Dr Paul Eggleton

Paul Eggleton
  • Merit Researcher, Entomology
  • Life Sciences department
  • Insects
Natural History Museum
Cromwell Road

Brief Biography

Employment history

Researcher and Head, Soil Biodiversity Group,  1992-present


BSc (Hons), Bristol University, 1985

PhD, Imperial College, 1990


Dr Eggleton's research group, the Soil Biodiversity Group (SBG) investigates patterns of diversity in soil macrofauna, their environmental correlates, and their relationship with ecological processes.

 At present the SBG is undertaking the following projects:

  • A large scale study of biodiversity patterns in the New Forest, southern England, particularly examining very diverse groups (e.g. soil and litter beetles, lichens) and their interaction with their environments and land management history.
  •  Studies of the interaction between ants and termites in tropical rain forests employing both spatial ecological and DNA techniques to pinpoint where ants are feeding on termites, where they are and which termites are being eaten. 
  • Systematic studies of the Termitidae, including developing a new subfamily level classification and taxonomic studies of the soldierless termites of Africa and South America
  • Studies of the biogeography and global functional structure of termites, continuing to explore the diversity anomaly of the enormously higher taxonomic and functional diversity of termites in tropical Africa than in other tropical regions. 
  • Studies of the role of termites in ecosystem processes, including developing a tropics-wide network of wood decomposition experiments, centred on a PhD project (Cecilia Dahljso) in Peru.
  • Descriptive ecology and natural history of UK earthworms (in conjunction with the Earthworm Society of Britain and OPAL) . 


Earthworm Society of Britain







Palin, OF, Eggleton, P, Malhi, Y, Girardin, CAJ, Rozas-Dávila, Parr, CL, 2011. Termite diversity along an Amazon-Andes elevation gradient, Peru. Biotropica, 43: 100-107.


Eggleton, P. 2011. An introduction to termites: biology, taxonomy and functional morphology. In, Bignell, DE, Roisin, Y & Lo, N. (eds) Biology of Termites: A Modern Synthesis. Springer, Dordrecht, pp. 1-26.


Lo, N & Eggleton, P. 2011. Termite phylogenetics and co-cladogenesis with symbionts. In, Bignell, DE, Roisin, Y & Lo, N. (eds) Biology of Termites: A Modern Synthesis. Springer, Dordrecht, pp. 27-50.


Jones, DT & Eggleton, P. 2011. Global biogeography of termites: a compilation of sources. In, Bignell, DE, Roisin, Y & Lo, N. (eds) Biology of Termites: A Modern Synthesis. Springer, Dordrecht, pp.  477-498.


Bone, J, Head, M, Jones, DT, Barraclough, D, Archer, M, Scheib, C, Flight, D, Eggleton, P & Voulvoulis, N. 2010. From chemical risk assessment to environmental quality management: the challenge for soil protection. Environmental Science & Technology, pub?.


Krell, F-K, Ross, E, Giusti, A, Krell-Westerwalbesloh, S, & Eggleton, P. Comparison of leaf litter and litter plus upper soil extractions in assessing forest floor macroinvetebrate fauna. Entomologist’s Monthly Magazine, 146, 71-82.


Fayle, TM, Turner, EC, Chey, VK, Chung, AYC, Eggleton, P, & Foster, WA. 2010. Oil palm expansion into rain forest greatly reduces ant biodiversity in canopy, epiphytes and leaf-litter. Basic & Applied Ecology, 11: 337-345.


Nobre, T, Eggleton, P., & Aanen, DK. 2010. Vertical transmission as the key to the colonization of Madagascar by fungus-growing termites? Proceeding of the Royal Society, series B, 277: 359-365


Fayle, TM, Chung, AYC, Dumbrell, AJ, Eggleton, P, Foster, WA. 2009. The effect of rain forest canopy architecture on the distribution of epiphytic ferns (Asplenium spp.) in Sabah, Malaysia. Biotropica, 41: 676-681.


Monaghan, MT,Wild, R, Elliot, M, Fujisawa, T, Balke, M, Inward, DJ, Lees, DC, Ranaivosolo, R, Eggleton, P, Barraclough, TG, Vogler, AP. 2009. Systematic Biology, 58: 298- 311.


Cornwell, WK, Cornelissen, JHC, Allison, SD, Bauhus, J, Eggleton, P, Preston, CM, Scarff, F, Weedon, JT, Wirth, C, Zanne, AE. Plant traits and wood fates across the globe: rotted, burned, or consumed? Global Change Biology, 15: 2431-2449.


