Prof Rony Huys

Dr Rony Huys
  • Researcher
  • Life Sciences department
  • Invertebrates
Natural History Museum
Cromwell Road
London
SW7 5BD

Biography

Employment history

2010 -                   Visiting Professor, Hanyang University, Seoul

1994 - present    Crustacea Researcher  (Band 3), Department of Zoology, NHM

1992 - 1994        Senior Scientific Officer, Department of Zoology, NHM

1988 - 1992        EU Research Associate, Delta Institute for Hydrobiological Research (NL)

1987 - 1988        Research Assistant, University of Gent (Belgium)

1984 - 1986        Research Assistant, University of Gent (Belgium)

1988 - 2002        Visiting Research Fellow, State University of Gent (Belgium)

1992 - 1996        Visiting Scientist, Netherlands Institute of Ecological Research

 

Qualifications

1995                    PhD cum laude, University of Gent (Belgium)

1984                    MSc, Zoology, University of Gent (Belgium)

Professional

Awards

1992

  • Royal Academy of Sciences and Arts of Belgium: Biannual Academy Prize for Biological Sciences

External comittees/associations

2011 - present   World Association of Copepodologists – President

2011                    Eleventh International Conference on Copepoda 2011, Mexico – Member of                              International Organizing Committee

2008 - 2011       World Association of Copepodologists – Vice-President

2002                    Eighth International Conference on Copepoda 2002, Taiwan – Member of                              International Organizing Committee

1999 - 2002        World Association of Copepodologists – Member of Executive Council

2000                     Associate Professorship/Curator of Crustacea at Zoological Museum -                                          University of Copenhagen – External member of Assessment Committee

 

World Association of Copepodologists 

Editorial Boards

  • 1998 - present     Zoological Journal of the Linnean Society – Associate Editor
Zoological Journal of the Linnean Society
  • 2001 - present     Meiofauna Marina – Member of Editorial Board
Meiofauna marina
  • 2002 - present     Cahiers de Biologie marine – Member of Editorial Board
Cahiers de Biologie marine
  • 2003 - 2004         Korean Journal of Environmental Biology – Member of Editorial Board
  • 2004 - present     The European Register of Marine Species – Associate Editor (Harpacticoida)
The European Register of Marine Species
  • 2007 - 2009     Organisms, Diversity and Evolution – Editor-in-Chief
Organisms, Diversity and Evolution

External collaborations

Prof. Wonchoel Lee - Hanyang University, Seoul, Korea

Prof. John B. J. Wells, University of Wellington, New Zealand

Prof. Susumu Ohtsuka - Hiroshima University, Japan

Dr John D. D. Bishop - Marine Biological Association, U.K.

Dr Diana M. P. Galassi - University of L'Aquila, Italy

Dr Süphan Karaytuğ - University of Mersin, Turkey

Dr Serdar Sak - University of Balikesir, Turkey

Dr Terue Kihara - DZMB, Forschungsinstitut Senckenberg, Wilhelmshaven, Germany

 

Symposia organized

2011

Eleventh International Conference on Copepoda, 10-15 July, Mérida, Mexico: Organiser of Symposium on Molecules versus Morphology - Why can't they be friends?

Eleventh International Conference on Copepoda, 10-15 July, Mérida, Mexico: Organiser of Symposium Symbiotic Copepods - a Celebration of the Life and Career of Ju-shey Ho

2005

Sixth International Crustacea Congress, 18-22 July, Glasgow, U.K.: Organiser of Symposium on Phylogeny of Crustacea.

2004

Twelfth International Meiofauna Conference, 11-16 July, Ravenna, Italy: Chairman of Systematics and Evolution Theme.

2002

 Eighth International Copepod Conference, 21-26 July, Keelung, Taiwan: Organiser of Symposium on Molecules vs Morphology: sorting the factual and the artefactual.

1998

Tenth International Meiofauna Conference, 27-31 July, Plymouth, U.K.: Chairman of Systematics and Phylogenetics Theme.

1995

Ninth International Meiofauna Conference, 3-7 July, Perpignan, France: Organiser of Symposium on Heterochrony and the Evolution of Meiofauna.

 

Training courses organized

2011

  • Pre-Conference Training Workshop on Morphology and Systematics of Copepods, 4-8 July, Chetumal, Quintana Roo, Mexico

Guest lectures

2011

  • Eleventh International Conference on Copepoda, 10-15 July, Mérida, Mexico
    • Integrating molecules and morphology: consensus or conflict in the symbiotic copepods?

2009

  • The Society for Integrative and Comparative Biology Annual Meeting, 3–7 January, Boston, U.S.A.
    • What can SSU rDNA do for copepod phylogeny and classification? (with J. Llewellyn-Hughes)
  • The Crustacean Society Summer Meeting and 47th Annual Meeting of the Carcinological Society of Japan, 20–24 September, Tokyo, Japan.
    • The evolution of parasitism in the Copepoda – new insights from 18S rDNA data (with J. Llewellyn-Hughes)
  • Darwin, Evolution and Life – The 3rd International Symposium of the National Institute of Biological Resources, 09 October, Incheon, Korea.
    • The evolution of parasitism in the Copepoda – new insights and future challenges

2008

  • Advances in Crustacean Phylogenetics, 7–11 October, Rostock, Germany.
    • Maxillopodan conundrum: insights from SSU rDNA and Bayesian inference (with J. Llewellyn-Hughes)

2005

  • Sixth International Crustacea Congress, 18-22 July, Glasgow, U.K.
    • Position of Tantulocarida and Mystacocarida as inferred from SSU rDNA data (with P.D. Olson & J. Llewellyn-Hughes)

2002

  • Eighth International Conference on Copepoda, 21-26 July, Keelung, Taiwan.
    • Ordinal relationships in the Copepoda: molecules versus morphology (with J. Llewellyn-Hughes and J. Spinks)

1999

  • Copepoda Diversity in the Neotropis, 21-22 July, São Sebastião, Brazil.
    • Marine Harpacticoida from South and Central America
  • Seventh International Conference on Copepoda, 25-31 July, Curitiba, Brazil.
    •  Heterochrony in copepods: pattern, process and significance
    • Sexual dimorphism - kingpin of harpacticoid evolution
    • Sexual dimorphism in calanoid copepods - morphology and function (with S. Ohtsuka)

1997

  • The Levant as a Biogeographic Bridge - Land, Sea and Air, 22-24 June, Jerusalem, Israel.
    • Evolutionary history of Canuellidae (Copepoda: Harpacticoida) and their radiation in the Levant

1995

  • Ninth International Meiofauna Conference, 3-7 July, Perpignan, France.
    • Systematics and evolution of the Cylindropsyllidae: a synthesis

1993

  • Phylogeny of and within the Maxillopoda (ASZ and TCS Winter Meeting), 27 December, Los Angeles, U.S.A
    • Tantulocaridan relationships and the significance of gonopore position and penile homology in maxillopodan phylogeny

Other principal seminars/lectures (last 15 years)          

2009

  • University of Adana, 23 March, Adana, Turkey;  University of the Philippines in the Visayas, 28 September, Miag-ao, Philippines; and, Hanyang University, 07 October, Seoul. Korea
    • Copepoda: an Introduction to the Insects of the Sea
  • Hanyang University, 08 October, Seoul, Korea.
    • The evolution of parasitism in the Copepoda

