apical system slightly anterior, tetrabasal, with four gonopores, no catenal plates, short posterior ocular plates (advanced form).
  • Petals narrow, open, tapering distally, extending two-thirds distance to ambitus, inner pore round, outer pore slightly elongated transversely; all ambulacral plates double pored.
  • Periproct supramarginal, longitudinal, opening into anal sulcus extending from periproct to posterior margin.
  • Peristome anterior, pentagonal, depressed, equant.
  • Bourrelets absent.
  • Phyllodes slightly widened, double pored, with an inner and an outer series of rounded pores in each half ambulacrum.
  • No buccal pores.
  • No naked zone in posterior interambulacrum orally.
Bajocian (Middle Jurassic) to Cenomanian (Late Cretaceous) of Europe.
Name gender masculine
Echinobrissus lorioli Cotteau, 1871, p. 236; by subsequent designation of Lambert, 1898, p. 468.
Species Included
  • C. lorioli (Cotteau, 1871); Late Bajocian-Callovian, France
  • C. micraulus (L. Agassiz, 1839): Callovian, Europe

Lambert & Thiery (1909-1925) list 53 nominal species within this genus.

Classification and/or Status

Irregularia; Neognathostomata; 'nucleolitid'.

Subjective junior synonym of Nucleolites Lamarck, 1801.


There are no diagnostic features to differentiate Clitopygus from Nucleolites.

G. Cotteau. 1867-1874. Paleontologie francaise ou description de fossiles de la France. Terrain Jurassiques. Tome IX. Echinoidea. G. Masson, Paris. 552 pp., pls. 1-142.

J. Lambert & P. Thiery. 1909-1925. Essai de nomenclature raisonnee des echinides. Libraire Septime Ferriere, Chaumont, 607 pp., 15 pls.

Pomel. A. 1883. Classification méthodique et genera des Échinides vivante et fossiles. Thèses présentées a la Faculté des Sciences de Paris pour obtenir le Grade de Docteur ès Sciences Naturelles, 503, 131 pp. Aldolphe Jourdan, Alger.