Eggleton, P, Inward, K, Smith, J, Jones, DT, Sherlock, E. 2009. A six year study of earthworm (Lumbricidae) populations in pasture woodland in southern England shows their responses to soil temperature and soil moisture. Soil Biology and Biochemistry, 41: 1857-1861.


Raheem, DC,  Naggs, F,  Chimonides, PDJ, Preece, RC and Eggleton, P. 2009. Fragmentation and pre-existing species turnover determine land-snail assemblages of tropical rain forest. Journal of Biogeography, 36: 1923 – 1938.  


Smith , J, Potts, SG, Woodcock, BA and Eggleton, P. 2009. The impact of two arable field margin management schemes on litter decomposition. Applied Soil Ecology, 41: 90-97.  


Carpenter, D; Hodson, ME; Eggleton, P, Kirk, C. 2008. .The role of earthworm communities in soil mineral weathering: a field experiment . Mineralogical Magazine   72: 33-36


Scholtz, O., Macleod, N., Eggleton, P. 2008. Termite soldier defence strategies: a reassessment of Prestwich’s classification using extended eigenshapes analyses of head capsule morphology. Zoological Journal of the Linnean Society, 153: 631-650.


Smith, J, Potts, S, Eggleton, P. 2008.  Evaluating the efficiency of sampling methods in assessing soil macrofauna communities in arable systems. European Journal of Soil Biology, 44: 271-276.


Smith, J, Potts, S, Eggleton, P. 2008. The value of sown grass margins for enhancing soil macrofaunal biodiversity in arable systems. Agriculture, Ecosytems and Environment, 127: 119-125


Raheem, D, Naggs, F., Preece, R., Mapatuna, Y., Kariyawasam, L. &  Eggleton, P. 2008. The Structure of the Land-snail Assemblages of Fragmented Lowland Rainforest and Village Home Gardens in Southern Sri Lanka. Journal of Applied Ecology, 45: 1019-1028


Smith, J., Potts, S., Woodcock, B,  Eggleton, P. 2008. Can arable field margins be managed to enhance their biodiversity, conservation and functional value for soil macrofauna? Journal of Applied Ecology. 45:  269–278.


Vanbergen, A.J., Watt, A.D., Mitchell, R., Truscott, A.M., Palmer, S.C.F., Ivits, E., Eggleton, P., Jones, T.H. & Sousa, J.P. 2007. Scale-specific correlations between habitat heterogeneity and soil fauna diversity along a landscape structure gradient. Oecologia, 153: 713-725.


Carpenter, D., Hodson, M.E., Eggleton, P. & Kirk, C. 2007. Earthworm induced mineral weathering: preliminary results. European Journal of Soil Biology, 43: S176-S183.


Inward, D.J.G., Vogler, A.P. & Eggleton, P. 2007. A comprehensive phylogenetic analysis of termites (Isoptera) illuminates key aspects of their evolutionary biology. Molecular Phylogenetics and Evolution, 44: 953-967.


Eggleton, P., Beccaloni, G. & Inward, D. 2007. Response to Lo et al.: “Save Isoptera” Biology Letters, 3: 564-565.


Inward, D. Beccaloni, G. & Eggleton, P. 2007. Death of an order: a comprehensive molecular phylogenetic study confirms that termites are eusocial cockroaches. Biology Letters, 3: 331-335.


Smith, J., Chapman, A. & Eggleton, P. 2006. Baseline biodiversity surveys of the soil macrofauna of London’s green spaces. Urban Ecosystems, 9: 337-349.


Jones, D.T., Davies, R.G. & Eggleton, P. 2006. Sampling termites in forest habitats: a reply to Roisin and Leponce. Austral Ecology, 31: 429-431.


Nobre, T., Nunes, L., Eggleton, P. & Bignell, D.E. 2006. Distribution and genetic variation of Reticulitermes (Isoptera: Rhinotermitidae) in Portugal. Heredity, 96: 403-409.


Eggleton, P. 2006. The termite gut habitat: its evolution and co-evolution. In Intestinal Microrganisms of Termites and Other Invertebrates (H. König & A. Varma, eds). Springer-Verlag, Berlin. Pp. 373-404.


Aanen, D.K. & Eggleton, P. 2005. Fungus-growing termites originated in African rain forests. Current Biology, 15: 851-855.