2008

  • Pre-conference workshop. Systematics, Morphology and Techniques for Identification of Copepods and their Larval Stages, 7-11 July, Khon Kaen, Thailand.
    • Marine benthic and meiofaunal harpacticoid copepods
    • Integrating morphology and molecular sequence data: some examples from the symbiotic Copepoda
  • Tenth International Conference on Copepoda, 13-19 July, Pattaya,Thailand.
    • Cyclopoids in different guises: testing the validity of the copepod order Thaumatopsylloida using morphology and small subunit rDNA (with J. Mackenzie-Dodds & J. Llewellyn-Hughes)

2007

  • Thirteenth International Meiofauna Conference, 11-16 July, Recife, Brazil.
    • Harpacticoids and the transition from the sea to freshwater habitats: some examples elucidated by SSU rDNA and Bayesian inference (with J. Llewellyn-Hughes & J. Mackenzie-Dodds)

2005

  • Sixth International Crustacea Congress, 18-22 July, Glasgow, U.K.
    • Phylogeny and zoogeography of the Tachidiidae Boeck, 1865 (Copepoda, Harpacticoida) (with S. Conroy-Dalton)
  • World Subterranean Biodiversity, 8-10 December, Villeurbanne, France.
    • Ectinosomatidae (Copepoda, Harpacticoida) in groundwater: radiation or multiple invasions (with D.M.P. Galassi & P. De Laurentiis)

2004 

  • Twelfth International Meiofauna Conference, 11-16 July, Ravenna, Italy.
    • Phylogeny and zoogeography of the Tachidiidae Boeck, 1865 (Copepoda: Harpacticoida) (with S. Conroy-Dalton)

2003

  • Korean Copepoda Workshop, Hanyang University, 10-11 July, Seoul, Korea.
    • Marine benthic and meiofaunal copepods
    • Parasitic copepods: an overview
  • The Crustacean Society Mid Year Meeting, 1-5 June, Williamsburg VA, U.S.A.
    • The Rhizocephala and the Tantulocarida: a comparison of life cycles and the infestation process (with J.T. Høeg, H. Glenner, N. Møbjerg & R.M. Kristensen)

2002

  • Pre-conference Workshop on Copepoda, 15-19 July, Kenting, Taiwan.
    • Marine benthic and meiofaunal copepods
    • Copepod phylogeny: morphology and molecular methods
  • Eighth International Conference on Copepoda, 21-26 July, Keelung, Taiwan.
    • Caudal ramus development in Copepoda (with S. Conroy-Dalton)

2001

  • Fifth International Crustacea Congress, 9-13 July, Melbourne, Australia.
    • Copepod Evolution: ten years on (with J. Llewellyn-Hughes & P. Foster)
    • Character transformation and polyphyly in the Ancorabolidae (Copepoda: Harpacticoida) (with S. Conroy-Dalton)

1999

  • Seventh International Conference on Copepoda, 25-31 July, Curitiba, Brazil.
    • New insights in ancorabolid phylogeny (with S. Conroy-Dalton)
    • Taxonomy of Oncaeidae (Copepoda, Poecilostomatoida) from the Red Sea (with R. Böttger-Schnack)

1998

  • Tenth International Meiofauna Conference, 27-31 July, Plymouth, U.K.
    • Evolution and ecological radiation of Darcythompsoniidae (with J.M. Gee)

1996

  • Sixth International Conference on Copepoda, 29 July-3 August, Oldenburg, Germany.
    • The taxonomic status of the deepwater Ameiridae and the case of disproportionate sex ratios in the deep-sea harpacticoids
    • The copepod antennule: its ontogeny and phylogeny (with G.A. Boxshall)
    • First data on hydrothermal vent harpacticoids (with S. Conroy-Dalton)
    • Species groups within the genus Oncaea (Copepoda, Poecilostomatoida) (with R. Böttger-Schnack)

Research

Interests

Copepods are the dominant metazoan group in the marine plankton, are extremely abundant in marine and freshwater sediments and are parasites on virtually every phylum of animals from sponges to chordates. The main theme of my research is the systematics and comparative anatomy of free-living and parasitic copepods, and the application of phylogenetic reconstruction to examine their evolution and ecological radiation, using morphology and molecular markers. Copepods are one of the best models to study fundamental phenomena like the evolution of parasitism and the marine-freshwater transition, and to test fundamental hypotheses such as the claim of oligomerization being the predominant mode of character transformation in Crustacea, and the enemy release hypothesis in invasion ecology.

Schematic representation of the primary habitat of each copepod order

Schematic representation of the primary habitat of each copepod order © Huys & Boxshall (1991)

Despite its hyperabundance and vast diversity, the phylogenetic position of the Copepoda within the Crustacea is still unresolved. Copepoda has traditionally been placed in the Maxillopoda which is widely regarded as the most controversial class of the Crustacea. Molecular analyses in particular have thus far failed to provide strong support for a monophyletic Maxillopoda or identified it as a paraphyletic taxon. Other workers have argued that there is merit to recognition of the Maxillopoda as a natural assemblage, despite the fact that there seem to be exceptions to every synapomorphy proposed. I have developed an interest in examining the relationships of the lesser known and molecularly undersampled maxillopodan lineages such as the Mystacocarida, Pentastomida, Branchiura and Tantulocarida. It is expected that the new (morphological and molecular) insights gained from these studies will contribute to a better understanding of the relationships of the Copepoda.

Projects

My research programme consists of four interrelated projects:

(1) Phylogenetic Relationships of Maxillopoda

Aim. To analyze the phylogenetic relationships between the various groups of the Maxillopoda, one of the six classes currently recognized within the Crustacea, by employing morphological and molecular data across all lineages.

(2) Systematics and evolutionary biology of harpacticoid copepods

Aim. To carry out revisionary work on various families of benthic copepods which will be used as the basis for a synthetic account of the systematics and phylogeny of the Harpacticoida.

SEM micrograph of cephalic region of Arenotopa sp. (Leptastacidae)

SEM micrograph of cephalic region of Arenotopa sp. (Leptastacidae). Photo by R. Huys

(3) Evolution of parasitism in Copepoda

Aims. To carry out morphological and molecular studies on symbiotic copepods in order to determine the cladogenetic events that gave rise to the main lineages and to identify the key innovations associated with them such as the origin of mesoparasitism and dwarf males; to place incertae sedis known thus far from juvenile stages; to generate a new ordinal phylogeny of the Copepoda.

Terebellides stroemi infested by 3 ovigerous females of highly modified copepod Melinnacheres

Terebellides stroemi (Polychaeta) infested by three ovigerous females of highly modified copepod Melinnacheres © M.O. MacNaughton

(4) Copepods and the transition from marine to freshwater habitats: a model for studying oligomerization

Aims. To use molecular data to analyze relationships within various freshwater lineages showing distinct progressive reduction in body morphology; to use the molecular trees to test the importance of oligomerization in copepod evolution and the widely accepted irreversibility of character transformations in this group.

Phylogenetic relationship of semiterrestrial Cancrincolidae - symbionts of landcrabs

Phylogenetic relationship of semiterrestrial Cancrincolidae - symbionts of landcrabs © Huys et al. (2009) Molecular Phylogenetics and Evolution, 51: 143–156.

Books

Copepod Evolution

HUYS, R. & G.A. Boxshall (1991). Copepod Evolution. The Ray Society, London. pp. 468.

