Continuous measures:
The threshold approach to measuring rarity and narrow endemism has been criticised as essentially arbitrary, because whatever threshold of range size is adopted, it can always miss potentially important species with marginally larger range sizes. An alternative approach is to use measures of rangesize rarity that are continuous functions of range size. This escapes the criticism of arbitrariness concerning the precise threshold used to determine which species are included, because all species make a contribution. Instead, rarity is included as a weighting for each species, which is higher for the more restricted species. However, arbitrariness is not avoided, because it remains in the choice of formula for the weighting, for which there is no obvious 'natural' formula. The examples show several popular measures of rangesize rarity applied to published records of British dragonflies among 10 km grid cells (Merritt, R., Moore, N. W. and Eversham, B. C. 1996, Atlas of the dragonflies of Britain and Ireland, HMSO, London.) (range sizes here are measured just within Britain, although global range size could be used). These give increasing weight to the species with the narrowest ranges from left to right (affecting the scorefrequency histograms in the upper row), using a difference measure, inverse measure, inversesquare measure, and an inverseminimumrange measure. Red cells have high rarity scores and blue cells have low rarity scores (in the top row the score classes on the colour scale are arranged for equal intervals of score values, whereas in the lower row they are arranged for equal numbers of grid cells) (below):
A further treatment that is sometimes used is to divide the inverse rangesize rarity measure by species richness, as a measure of the mean rarity among the species of a grid cell (ref 5). This must be interpreted with care (see also ref 14). It might also be more appropriate for scores such as these with a highly skewed frequency distribution of values to use the median rather than the mean of the inverse range sizes, although the results of comparisons among cells would be broadly similar. A different example below shows the application of mean rarity from work with the Flora Neotropica data for five plant families (Dichapetalaceae, Lecythidaceae, Caryocaraceae, Chrysobalanaceae and Proteaceae genus Panopsis) among onedegree grid cells, with red for high scores and blue for low scores (the mean rarity scores have been smoothed by neighbourhood averaging) (below):
