The Echinoid Directory

Key to the major clades of Palaeozoic echinoid (1)

1a. Test composed entirely of ambulacral plates, with a perradial column of imperforate plates flanked by one or more columns of perforate plates. Plates always thick and sutures vertical.
Bothriocidaridae

1b. Test with imperforate series of interambulacral plates separating ambulacral zones. Plates commonly thin and imbricate, though they can be thick and tesselate.

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2a. Test with perradial column of imperforate plates in addition to paired perforate ambulacral plate columns and paired interambulacral plate columns.
Bromidechinus

2b. Test composed of alternating zones of perforate ambulacral plate columns and imperforate interambulacral plate columns.

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3a. Single column of interambulacral plates alternates with ambulacral zones.
Cravenechinidae

3b. At least three columns and up to 30 columns of interambulacral plates in each zone.

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4a. Marked difference in ambulacral structure and pore morphology between oral and aboral surfaces. Oral pore-pairs enlarged circular structures with prominent peripodial ring.
Proterocidaridae
4b. Ambulacra not expanded adorally and pore pairs small and undifferentiated, morphologically identical on oral and aboral surfaces.
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5a. Interambulacral plates thick and hexagonal, with vertical sutures and with a fine uniform covering of granules.

Palaechinidae

5b . Interambulacral plates relatively thin and clearly imbricate with oblique sutures; almost always bearing distinct tubercles.

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