 

 

HUYS, R., J.M. Gee, C.G. Moore & R. Hamond (1996).  Marine and Brackish Water Harpacticoid Copepods: Part I. Keys and notes for the identification of genera. Synopses of the British Fauna (New Series). Published for the Linnean Society of London and the Estuarine and Coastal Sciences Association. i-vii, 1-352.

 

Field Studies Council

 

Synopses of the British Fauna (New Series)

Other publications

Most publications are available as PDFs upon request

2010
  • Back, J.,  R. HUYS & W. Lee ( 2010 ) A new species of the genus Tegastes (Copepoda: Harpacticoida: Tegastidae) from hydrothermal vents in the Okinawa Trough. Zoological Science 27 : 678-688 . 10.2108/zsj.27.678
  • El Nagar, A., R. HUYS & J.D.D. Bishop ( 2010 ) Widespread occurrence of the Southern Hemisphere ascidian Corella eumyota Traustedt, 1882 on the Atlantic coast of Iberia. Aquatic Invasions 5 : 169-173 . Aquatic Invasions | doi: 10.3391/ai.2010.5.2.06
  • HUYS, R. & T.C. Kihara ( 2010 ) Systematics and phylogeny of Cristacoxidae (Copepoda, Harpacticoida): a review Zootaxa 2568 : 1-38 . Zootaxa
  • Selden, P.A., R. HUYS, M.H. Stephenson, A.P. Heward & P.N. Taylor ( 2010 ) Crustaceans from bitumen clast in Carboniferous glacial diamictite extend fossil record of copepods Nature Communications 1(50) : 1-6 . Nature Communications | doi:10.1038/ncomms1049
  • Veit-Köhler, G., M. De Troch, M. Grego, W. Bonne, G. De Smet, C. Folkers, K.H. George, C. Guotong, R. Herman, R. HUYS, N. Lampadariou, J. Laudien, P. Martínez Arbizu, A. Rose, M. Schratzberger, S. Seifried, P., J. Vanaverbeke, E. Vanden Berghe, M. Vincx, B. Vrišer & L. Vandepitte ( 2010 ) Large-scale diversity and biogeography of benthic copepods in European waters. Marine Biology 157 : 1819-1835 . doi: 10.1007/s00227-010-1454-0
2009
  • Galassi, D.M.P., R. HUYS & J.W. Reid ( 2009 ) Diversity, ecology and evolution of groundwater copepods. Freshwater Biology 54 : 691-708 . doi:10.1111/j.1365-2427.2009.02185.x
  • HUYS, R. ( 2009 ) On the junior subjective synonyms of Coullia Hamond, 1973 (Copepoda, Harpacticoida, Laophontidae): an update and key to species and related genera. ZooKeys 5 : 33-40 . ZooKeys | doi:10.3897/zookeys.5.64
  • HUYS, R. ( 2009 ) Unresolved cases of type fixation, synonymy and homonymy in harpacticoid copepod nomenclature (Crustacea: Copepoda). Zootaxa 2183 : 1-99 . Zootaxa
  • HUYS, R. & P.F. Clark ( 2009 ) Case 3467. ZOSIMIDAE Seifried, 2003 (Crustacea, Copepoda, Harpacticoida): proposed emendation of spelling to ZOSIMEIDAE to remove homonymy with ZOSIMINAE Alcock, 1898 (Crustacea, Decapoda, XANTHIDAE). Bulletin of Zoological Nomenclature 66 : 24-29 . Bulletin of Zoological Nomenclature
  • HUYS, R. & W. Lee ( 2009 ) Proposal of Marbefia, gen. n. and Inermiphonte, gen. n., including updated keys to the species of Pseudonychocamptus Lang, 1944 and Paralaophonte Lang, 1948 (Copepoda, Harpacticoida, Laophontidae). ZooKeys 23 : 1-38 . doi:10.3897/zookeys.23.168
  • HUYS, R., J. MacKenzie-Dodds & J. Llewellyn-Hughes ( 2009 ) Cancrincolidae (Copepoda, Harpacticoida) associated with land crabs: a semiterrestrial leaf of the ameirid tree. Molecular Phylogenetics and Evolution 51 : 143-156 . doi:10.1016/j.ympev.2008.12.007
  • Kihara, T.C. & R. HUYS ( 2009 ) A new genus of Ectinosomatidae (Copepoda: Harpacticoida) from sublittoral sediments in Ubatuba, São Paulo State (Brazil), an updated key to genera and notes on Noodtiella Wells, 1965. ZooKeys 17 : 57-88 . doi:10.3897/zookeys.17.202
  • Kihara, T.C. & R. HUYS ( 2009 ) Corrigenda: Kihara TC & Huys R (2009) A new genus of Ectinosomatidae (Copepoda, Harpacticoida) from sublittoral sediments in Ubatuba, São Paulo State (Brazil), an updated key to genera and notes on Noodtiella Wells, 1965. ZooKeys 18 : 181-182 . doi:10.3897/zookeys.18.249
  • Kihara, T.C. & R. HUYS ( 2009 ) Contributions to the taxonomy of the Normanellidae (Copepoda, Harpacticoida): description of a new genus from the Brazilian continental shelf and re-assignment of Pseudocletodes vararensis Scott & Scott, 1893 (ex Nannopodidae). Zootaxa 2233 : 1-38 . Zootaxa
2008
  • Bandera, M.E. & R. HUYS ( 2008 ) Proposal of new genus for Asterocheres mucronipes Stock, 1960 (Copepoda, Siphonostomatoida, Asterocheridae), an associate of the scleractinian coral Astroides calycularis (Pallas, 1766) in the Strait of Gibraltar. Zoological Journal of the Linnean Society 152 : 635-653 . doi:10.1111/j.1096-3642.2008.00375.x
  • HUYS, R. ( 2008 ) Case 3470. Halectinosoma Vervoort, 1962 (Crustacea, Copepoda, Harpacticoida): proposed conservation of usage by designation of Ectinosoma chrystalii Scott & Scott, 1894 as the type species and by giving it precedence over Pararenosetella Lang, 1944. Bulletin of Zoological Nomenclature 65 : 276-281 . Bulletin of Zoological Nomenclature
  • HUYS, R. ( 2008 ) Case 3471. Heterolaophonte Lang, 1948 (Crustacea, Copepoda, Harpacticoida): proposed precedence over Mesolaophonte Nicholls, 1941 and Monolaophonte Nicholls, 1941. Bulletin of Zoological Nomenclature 65 : 282-287 . Bulletin of Zoological Nomenclature
  • HUYS, R. & F.-H. Mu ( 2008 ) Description of a new species of Onychostenhelia Itô (Copepoda, Harpacticoida, Miraciidae) from the Bohai Sea, China. Zootaxa 1706 : 51-68 . Zootaxa
  • Sak, R., S. Karaytuğ & R. HUYS ( 2008 ) A review of Pseudoleptomesochrella Lang, 1965 (Copepoda, Harpacticoida, Ameiridae), including a redescription of P. halophila (Noodt, 1952) from the Black Sea and a key to species. Zootaxa 1758 : 45-60 . Zootaxa
  • Sak, R., S. Karaytuğ & R. HUYS ( 2008 ) Ciplakastacus gen. nov., a new primitive genus of Leptastacidae (Copepoda, Harpacticoida) from the Mediterranean coast of Turkey. Journal of Natural History 42 : 2443-2459 . doi:10.1080/00222930802277632
