The Echinoid Directory

Breynia Desor, in Agassiz & Desor 1847, p. 12

[?Spatangomorpha Boehm, 1882, p. 367, type species Spatangomorpha eximia Boehm, 1882 ]

Diagnostic Features
  • Test ovate with slight anterior sulcus; base slightly keeled towards rear.
  • Apical disc ethmolytic, with 4 gonopores.
  • Anterior ambulacrum moderately wide and a little depressed; pore-pairs and tube-feet specialized.
  • Tubercles adjacent to frontal groove enlarged and in rows.
  • Other ambulacra petaloid and flush; widening towards apex and closed distally. Almost arcuate laterally.
  • Periproct inframarginal, on short near-vertical truncate face.
  • Peristome wider than long; kidney-shaped.
  • Labral plate narrow and elongate; only just contacting sternal plates; paired sternal plates narrow and triangular; fully tuberculate.
  • Aboral tuberculation heterogenous with scattered coarse tubercules in all but the posterior interambulacrum. Primary tubercles deeply sunken. Oral tubercles relatively small and without sunken areoles. Arranged in distinct rows.
  • Well-developed shield-shaped subanal fasciole, an inner fasciole and a peripetalous fasciole that is not indented in the interambulacra.

Oligocene to Recent, Indo-Pacific (India, Australia, Indonesia).

Shallow water sands to shell gravels - infaunal.

Name gender feminine
Spatangus australasiae Leach, 1815, p. 68, by original designation.
Species Included
  • B. australasiae (Leach, 1815); Recent, Australia.
  • B. desorii Gray, 1851; Recent, Australia.
  • B. elegans Mortensen, 1948; Recent, Malay archipelago.
  • B. eximia Boehm, 1882; Miocene, Java.
  • B. vredenburgi Anderson, 1907; Recent, Andamans Islands.
  • B. carinata Haime, 1853;Oligo-Miocene, western Sind, Taiwan, western Australia.
  • B. magna (Herklots); Miocene, Java.
  • B. meslei Gauthier, in Cotteau, 1894; Oligocene, Egypt.
  • B. multituberculata Vredenburg, 1906; Oligocene, India.
  • B. sundaica Gerth, 1922; Miocene, Java.
  • B. paucituberculata (Gerth, 1922); Miocene, Madura.
  • B. birmanica Vredenburg, 1932; Oligocene, Burma.
  • B. cordata Hayasaka, 1947; Pliocene, Taiwan.
  • B. testudinaria Hayasaka, 1947; Pliocene, Taiwan, Japan.
Classification and/or Status

Spatangoida, Micrasterina, Loveniidae.


Distinguished from Lovenia by having both a peripetalous and inner fasciole, and by having the sternal plates of the plastron fully tuberculate. Distinguished from Echinocardium by having prominent aboral primary tubercles in interambulacral zones with sunken areoles, a longer labral plate extending well beyond the first ambulacral plate, and a peripetalous fasciole.

Agassiz, L. & Desor, E. 1846-1847. Catalogue raisonné des familles, des genres, et des espèces de la classe des échinodermes. Annales des Sciences Naturelles, Troisième Série, Zoologie: 6 (1846), 305-374, pls 15-16; 7 (1847), 129-168; 8 (1847), 5-35, 355-380.