Eggleton, P., Vanbergen, A.J., Jones, D.T., Lambert, M.C., Rockett, C., Hammond, P.M., Beccaloni, J., Marriott, D., Ross, E. & Giusti, A. 2005. Assemblages of soil macrofauna across a Scottish land-use intensification gradient: influences of habitat quality, heterogeneity and area. Journal of Applied Ecology, 42: 1153-1164.


Krell, F-T., Chung, A.Y.C., Ross, E., Eggleton, P., Giusti, A., Inward, K. & Krell-Westerwalbesloh, S. 2005. Quantitative extraction of macro-invertebrates from temperate and tropical leaf-litter and soil: efficiency and time-dependent taxonomic biases of the Winkler extraction. Pedobiologia, 49: 175-186.


Donovan, S.E., Purdy, K.J., Kane, M.D. & Eggleton, P. 2004. Comparison of Euryarchaea strains in the guts and food-soil of the soil-feeding termite Cubitermes fungifaber across different soil-types. Applied and Environmental Microbiology, 70: 3884-3892.


Jones, D.T., Verkerk, R.H.J, & Eggleton, P. 2004. Methods for sampling termites. In Insect Sampling in Forest Ecosystems (S. Leather, ed.). Blackwell, Oxford. Pp. 221-253.


Davies, R.G., Eggleton, P., Jones, D.T., Gathorne-Hardy, F.J. & Hernandez, L.M.  2003. Evolution of termite functional diversity: analysis and synthesis of local ecological and regional influences on local species richness. Journal of Biogeography, 30: 847-877.


Davies, R.G., Hernandez, L., Eggleton, P., Didham, R.K., Fagan, L. & Winchester, N.  2003. Assemblage composition responses of termites to fragmentation at a land-bridge island site in French Guiana. Journal of Tropical Ecology, 19: 509-524.


Jones, D.T., Susilo, F.X., Bignell, D.E., Hardiwinoto, S., Gillison, A.N. & Eggleton, P. 2003. Termite assemblage collapse along a land-use intensification gradient in lowland central Sumatra, Indonesia. Journal of Applied Ecology, 40: 380-391.


Eggleton, P. & Davies, R.G. 2003. Isoptera, Termites. In The Natural History of Madagascar (S.M. Goodman & J.P. Benstead eds). University of Chicago Press. Pp 654-660.


Krell, F-T., Krell-Westerwalbesloh, S., Weiß, I., Eggleton, P., & Linsenmair, K.E. 2003. Spatial separation of Afrotropical dung beetle guilds: a trade-off between competitive superiority and energetic constraints (Coleoptera: Scarabaeidae). Ecography, 26: 210-222.


Aanen, D.K., Eggleton, P., Rouland-Lefèvre C., Guldberg-Frøslev, T., Rosendahl, S. & Boomsma, J.J. 2002. The evolution of fungus-growing termites and their mutualistic fungal symbionts. Proceedings of the National Academy of Science, 99: 14887-14892. 


Gathorne-Hardy, F. J., Davies, R.G., Eggleton, P., Jones, D.T. & Syaukani. 2002. Quaternary rainforest refugia in south-east Asia: using termites (Isoptera) as indicators. Biological Journal of the Linnean Society, 75: 453-466.


Eggleton, P., Bignell, D.E., Hauser, S., Dibog, L., Norgrove, L. & Madong, B. 2002. Termite diversity across an anthropogenic disturbance gradient in the humid forest zone of West Africa. Agriculture, Ecosystems and Environment, 90: 189-202.


Davies, R.G. 2002. Feeding group responses of a Neotropical termite assemblage to rain forest fragmentation. Oecologia, 133: 233-242.


Donovan, S.E., Eggleton, P., & Martin, A. 2002. Species composition of termites of the Nyika plateau forests, northern Malawi, over an altitudinal gradient. African Journal of Ecology, 40: 379-385.


Eggleton, P., Davies, R.G., Connetable, S., Bignell, D.E. & Rouland, C. 2002. The termites of the Mayombe Forest Reserve, Congo (Brazzaville): transect sampling reveals an extremely high diversity of ground-nesting soil feeders. Journal of Natural History, 36: 1239-1246.


Chung, A.Y.C., Eggleton, P., Speight, M.R., Hammond, P.M. & Chey, V.K. 2002. A survey on defoliation and phytophagous insects in four habitat types in Sabah, Malaysia. Journal of Tropical Forest Science 14: 116-130.