  • Sak, S., R. HUYS & S. Karaytuğ ( 2008 ) Disentangling the subgeneric division of Arenopontia Kunz, 1937: resurrection of Psammoleptastacus Pennak, 1942, re-examination of Neoleptastacus spinicaudatus Nicholls, 1945 and proposal of two new genera and a new generic classification (Copepoda, Harpacticoida, Arenopontiidae). Zoological Journal of the Linnean Society 152 : 409-458 . doi:10.1111/j.1096-3642.2007.00373.x
  • Willems, W.R., M. Curini-Galletti, T.J. Ferrero, D. Fontaneto, I. Heiner, R. HUYS, V.N. Ivanenko, R.M. Kristensen, T. Kånneby, M.O. MacNaughton, P. Martínez Arbizu, M.A. Todaro, W. Sterrer & U. Jondelius ( 2008 ) Meiofauna of the Koster-area, results from a workshop at the Sven Lovén Centre for Marine Sciences (Tjärnö, Sweden). Meiofauna Marina 17 : 1-34 .
2007
  • HUYS, R. & R. Böttger-Schnack ( 2007 ) Taxonomy of Oncaeidae (Copepoda, Cyclopoida) from the Red Sea. - VIII. Morphology and phylogenetic position of Oncaea tregoubovi Shmeleva, 1968 and the closely related O. prendeli Shmeleva, 1966 from the Mediterranean Sea. Mitteilungen aus den hamburgischen zoologischen Museum und Institut 104 : 89-127 .
  • HUYS, R., J. Llewellyn-Hughes, S. Conroy-Dalton, P.D. Olson, J. Spinks & D.A. Johnston ( 2007 ) Extraordinary host switching in siphonostomatoid copepods and the demise of the Monstrilloida: integrating molecular data, ontogeny and antennulary morphology. Molecular Phylogenetics and Evolution 43 : 368-378 . doi:10.1016/j.ympev.2007.02.004
2006
  • HUYS, R. & S. Conroy-Dalton ( 2006 ) Revision of the genus Evansula T. Scott, 1906 (Copepoda, Harpacticoida, Cylindropsyllidae) with a description of three new species. Zoological Journal of the Linnean Society 147 : 419-472 . doi:10.1111/j.1096-3642.2006.00227.x
  • HUYS, R. & S. Conroy-Dalton ( 2006 ) Comparative analysis of vermiform genera (Copepoda: Harpacticoida: Cylindropsyllidae), including descriptions of Boreovermis, new genus and Bolbotelos, new genus, from the southern North Sea. Journal of Crustacean Biology 26 : 206-223 . doi:10.1651/C-2609.1
  • HUYS, R., J. Llewellyn-Hughes, P.D. Olson & K. Nagasawa ( 2006 ) Small subunit rDNA and Bayesian inference reveal Pectenophilus ornatus (Copepoda incertae sedis) as highly transformed Mytilicolidae, and support assignment of Chondracanthidae and Xarifiidae to Lichomolgoidea (Cyclopoida). Biological Journal of the Linnean Society 87 : 403-425 . doi:10.1111/j.1095-8312.2005.00579.x
2005
  • HUYS, R. & S. Conroy-Dalton ( 2005 ) Aquilastacus gen. nov. from the southern North Sea and the taxonomic position of Leptastacus operculatus Masry, 1970 (Copepoda: Harpacticoida: Leptastacidae). Cahiers de Biologie marine 46 : 347-363 . Cahiers de Biologie marine
  • HUYS, R., S. Karaytuğ & V. Cottarelli ( 2005 ) On the synonymy of Delamarella Chappuis and Latiremus Božić (Copepoda, Harpacticoida, Latiremidae), including the description of D. obscura sp. nov. from the Black Sea. Zoological Journal of the Linnean Society 145 : 263-281 . doi:10.1111/j.1096-3642.2005.00188.x
  • HUYS, R., S. Ohtsuka, S. Conroy-Dalton & Y. Kikuchi ( 2005 ) Description of two new species of Neotachidius Shen & Tai, 1963 (Copepoda, Harpacticoida, Tachidiidae) from Korean brackish waters and proposal of a new genus for Tachidius (Tachidius) vicinospinalis Shen & Tai, 1964. Zoological Journal of the Linnean Society 143 : 133-159 . doi:10.1111/j.1096-3642.2004.00148.x
  • Ivanenko, V.N., D. Defaye & R. HUYS ( 2005 ) A new species of Ambilimbus nom. nov., a replacement name for Amphicrossus Huys, 1991 (Copepoda, Cyclopoida, Erebonasteridae), from the Mid-Atlantic Ridge at 36°N (hydrothermal vent site Rainbow). Marine Biology Research 1 : 233-241 . doi:10.1080/17451000510019097
2004
  • Böttger-Schnack, R. & R. HUYS ( 2004 ) Size polymorphism in Oncaea venusta Philippi, 1843 and the validity of O. frosti Heron, 2002: a commentary. Hydrobiologia 513 : 1-5 . doi:10.1023/B:hydr.0000018301.50664.3d
  • HUYS, R. ( 2004 ) Review of Damkaer, D.M. (2002). The copepodologist’s cabinet: a biographical and bibliographical history. Volume 1, Aristotle to Alexander von Nordmann (330 BC to AD 1832). Archives of natural History 31 : 187-188 . doi:10.3366/anh.2004.31.1.187
  • HUYS, R. & S.J. Song ( 2004 ) The Ismardiidae Leigh-Sharpe (Copepoda, Harpacticoida incertae sedis): enigmatic as ever? Journal of Crustacean Biology 24 : 37-53 . doi:10.1651/C-2412
  • Karaytuğ, S. & R. HUYS ( 2004 ) Taxonomic position of and generic distinction between Parepactophanes Kunz, 1935 and Taurocletodes Kunz, 1975 (Copepoda, Canthocamptidae incertae sedis), with description of a new species from the Black Sea. Zoological Journal of the Linnean Society 140 : 469-486 . doi:10.1111/j.1096-3642.2003.00101.x
  • Mu, F.-h. & R. HUYS ( 2004 ) Canuellidae (Copepoda, Harpacticoida) from the Bohai Sea, China. Journal of natural History 38 : 1-36 . doi:10.1080/00222930210138935
2003
2002
  • HUYS, R., P.J. López-González, Á.A. Luque & E. Roldán ( 2002 ) Brooding in cocculiniform limpets (Gastropoda) and familial distinctiveness of the Nucellicolidae (Copepoda): misconceptions reviewed from a chitonophilid perspective. Biological Journal of the Linnean Society 75 : 187-217 . doi:10.1046/j.1095-8312.2002.00010.x
  • Lee, W. & R. HUYS ( 2002 ) A new genus of groundwater Ameiridae (Copepoda: Harpacticoida) from boreholes in Western Australia and the artificial status of Stygonitocrella Petkovski, 1976. Bulletin of the Natural History Museum (Zoology) 68 : 39-50 . doi:10.1017/S0968047002000055
  • Mu, F.-h. & R. HUYS ( 2002 ) New species of Stenhelia (Copepoda, Harpacticoida, Diosaccidae) from the Bohai Sea (China) with notes on subgeneric division and phylogenetic relationships. Cahiers de Biologie marine 43 : 179-206 . Cahiers de Biologie marine
