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1b. Ambulacral pore pairs forming a single irregular column on oral surface; in some forming three rows adapically. |
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2a.Ambulacra with outer accessory plate large and in contact with adradial suture. Ambulacral zones almost as wide as the interambulacral zones at the ambitus. |
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3b. Buccal notches and gills present. Primary ambulacral element in each triad always divided so as to create an inner column of imperforate perradial plates. |
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4b. No such membranous gaps; interambulacral plates fully abutting. |
| 5a.Primary tubercles consistently crenulate. |
5b. Primary tubercles non-crenulate or predominantly non-crenulate. Go to 6 |
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6a.Large primary tubercles developed orally in interambulacra along adradial margin and interradially, and also present in ambulacral zones. |
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7a. Ambulacral plates on peristome arranged quadriserially in each zone. Genital pores open in a membranous gap, and do not pierce the genital plates.
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7b. Ambulacral plates on peristome arranged biserially in each zone. Gonopores open within the genital plates. |
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8a. Ophicephalous pedicellariae present. Membranous gaps between interambulacral plates only obvious on the oral surface. |
8b. Ophicephalous pedicellariae absent. Membranous gaps in interambulacral zones present on both oral and apical surfaces. |