Brauman, A., Doré, J., Breznak, J., Eggleton, P., Bignell, D.E. & Kane, M.D. 2001. Molecular phylogenetic profiling of prokaryotic communities in termites with different feeding habits. FEMS Microbiology Ecology, 35: 27-36.


Gathorne-Hardy, F., Syaukani & Eggleton, P. 2001. The effect of altitude and rainfall on the composition of termites  (Isoptera) of the Leuser Ecosystem (Sumatra, Indonesia). Journal of Tropical Ecology, 17: 379-393.


Donovan, S.E., Eggleton, P. & Bignell, D.E. 2001. Gut content analysis and a new feeding group classification of termites (Isoptera). Ecological Entomology, 26: 356-366.


Donovan, S.E., Eggleton, P., Dubbin, W.E., Batchelder, M. & Dibog, L. 2001. The effect of a soil-feeding termite, Cubitermes fungifaber (Isoptera: Termitidae) on soil properties: termites may be an important source of soil microhabitat heterogeneity in tropical forests. Pedobiologia, 45: 1-11.


Eggleton, P. 2001. Termites and trees: a review of recent advances in termite phylogenetics. Insectes Sociaux, 48: 187-193.


Eggleton, P. & Tayasu, I. 2001. Feeding groups, lifetypes and the global ecology of termites. Ecological Research, 16: 941-960.


Chung, Y.C., Speight, M.R., Eggleton, P., Hammond, P.M. & Chey, V.K. 2001. Impact of simulated defoliation on selected commercially important tree species in Sabah, Malaysia. The Malaysian Forester, 63: 113-125.


Kambhampati, S. & Eggleton, P. 2000. Phylogenetics and Taxonomy. In: Abe, T., Bignell, D. E. & Higashi, M. (eds.),Termites: Evolution, Sociality, Symbiosis, Ecology. Kluwer Academic Publishers. Pp 1-23.


Eggleton, P. 2000. Global diversity patterns. In: Abe, T., Bignell, D. E. & Higashi, M. (eds.), Termites: Evolution, Sociality, Symbiosis, Ecology. Kluwer Academic Publishers. Pp. 25-51.


Bignell, D.E. & Eggleton, P. 2000. Termites in Ecosystems. In: Abe, T., Bignell, D. E. & Higashi, M. (eds.),Termites: Evolution, Sociality, Symbiosis, Ecology. Kluwer Academic Publishers. Pp. 363-387.


Donovan, S.E., Jones, D.T., Sands, W.A. & Eggleton, P. 2000. Morphological phylogenetics of termites. Biological Journal of the Linnean Society, 70: 467-513


Jones, D.T. & Eggleton, P. 2000. Sampling termite assemblages in tropical forests: testing a rapid biodiversity assessment protocol. Journal of Applied Ecology, 37: 191-203.


Chung, A.Y.C., Eggleton, P., Speight, M.R., Hammond, P.M. & Chey, V.K. 2000. The diversity of beetle assemblages in different habitat types in Sabah, Malaysia. Bulletin of Entomological Research, 90: 475-496.


Chung, A.Y.C., Chey, V.K., Eggleton, P., Hammond, P.M. & Speight, M.R. 2000. Variation in beetle (Coleoptera) diversity at different heights of tree canopy in a native forest and forest plantation in Sabah, Malaysia. Journal of Tropical Forest Science, 13: 369-385


Gathorne-Hardy, F., Collins, N.M., Buxton, R.D. & Eggleton, P. 2000. A faunistic review of the termites (Insecta: Isoptera) of Sulawesi, including an updated checklist of the species. Malayan Nature Journal, 54: 347-353.


Chung, A.Y.C., Hammond, P.M., Eggleton, P., Speight, M.R. & Chey, V.K. 2000. A general survey of the Staphylinidae (Insecta: Coleoptera) assemblage in Sabah, Malaysia. Malayan Nature Journal, 54: 355-368.


Chung, A.Y.C, Eggleton, P., Hammond, P.M., Speight, M.R. & Chey, V.K. 2000. Beetle (Coleoptera) sampling using the Winkler method, flight-interception-trapping and mist-blowing in Sabah. Malaysia. Sabah Parks Nature Journal, 3: 63-78.


Eggleton, P., Homathevi, R., Jones, D.T., MacDonald, J., Jeeva, D., Bignell, D.E., Davies, R.G. & Maryati, M. 1999. Termite assemblages, forest disturbance, and greenhouse gas fluxes in Sabah, East Malaysia. Philosophical Transactions of the Royal Society, series B, 354: 1791-1802.