2001
  • Böttger-Schnack, S. & R. HUYS ( 2001 ) Taxonomy of Oncaeidae (Copepoda, Poecilostomatoida) from the Red Sea. III. Morphology and phylogenetic position of Oncaea subtilis Giesbrecht, 1892. In: R.M. Lopes, J.W. Reid & C.E.F. Rocha (eds), Copepoda: Developments in Ecology, Biology and Systematics. Hydrobiologia 453/454 : 467-481 . doi:10.1023/A:1013114706718
  • Coomans, A., R. HUYS, J. Heyns & M. Luc ( 2001 ) Character analysis, phylogeny and biogeography of the genus Xiphinema Cobb, 1913 (Nematoda: Longidoridae). Annales du Musée royal de l'Afrique Central (Sciences zoologiques) 287 : i-iv + 1-240 .
  • HUYS, R. ( 2001 ) Splanchnotrophid systematics: a case of polyphyly amd taxonomic myopia. Journal of Crustacean Biology 21 : 106-156 . doi:10.1651/0278-0372(2001)021[0106:SSACOP]2.0.CO;2
  • HUYS, R. ( 2001 ) Harpacticoida. In: Costello, M.J., C. Emblow & R. White (eds). European register of marine species. A check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels 50 : 268-280 .
  • HUYS, R. & G.A. Boxshall ( 2001 ) An appreciation of the contribution of Arthur Humes to copepod systematics. Journal of Crustacean Biology 21 : 13-27 . doi:10.1651/0278-0372(2001)021[0013:AAOTCO]2.0.CO;2
  • Lee, W. & R. HUYS ( 2001 ) New Aegisthidae (Copepoda: Harpacticoida) from western Pacific cold seeps and hydrothermal vents. [Erratum]. Zoological Journal of the Linnean Society 131 : 249 . doi:10.1111/j.1096-3642.2001.tb01317.x
  • Ohtsuka, S. & R. HUYS ( 2001 ) Sexual dimorphism in calanoid copepods: morphology and function. In: R.M. Lopes, J.W. Reid & C.E.F. Rocha (eds), Copepoda: Developments in Ecology, Biology and Systematics. Hydrobiologia 453/454 : 441-466 . doi:10.1023/A:1013162605809
2000
  • Conroy-Dalton, S. & R. HUYS ( 2000 ) Systematics and phylogeny of the Ancorabolidae (Copepoda, Harpacticoida). I. The Ancorabolus-lineage, with the description of three new genera. Cahiers de Biologie marine 41 : 343-397 . Cahiers de Biologie marine
  • HUYS, R. & S. Conroy-Dalton ( 2000 ) Generic concepts in the Clytemnestridae (Copepoda: Harpacticoida): revision and revival. Bulletin of the Natural History Museum (Zoology) 66 : 1-48 .
  • HUYS, R. & W. Lee ( 2000 ) Basal resolution of laophontid phylogeny and the paraphyly of Esola Edwards. Bulletin of the Natural History Museum (Zoology) 66 : 49-107 .
  • Lee, W. & R. HUYS ( 2000 ) New Aegisthidae (Copepoda: Harpacticoida) from western Pacific cold seeps and hydrothermal vents. Zoological Journal of the Linnean Society 129 : 1-71 . doi:10.1111/j.1096-3642.2000.tb00008.x
  • López-González, P.J., J. Bresciani, R. HUYS, A. F. González, A. Guerra & S. Pascual ( 2000 ) Discovery of Genesis vulcanoctopusi gen. et sp. nov. (Copepoda: Tisbidae) on a hydrothermal vent octopod and a reinterpretation of the life cycle of cholidyinid harpacticoids. Cahiers de Biologie marine 41 : 241-253 . Cahiers de Biologie marine
1999
  • Bouck, L., D. Thistle & R. HUYS ( 1999 ) Systematics and phylogeny of Zausodes C.B. Wilson (Copepoda, Harpacticoida, Harpacticidae), including three new species from the northern Gulf of Mexico. Bulletin of the Natural History Museum (Zoology) 65 : 73-122 .
  • Conroy-Dalton, S. & R. HUYS ( 1999 ) A new genus of Aegisthidae (Copepoda, Harpacticoida) from hydrothermal vents on the Galapagos Rift. Journal of Crustacean Biology 19 : 408-431 . Journal of Crustacean Biology
  • HUYS, R. & W. Lee ( 1999 ) On the relationships of the Normanellidae and the recognition of Cletopsyllidae grad. nov. (Copepoda, Harpacticoida). Zoologischer Anzeiger 237 : 267-290 .
  • Lee, W. & R. HUYS ( 1999 ) New Normanellidae (Copepoda: Harpacticoida) from western Pacific cold seeps including a review of the genus Normanella. Cahiers de Biologie marine 40 : 203-262 . Cahiers de Biologie marine
  • Lee, W. & R. HUYS ( 1999 ) Bathylaophonte gen. nov. from deep-sea hydrothermal vents and the polyphyly of Paronychocamptus (Copepoda: Harpacticoida). Cahiers de Biologie marine 40 : 293-328 . Cahiers de Biologie marine
  • Lee, W. & R. HUYS ( 1999 ) New Tachidiella (Copepoda, Harpacticoida, Tisbidae) from the Antarctic and Norway including a review of the genus. Zoosystema 21 : 419-444 . Zoosystema
1998
  • Boxshall, G.A. & R. HUYS ( 1998 ) The ontogeny and phylogeny of copepod antennules. Philosophical Transactions of the Royal Society of London B353 : 765-786 . doi:10.1098/rstb.1998.0242
  • Böttger-Schnack, R. & R. HUYS ( 1998 ) Species groups within the genus Oncaea Philippi, 1843 (Copepoda, Poecilostomatoida). Journal of marine Systems 15 : 369-371 . doi:10.1016/S0924-7963(97)00072-9
  • Conroy-Dalton, S. & R. HUYS ( 1998 ) Towards a revision of Ameira Boeck, 1865 (Harpacticoida, Ameiridae): reinstatement of Psammameira Noodt, 1952. Zoologica Scripta 27 : 247-261 . doi:10.1111/j.1463-6409.1998.tb00439.x
  • HUYS, R. & T.M. Iliffe ( 1998 ) Novocriniidae, a new family of harpacticoid copepods from anchihaline caves in Belize. Zoologica Scripta 27 : 1-15 . doi:10.1111/j.1463-6409.1998.tb00425.x
  • Todaro, M.A. & R. HUYS ( 1998 ) La meiofauna delle secche della Meloria: osservazioni su un nuovo Leptastacidae (Copepoda, Harpacticoida). Meiofauna from the Meloria Shoals: observations on a new Leptastacidae (Copepoda, Harpacticoida). Biologia Marina Mediterranea 5 : 591-595 .
1997
  • Böttger-Schnack, R. & R. HUYS ( 1997 ) Archioncaea arabica gen. et sp. nov., a remarkable oncaeid (Copepoda: Poecilostomatoida) from the northern Arabian Sea. Cahiers de Biologie marine 38 : 79-89 .
  • Böttger-Schnack, R. & R. HUYS ( 1997 ) Morphological observations on Oncaea mediterranea (Claus, 1863) (Copepoda: Poecilostomatoida) with a comparison of Red Sea and eastern Mediterranean populations. Bulletin of the Natural History Museum (Zoology) 63 : 137-147 .
