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apical disc of standard holasteroid structure, with four gonopores;
frontal groove present from ambitus to peristome as shallow furrow; usually also extending adapically;
plastron plating meridosternous with sternal plate similar in size to episternal plates and with episternals biserially offset. In some taxa episternals reaching opposite adradial suture making plastron metasternous;
pores in anterior ambulacrum small and differentiated from those in the paired ambulacra aborally.
Range
Cretaceous (Hauterivian to Maastrichtian); northern hemisphere.