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This year's Grand title winner captures the vast yet fragile beauty of the Earth's rainforests.
The image Entwined lives claimed the title over almost 50,000 entries from 95 countries. It is on show with 99 other shots at the Wildlife Photographer of the Year exhibition.
It shows an orang-utan climbing 30 metres above the canopy in Borneo's Gunung Palung National Park. The park is one of the few protected orang-utan strongholds on the island.
American photographer Tim Laman caught the young male's determined scramble to reach a crop of figs at the top of a thick tree.
The animal had already feasted on the crop once and would have had a mental map of the fruiting trees in his range.
Tim knew the orang-utan would return to eat the rest. But the photographer had to spend three days climbing up and down by rope, placing several GoPro cameras in position to capture the shot.
He triggered the cameras remotely to get a chance of not only a wide‐angle view of the forest below, but also a shot of the orang-utan's face from above. This image was the one he had long visualised, looking down on the orang-utan within its forest home.
Gideon Knight from the UK won the Young Wildlife Photographer of the Year title for his image The Moon and the crow.
The 16-year-old turned an ordinary London scene into something more magical, showing a crow silhouetted against a full Moon.
Gideon took the photo near his home, in London's Valentines Park, which he visits regularly to take photographs.
The twigs of the sycamore tree silhouetted against the sky 'made it feel almost supernatural, like something out of a fairy tale,' says Gideon.
Positioning himself on an opposite slope, he tried to capture the perfect composition - but the crow kept moving, forcing Gideon to keep moving too.
Then just as the light was about to fade beyond the point that photography was possible, the birds and the Moon aligned.
The two images were selected from 16 category winners, showing nature at its finest, from displays of animal behaviour to exotic landscapes.