  • Conroy-Dalton, S. & R. HUYS ( 1997 ) Towards a revision of Ameira Boeck, 1865 (Harpacticoida: Ameiridae): re-examination of the A. tenella-group and the establishment of Filexilia gen. n. and Glabrameira gen. n. Zoologica Scripta 25 : 317-339 . doi:10.1111/j.1463-6409.1996.tb00169.x
  • HUYS, R. & M.A. Todaro ( 1997 ) Meloriastacus ctenidis gen. et sp. nov.: a primitive interstitial copepod (Harpacticoida, Leptastacidae) from Tuscany. Italian Journal of Zoology 64 : 181-196 . doi:10.1080/11250009709356193
  • HUYS, R. & P. Bodin ( 1997 ) First record of Acanthocephala in marine copepods. Ophelia 46 : 217-231 .
  • HUYS, R. & R. Böttger-Schnack ( 1997 ) On the diphyletic origin of the Oncaeidae Giesbrecht, 1892 (Copepoda: Poecilostomatoida) with a phylogenetic analysis of the Lubbockiidae fam. nov. Zoologischer Anzeiger 235 : 243-261 .
  • HUYS, R. & S. Conroy-Dalton ( 1997 ) Discovery of hydrothermal vent Tantulocarida on a new genus of Argestidae (Copepoda: Harpacticoida). Cahiers de Biologie marine 38 : 235-249 .
  • HUYS, R., N. Møbjerg & R.M. Kristensen ( 1997 ) New tantulocarid Polynyapodella ambrosei gen. et sp. nov. (Basipodellidae) from the Northeast Water Polynya (Greenland) with emphasis on the phylogeny of its host genus Cervinia (Copepoda: Harpacticoida). Cahiers de Biologie marine 38 : 181-199 .
1996
  • Gee, J.M. & R. HUYS ( 1996 ) An appraisal of the taxonomic position of Enhydrosoma buchholzi (Boeck, 1872), E. bifurcarostratum Shen & Tai, 1965, E. barnishi Wells, 1967 and E. vervoorti Fiers, 1987 with definition of two new genera (Copepoda, Harpacticoida, Cletodidae). Sarsia 81 : 161-191 . Sarsia
  • HUYS, R. ( 1996 ) Biuncus nom. nov., a replacement name for Singularia Huys, 1995 (Copepoda: Harpacticoida: Paramesochridae). Journal of natural History 30 : 1261 . doi:10.1080/00222939600770691
  • HUYS, R. ( 1996 ) Superornatiremidae fam. nov. (Copepoda: Harpacticoida): an enigmatic family from North Atlantic anchihaline caves. Scientia marina 60 : 497-542 . Scientia marina
  • HUYS, R. & J.M. Gee ( 1996 ) Prionos gen. nov. from the meiofauna of a Malaysian mangrove forest and the status of Psammis borealis (Copepoda, Harpacticoida, Paranannopidae). Cahiers de Biologie marine 37 : 227-248 .
  • HUYS, R. & J.M. Gee ( 1996 ) Sentiropsis, Peltisenia and Afrosenia: three new genera of Paranannopidae (Copepoda: Harpacticoida). Cahiers de Biologie marine 37 : 49-75 .
  • HUYS, R. & S. Conroy-Dalton ( 1996 ) A revision of Leptopontia T. Scott (Copepoda: Harpacticoida) with description of six new species. Zoological Journal of the Linnean Society 118 : 197-239 . doi:10.1111/j.1096-3642.1996.tb01265.x
  • HUYS, R., J.-Y. Bodiou & P. Bodin ( 1996 ) A revision of Psamathea (Harpacticoida: Leptastacidae) with description of P. brittanica sp. nov. Vie et Milieu 46 : 7-19 .
1995
  • HUYS, R. ( 1995 ) Some remarks on the taxonomic status of Paraschizopera Wells, 1981 (Copepoda: Harpacticoida). Hydrobiologia 308 : 23-28 . doi:10.1007/BF00037783
  • HUYS, R. ( 1995 ) A new genus of Paramesochridae (Copepoda: Harpacticoida) from Amphioxus-sand in Elat, Israel. Journal of natural History 29 : 673-684 . doi:10.1080/00222939500770231
  • HUYS, R. ( 1995 ) A new genus of Canuellidae (Copepoda: Harpacticoida) associated with Atlantic bathyal sea urchins. Zoologica Scripta 24 : 225-243 . doi:10.1111/j.1463-6409.1995.tb00401.x
1994
  • Boxshall, G.A. & R. HUYS ( 1994 ) Asterocheres reginae, a new species of parasitic copepod (Siphonostomatoida: Asterocheridae) from a sponge in Belize. Systematic Parasitology 27 : 19-33 . doi:10.1007/BF02185665
  • Gee, J.M. & R. HUYS ( 1994 ) Paranannopidae (Copepoda: Harpacticoida) from sublittoral soft sediments in Spitsbergen. Journal of natural History 28 : 1007-1086 . doi:10.1080/00222939400770541
  • HUYS, R. & R. Böttger-Schnack ( 1994 ) Taxonomy, biology and phylogeny of Miraciidae (Copepoda: Harpacticoida). Sarsia 79 : 207-283 . Sarsia
  • HUYS, R., S. Ohtsuka & G.A. Boxshall ( 1994 ) A new tantulocaridan (Crustacea: Maxillopoda) parasitic on calanoid, harpacticoid and cyclopoid hosts. Publications of the Seto marine biological Laboratory 36 : 197-209 .
1993
  • HUYS, R. ( 1993 ) Styracothoracidae (Copepoda: Harpacticoida), a new family from the Philippine deep sea. Journal of Crustacean Biology 13 : 769-783 . Journal of Crustacean Biology
  • HUYS, R. & J.M. Gee ( 1993 ) A revision of Danielssenia Boeck and Psammis Sars with the establishment of two new genera Archisenia and Bathypsammis (Harpacticoida: Paranannopidae). Bulletin of the Natural History Museum (Zoology) 59 : 45-81 .
  • HUYS, R. & K.A. Willems ( 1993 ) A revision of Cylinula and two new species of Cylindropsyllus (Copepoda, Harpacticoida, Cylindropsyllidae). Zoologica Scripta 22 : 347-362 . doi:10.1111/j.1463-6409.1993.tb00364.x
  • HUYS, R. & S. Conroy-Dalton ( 1993 ) Willemsia gen. nov. and Boreopontia Willems, 1981 revisited (Harpacticoida: Cylindropsyllidae). Sarsia 78 : 273-300 . Sarsia
  • HUYS, R. & S. Ohtsuka ( 1993 ) A new species of Syrticola Willems & Claeys, 1982 (Copepoda: Harpacticoida) from Japan with notes on the type species. Bulletin of the Natural History Museum (Zoology) 59 : 83-94 .
  • HUYS, R., G.A. Boxshall & J.-P. Casanova ( 1993 ) A redescription of Cumoniscus Bonnier, 1903 (Crustacea: Tantulocarida) from a cumacean host in the Bay of Biscay. Systematic Parasitology 26 : 75-79 . doi:10.1007/BF00009651
  • HUYS, R., G.A. Boxshall & R.J. Lincoln ( 1993 ) The tantulocarid life cycle: the circle closed? Journal of Crustacean Biology 13 : 432-442 . Journal of Crustacean Biology
1992
  • Boxshall, G.A. & R. HUYS ( 1992 ) A homage to homology: patterns of copepod evolution. Acta Zoologica 73 : 327-334 .
