Eggs are usually laid close to water on objects such as stones or vegetation.
The larvae usually develop in soil where they prey on a variety of other invertebrates.
The life cycle of some species can take up to three years.
The pupal stage is short and the strong-flying adults soon emerge.
Males are vegetarians feeding mainly on nectar.
Females need a blood meal for the development of eggs.
Most species adults only live for approximately one month