Characters and data matrix supporting this cladogram
Apical disc
1. Ocular and genital plates form a contiguous ring in adults: no (0); yes (1)
2. Genital plates subdivided into a mass of small platelets: no (0); yes (1)
3. Ocular plates very elongate and extended: no (0); yes (1).
4. Gonopores open within genital plate (0); open in distal membranous gap (1)
5. periproctal plates: contiguous pavement (0); separated and embedded in membrane (1)
Ambulacra
6. Ambulacral zones: 2/3 or so width of interambulacra at subambitus (0); almost as wide or wider than interambulara at subambitus (1)
7. Smaller elements reduced to minute, ovate accessory plates: no (0); yes one is (1); yes both are (2).
8. On oral surface ambulacral plates occluded from adradial suture: no (0); inner plate occluded (1); both elements occluded (2).
9. On aboral surface primary element occluded from the adradial suture: no (0); yes (1)
10. On oral surface triads heterogeneous and tending to form multicomponent elements: no (0); yes (1).
11. Primary element divided medially into two pieces: no (0); yes (1).
12. On oral surface, pore-pairs marginal (0); forming a band removed from the adradial suture (1)
13. Pore-pairs adorally forming a broad marginal phyllode; no (0); yes (1).
14. Only one accessory plate on ambital and adoral elements: no (0); yes
15. All pores/tube-feet equally developed (0); some pore-pairs reduced (1)
16. Pore-pairs in three discrete columns at ambitus and aborally: no (0); yes (1).
Peristome
17. Buccal plates present: no (0); yes (1).
18. Plates arranged uniserially (0); biserially (1) in each half ambulacrum.
19. Buccal notches: absent (0); feeble (1); well developed (2).
Interambulacra
20. Membranous gaps present orally in interambulacral zones: no (0); yes (1).
21. Membranous gaps present adapically in interambulacral zones: no (0); yes (1).
Tuberculation
22. Primary tubercles: non-crenulate (0); crenulate (1).
23. Areole not sunken (0); sunken (1).
24. Oral tubercles much larger than aboral: no (0); yes (1).
25. Multiple equal tubercles on interambulacral plates: no (0); yes (1).
26. Distinct adradial column of tubercles: no (0); yes (1).
Spines and pedicellariae
27. Oral spines end: in rounded point with fleshy sac (0); in hyaline hoof (1); in flattened blade (2).
28. Sphaeridia: on both accessory plates (0); on inner plate only (1).
29. Tridentate pedicellariae: open blade-like valves with contiguous edges (0);
30. Tridentate pwedicellariae with involute neck and wide lateral gap (1)
30. Ophicephalous pedicellariae: absent (0); present (1).
31. Dactylous pedicellariae: absent (0); present (1).
Teeth
32. CLNP/LLNO system differentiated: no (0); yes (1).