  • Heip, C., R. HUYS & R. Alkemade ( 1992 ) Community structure and functional roles of meiofauna in the North Sea. Netherlands Journal of Aquatic Ecology 26 : 31-41 .
  • Humes, A.G. & R. HUYS ( 1992 ) Copepoda (Poecilostomatoida and Siphonostomatoida) from deep-sea hydrothermal vent areas off British Columbia, including Amphicrossus altalis, a new species of Erebonasteridae, with notes on the taxonomic position of the genus Tychidion Humes. Canadian Journal of Zoology 70 : 1369-1380 . doi:10.1139/z92-193
  • HUYS, R. ( 1992 ) The amphiatlantic distribution of Leptastacus macronyx (T. Scott, 1892) (Copepoda: Harpacticoida): a paradigm of taxonomic confusion; and, a cladistic approach to the classification of the Leptastacidae Lang, 1948. Mededelingen van de Koninklijke Academie voor Wetenschappen, Letteren en Schone Kunsten van België 54(4) : 21-196 .
  • HUYS, R., G.A. Boxshall & R. Böttger-Schnack ( 1992 ) On the discovery of the male of Mormonilla Giesbrecht, 1891 (Copepoda, Mormonilloida). Bulletin of the British Museum of Natural History (Zoology) 58 : 157-170 .
  • HUYS, R., P.F. Andersen & R.M. Kristensen ( 1992 ) Tantulacus hoegi gen. et sp. nov. (Tantulocarida: Deoterthridae) from the meiobenthos of the Faroe Bank, North Atlantic. Sarsia 76 : 287-297 . Sarsia
  • HUYS, R., P.M.J. Herman, C.H.R. Heip & K. Soetaert ( 1992 ) The meiobenthos of the North Sea: density, biomass trends and distribution of copepod communities. ICES Journal of marine Science 49 : 23-44 . doi:10.1093/icesjms/49.1.23
  • HUYS, R., S. Ohtsuka, G.A. Boxshall & T. Itô ( 1992 ) Itoitantulus misophricola gen. et sp. nov.: First record of Tantulocarida (Crustacea: Maxillopoda) in the North Pacific region. Zoological Science 9 : 875-886 .
  • Ohtsuka, S., R. HUYS, G.A. Boxshall & T. Itô ( 1992 ) Misophriopsis okinawensis sp. nov. (Crustacea: Copepoda) from hyperbenthic waters off Okinawa, South Japan, with definitions of related genera Misophria Boeck, 1864 and Stygomisophria gen. nov. Zoological Science 9 : 859-874 .
1991
  • Gee, J.M. & R. HUYS ( 1991 ) A review of Paranannopidae (Copepoda: Harpacticoida) with claviform aesthetascs on oral appendages. Journal of natural History 25 : 1135-1169 . doi:10.1080/00222939100770741
  • HUYS, R. ( 1991 ) Tantulocarida (Crustacea: Maxillopoda): a new taxon from the temporary meiobenthos. P.S.Z.N.I.: Marine Ecology 12 : 1-34 . doi:10.1111/j.1439-0485.1991.tb00080.x
  • HUYS, R. ( 1991 ) Crustacea Copepoda: Amphicrossus pacificus gen. et sp. nov., an erebonasterid copepod (Poecilostomatoida) from the New Caledonian continental shelf. In: Crosnier, A. (ed.), Résultats des Campagnes MUSORSTOM, Volume 9. Mémoires du Muséum national d’Histoire naturelle, (A) 152 : 63-77 .
  • Smol, N., R. HUYS & M. Vincx ( 1991 ) A 4-years analysis of the meiofauna community of a dumping site for TiO2-waste off the Dutch coast. Chemistry and Ecology 5 : 197-215 . doi:10.1080/02757549108035250
1990
  • Gee, J.M. & R. HUYS ( 1990 ) The rediscovery of Danielssenia intermedia Wells 1965 (Copepoda, Harpacticoida): a missing link between the "danielsseniid" genera and Paranannopus Lang 1936 (Paranannopidae). Journal of natural History 24 : 1549-1571 . doi:10.1080/00222939000770891
  • Heip, C., R. HUYS, M. Vincx, A. Vanreusel, N. Smol, R. Herman & P.M.J. Herman ( 1990 ) Composition, distribution, biomass and production of North Sea meiofauna. Netherlands Journal of Sea Research 26 : 333-342 . doi:10.1016/0077-7579(90)90095-X
  • HUYS, R. ( 1990 ) Campyloxiphos dineti gen. et spec. nov. from off Namibia and a redefinition of the Deoterthridae Boxshall & Lincoln (Crustacea: Tantulocarida). Journal of natural History 24 : 415-432 . doi:10.1080/00222939000770311
  • HUYS, R. ( 1990 ) Adenopleurella, new genus, Proceropes, new genus, Sarsocletodes Wilson (ex Laophontidae) and Miroslavia Apostolov (ex Cletodidae): representatives of a new family (Copepoda: Harpacticoida). Journal of Crustacean Biology 10 : 340-363 . Journal of Crustacean Biology
  • HUYS, R. ( 1990 ) A new harpacticoid copepod family collected from Australian sponges and the status of the subfamily Rhynchothalestrinae Lang. Zoological Journal of the Linnean Society 99 : 51-115 . doi:10.1111/j.1096-3642.1990.tb01222.x
  • HUYS, R. ( 1990 ) Pholenota spatulifera Vervoort (Copepoda: Harpacticoida): aberrant laophontid or specialized diosaccid? Journal of natural History 24 : 635-646 . doi:10.1516/H246-1M40-2R04-2Q33
  • HUYS, R. ( 1990 ) Amsterdam Expeditions to the West Indian Islands, Report 64. A new family of harpacticoid copepods and an analysis of the phylogenetic relationships within the Laophontoidea T. Scott. Bijdragen tot de Dierkunde 60 : 79-120 .
  • HUYS, R. ( 1990 ) Coralliotantulus coomansi gen. et sp. n.: First record of a tantulocaridan (Crustacea: Maxillopoda) from shallow subtidal sands in tropical waters. Stygologia 5 : 183-198 .
  • HUYS, R. ( 1990 ) Allocation of the Mantridae Leigh-Sharpe to the Cyclopoida (Crustacea: Copepoda) with notes on Nearchinotodelphys Ummerkutty. Bijdragen tot de Dierkunde 60 : 283-291 .
  • HUYS, R. & G.A. Boxshall ( 1990 ) The rediscovery of Cyclopicina longifurcata (Scott) (Copepoda: Cyclopinidae) in deep water in the North Atlantic, with a key to genera of the subfamily Cyclopininae. Sarsia 75 : 17-32 . Sarsia
  • HUYS, R. & J.M. Gee ( 1990 ) A revision of Thompsonulidae Lang, 1944 (Copepoda: Harpacticoida). Zoological Journal of the Linnean Society 99 : 1-49 . doi:10.1111/j.1096-3642.1990.tb01221.x
  • HUYS, R. & G.A. Boxshall ( 1990 ) Discovery of Centobnaster humesi, new genus, new species (Erebonasteridae), the most primitive poecilostomatoid copepod known, in New Caledonian deep waters. Journal of Crustacean Biology 10 : 504-519 . Journal of Crustacean Biology