Davies, R.G., Eggleton, P., Dibog, L., Lawton, J.H., Bignell, D.E., Brauman, A., Hartmann, C., Nunes, L., Holt, J. & Rouland, C. 1999. Successional response of a tropical forest termite assemblage to experimental habitat perturbation. Journal of Applied Ecology, 36: 946-962.


MacDonald, J., Eggleton, P., Jeeva, D., Bignell, D.E., Lawton, J.H., Davies, R.G. & Maryati, M. 1999. The effect of termites and disturbance on the CH4 budget in tropical forests in Cameroon and Borneo. Global Change Biology, 5: 869-881.


Eggleton, P. 1999. Termite species description rates and the state of termite taxonomy. Insectes Sociaux, 46: 1-5.


Jeeva, D., Bignell, D.E., Eggleton, P. & Marayti, M. 1999. Respiratory gas exchanges of termites from the Sabah (Borneo) assemblage. Physiological Entomology, 24: 11-17.


Dibog, L., Eggleton, P., Norgrove, L., Bignell, D.E. & Hauser, S. 1999. Effects of canopy retention on the abundance and species richness of termites in an agrisilvicultural system in southern Cameroon. Bulletin of Entomological Research, 89: 125-132.


Eggleton, P., Davies, R.G., Kane, M.D. & Kambhampati, S. 1999. A checklist of termites from Kaieteur National Park (Guyana). Proceedings of the Entomological Society of Washington, 101: 687-689.


Lawton, J.H., Bignell, D.E., Bolton, B., Bloemers, G., Eggleton, P., Hammond, P.M., Hodda, M., Holt, R.D., Larsen, T.B., Mawdsley, N.A., Stork, N.E., Srivastava, D.S. & Watt, A.D. 1998. Biodiversity inventories, indicator taxa and effects of habitat modification in a Cameroon tropical forest. Nature, 391: 72-76.  


Eggleton, P., Davies, R.G. & Bignell, D.E. 1998. Body size and energy use in termites (Isoptera): the responses of soil-feeders and wood-feeders differ in a tropical forest assemblage. Oikos, 81: 525-530.


MacDonald, J.A., Eggleton, P., Bignell, D.E. & Forzi, F. 1998. Methane emission by termites and oxidation by soils across a forest disturbance gradient in the Mbalmayo Forest Reserve, Cameroon. Global Change Biology, 4: 409-418.


Didham, R.K., Lawton, J.H., Hammond, P.M., Eggleton, P. & Stork, N.E. 1998. Beetle species responses to tropical forest fragmentation. Ecological Monographs, 68: 295-333


Bignell, D.E. & Eggleton, P. 1998. Termites. In, (Callow, P., ed.) The Encyclopedia of Ecology and Environment Management, pp. 744-746.


Dibog, L., Eggleton, P. & Forzi, F. 1998. The seasonality of a termite assemblage in a humid tropical forest, Mbalmayo, southern Cameroon. Journal of Tropical Ecology, 14: 841-850.


Brauman, A., Doré, J., Eggleton, P., Bignell, D.E. & Kane, M.D. 1998. Analyse phylogénetique des communautés microbienne de tube digestif de termites à regimes alimentaires différenciés. Microorganismes Aaerobies, (ed. J.L. Tholozan). 1998, Acte du Collogue Société Francais de Microbiologie. 245-355.


Eggleton, P. & Bignell, D.E. 1997. The incidence of secondary occupation of epigeal termite (Isoptera) mounds by other termites in the Mbalmayo Forest Reserve, southern Cameroon, and its biological significance. Journal of African Zoology, 111: 489-498.


Tayasu, H., Abe, T., Eggleton, P. & Bignell, D.E. 1997. Nitrogen and carbon isotope ratios in termites (Isoptera): an indicator of trophic habit along the gradient from wood-feeding to soil-feeding. Ecological Entomology, 21: 279-288.


Watt, A., Stork, N.E., Eggleton, P. & Srivastava, D., Larsen, T.B., Brendell, M.J.D. & Bignell, D.E. 1997. Impact of forest loss and regeneration on insect abundance and diversity. In Insects and Forests (eds. Watt, A., Stork, N.E., & Hunter, M.). Chapman & Hall, London, pp. 273-286.


Bignell, D.E., Eggleton, P., Thomas, K. & Nunes, L. 1997. Termites as mediators of carbon fluxes in tropical forest: budgets for carbon dioxide and methane emissions. In Insects and Forests (eds. Watt, A., Stork, N.E. & Hunter, M.). Chapman & Hall, London, pp. 119-134.


Nunes, L., Bignell, D.E., Lo, N. & Eggleton, P. 1997. On the respiratory quotient (RQ) of termites (Insecta: Isoptera). Journal of Insect Physiology, 43: 749-758.


Eggleton, P., Homathevi, R., Jeeva, D., Jones, D.T., Davies, R.G. & Maryati, M. 1997. The species richness and composition of termites (Isoptera) in primary and regenerating lowland dipterocarp forest in Sabah, east Malaysia. Ecotropica, 3:119-128.


Eggleton, P., Bignell, D.E., Sands, W.A., Mawdsley, N.A., Lawton, J.H., Wood, T.G. & Bignell, N.C. 1996. The diversity, abundance and biomass of termites under differing levels of disturbance in the Mbalmayo Forest Reserve, southern Cameroon. Philosophical Transactions of the Royal Society of London, series B, 351: 51-68.


Lawton, J.H., Bignell, D.E., Bloemers, G.F., Eggleton, P. & Hodda, M.E. 1996. Carbon flux and diversity of nematodes and termites in Cameroon forest soils. Biodiversity and Conservation, 5: 261-273. 


Sleaford, F., Bignell, D.E. & Eggleton, P. 1996. A pilot analysis of gut contents in termites from the Mbalmayo Forest Reserve, Cameroon. Ecological Entomology, 21: 279-288.


Eggleton, P., Bignell, D.E., Sands, W.A., Waite, B. & Wood, T.G. 1995. The species richness of termites (Isoptera) under differing levels of forest disturbance in the Mbalmayo Forest Reserve, southern Cameroon. Journal of Tropical Ecology, 11: 85-98.


Bignell, D.E. & Eggleton, P. 1995. On the elevated gut pH of higher termites (Isoptera: Termitidae). Insectes Sociaux, 42: 57-69.


Gaston, KJ, Williams, PH, Eggleton, P, & Humphries, CJ. Large scale patterns of biodiversity: spatial variation in family richness. Proceeding of the Royal Society: Biological Sciences, 260: 149-154.


Eggleton, P. & Bignell, D.E. 1995. Monitoring the response of tropical insects to changes in the environment: troubles with termites. In, Insects in a changing environment (R. Harrington & N.E. Stork, eds), pp. 473-497. Academic Press.


Eggleton, P. & Vane‑Wright, R.I. 1994. Some principles of phylogenetics and their consequences for comparative biology. In, Phylogenetics and Ecology (P. Eggleton & R.I. Vane‑Wright, eds). Academic Press, London. Pp. 345-366.


Eggleton, P., Williams, P.H. & Gaston, K.J. 1994. Explaining termite global richness: productivity or history? Biodiversity and Conservation, 3: 318-330.


Eggleton, P. 1994. Termites live in pear‑shaped world: a response to Platnick. Journal of Natural History, 28: 1209-1212.


Eggleton, P. & Belshaw R. 1993. Differences between Diptera, Hymenoptera and Coleoptera parasitoids: provisional phylogenetic expla­nations. Biological Journal of the Linnean Society, 48: 213‑226.


Stork N.E. & Eggleton P. 1992. Invertebrates as determinants and indicators of soil quality. American Journal of Alternative Agriculture, 7: 38‑47.


Eggleton, P. & Gaston, K.J. 1992. Tachinid host ranges: a reappraisal (Diptera: Tachinidae). Entomologist's Gazette, 43: 139‑143.


Eggleton, P.. & Belshaw R.  1992. Insect parasitoids: an evolutionary overview. Philosophical Transactions of the Royal Society of London, series B, 331: 1‑20.


Eggleton, P. 1991. Patterns in male mating strategies of the Rhyssinae (Hymenoptera: Ichneumonidae). Animal Behaviour, 41: 829‑837.


Eggleton, P. & Gaston K.J. 1990. Parasitoid species and assemblages: convenient definitions or misleading compromises? Oikos, 59: 417‑421


Eggleton, P. 1990. The male reproductive strategy of Lytarmes maculipennis (Hymenoptera: Ichneumonidae). Ecological Entomology, 15: 357‑360.