  • HUYS, R., C.H.R. Heip, P.M.J. Herman ( 1990 ) The meiobenthos of the North Sea: preliminary results of the North Sea Benthos Survey. ICES Committee Meeting Papers and Reports C.M. 1990/Mini: 8 : 1-19 .
1989
  • Boxshall, G.A. & R. HUYS ( 1989 ) New tantulocarid, Stygotantulus stocki, parasitic on harpacticoid copepods, with an analysis of the phylogenetic relationships within the Maxillopoda. Journal of Crustacean Biology 9 : 126-140 . Journal of Crustacean Biology
  • Boxshall, G.A. & R. HUYS ( 1989 ) A new family of deep-sea planktonic copepods, the Paralubbockiidae (Copepoda: Poecilostomatoida). Biological Oceanography 6 : 163-173 .
  • Boxshall, G.A., R. HUYS & R.J. Lincoln ( 1989 ) A new species of Microdajus (Crustacea: Tantulocarida) parasitic on a tanaid in the northeastern Atlantic, with observations on M. langi Greve. Systematic Parasitology 14 : 17-30 . Systematic Parasitology
  • HUYS, R. ( 1989 ) Dicrotrichura tricincta gen. et spec. nov.: a new tantulocaridan (Crustacea: Maxillopoda) from the Mediterranean deep waters off Corsica. Bijdragen tot de Dierkunde 59 : 243-249 .
  • HUYS, R. & D. Thistle ( 1989 ) Bathycamptus eckmani gen. et spec. nov. (Copepoda, Harpacticoida) with a review of the taxonomic status of certain other deepwater harpacticoids. Hydrobiologia 185 : 101-126 . doi:10.1007/BF00010809
  • HUYS, R. & K.A. Willems ( 1989 ) Laophontopsis Sars and the taxonomic concept of the Normanellinae (Copepoda: Harpacticoida): a revision. Bijdragen tot de Dierkunde 59 : 203-227 .
  • HUYS, R. & A. Coomans ( 1989 ) Echinoderes higginsi sp.n. (Kinorhyncha, Cyclorhagida) from the southern North Sea with a key to the genus Echinoderes Claparède. Zoologica Scripta 18 : 211-221 . doi:10.1111/j.1463-6409.1989.tb00446.x
1988
  • Heip, C., R.M. Warwick, M.R. Carr, P.M.J. Herman, R. HUYS, N. Smol & K. Van Holsbeke ( 1988 ) Analysis of community attributes of the benthic meiofauna of Frierfjord/Langesundfjord. Marine Ecology Progress Series 46 : 171-180 . Marine Ecology Progress Series
  • HUYS, R. ( 1988 ) Studies on the Cylindropsyllidae (Copepoda, Harpacticoida). 2. Revision of the genus Cylindropsyllus Brady. Zoologica Scripta 17 : 253-276 . doi:10.1111/j.1463-6409.1988.tb00100.x
  • HUYS, R. ( 1988 ) A redescription of the presumed associated Caligopsyllus primus Kunz, 1975 (Harpacticoida, Paramesochridae) with emphasis on its phylogenetic affinity with Apodopsyllus Kunz, 1962. Hydrobiologia 162 : 3-19 . doi:10.1007/BF00014330
  • HUYS, R. ( 1988 ) Gelyelloida, a new order of stygobiont copepods from European karstic systems. Hydrobiologia 167/168 : 485-495 . doi:10.1007/BF00026343
  • HUYS, R. ( 1988 ) Stygofauna of the Canary Islands, 10. Rotundiclipeidae fam. nov. (Copepoda, Harpacticoida) from an anchihaline cave on Tenerife, Canary Islands. Stygologia 4 : 42-63 .
  • HUYS, R. ( 1988 ) Sexual dimorphism in aegisthid cephalosomic appendages (Copepoda, Harpacticoida): a reappraisal. Bijdragen tot de Dierkunde 58 : 114-136 .
  • HUYS, R. ( 1988 ) On the identity of the Namakosiramiidae Ho & Perkins 1977 (Crustacea, Copepoda), including a review of harpacticoid associates of Echinodermata. Journal of natural History 22 : 1517-1532 . doi:10.1080/00222938800770941
  • HUYS, R. ( 1988 ) Stygofauna of the Canary Islands, 11. Boxshallia bulbantennulata gen. et spec. nov. (Copepoda: Misophrioida) from an anchihaline lava pool on Lanzarote, Canary Islands.
    Stygologia 4 : 138-154 .
  • HUYS, R. & G.A. Boxshall ( 1988 ) A new genus and species of tantulocaridan (Crustacea: Tantulocarida) parasitic on a harpacticoid copepod from the Skagerrak. Sarsia 73 : 205-211 . Sarsia
  • HUYS, R. & H. Kunz ( 1988 ) On the generic boundaries within the marine interstitial Latiremidae (Copepoda: Harpacticoida). Stygologia 4 : 292-305 .
1987
  • HUYS, R. ( 1987 ) Paramesochra T. Scott, 1892 (Copepoda, Harpacticoida): a revised key, including a new species from the SW Dutch coast and some remarks on the phylogeny of the Paramesochridae. Hydrobiologia 144 : 193-210 . doi:10.1007/BF00005554
  • HUYS, R. ( 1987 ) Studies on the Cylindropsyllidae (Copepoda, Harpacticoida). 1. The status of Leptastacus laticaudatus Nicholls. Zoologica Scripta 16 : 155-166 . doi:10.1111/j.1463-6409.1987.tb00062.x
  • HUYS, R. ( 1987 ) Some morphological observations on the Neobradyidae Olofsson, 1917 (Copepoda, Harpacticoida) including the redescription of Antarcticobradya tenuis (Brady, 1910) comb. nov.
    Bulletin de l’Institut royal des Sciences naturelles de Belgique, Biologie 57 : 133-148 .
1986
  • HUYS, R., R.L. Herman & C. Heip ( 1986 ) Seasonal fluctuations in vertical distribution and breeding activity of a subtidal harpacticoid community in the Southern Bight, North Sea. Netherlands Journal of Sea Research 20 : 375-383 . doi:10.1016/0077-7579(86)90004-9
1985
  • Heip, C., O. Hamerlynck, P. Herman, R. HUYS, M. Vincx & G. Vranken ( 1985 ) Meiobenthos in the Southern Bight of the North Sea. In: Van Grieken, R. & R. Wollast (eds), Progress in Belgian Oceanographic Research, Antwerp: 264-266 .
  • HUYS, R. & R. Herman ( 1985 ) Seasonal fluctuations in vertical distribution and breeding activity of a subtidal harpacticoid community in the Southern Bight, North Sea. Some preliminary results. In: Van Grieken, R. & R. Wollast (eds), Progress in Belgian Oceanographic Research, Antwerp: 279-291 .
  • Martens, M., P. Jouk, R. HUYS & R. Herman ( 1985 ) A short note on the relative abundance of the Turbellaria in the meiofauna of sandy habitats in the Southern Bight of the North Sea and on Belgian beaches. In: Van Grieken, R. & R. Wollast (eds), Progress in Belgian Oceanographic Research, Antwerp: 341-342 .