Dr Paul Clark

Biography

Employment history

2008-present     Crustacea Research Group

2001-2008         Grade 4, Decapoda Worker and Contract Support

1992-2001         Grade 4, Curation Manager, Inverts II

1990-1992         Senior Scientific Officer

1986-1990         Higher Scientific Officer

1978-1985         Scientific Officer

1974-1977         Assistant Scientific Officer

1974                   Joined the Crustacea Section,
                             Department of Zoology, Natural History Museum, London

1973-1974         Computer Operator,
                             Seismograph Services Ltd (UK)

1968-1969         Laboratory Assistant, Chemotheraphy,
                              Wellcome Research Laboratories              

Academic qualifications

2007                   PhD

1980                   MSc Modern Taxonomy

1973                   BSc (General) Physiology & Zoology

MSc Thesis

Clark, PF 1980. British spider crabs of the genus Inachus; a morphological study of larval development. Polytechnic of Central London/City of London Polytechnic, MSc. Modern Taxonomy (CNAA) pp. 1-140, figs 1-29, pls 1-14. 

PhD Thesis

Clark, PF 2007. Xanthoid crab (Crustacea: Decapoda:Brachyura) classification and phylogeny: an appraisal using the chaetotaxy of first stage zoeae. Thesis pp1-473; Appendix I Larval Descriptions pp 1-227; Appendix II Larval Figures Acantholobulus bermudensis to Lophozozymus pictor pp 1-308; Appendix III Larval Figures Lybia plumosa to Zozymodes xanthoides pp 309-590. University Marine Station Millport, University of London.

Professional Roles

Honorary position

Honorary Research Associate of the Raffles Museum, Singapore, November 2002.

Editorial Boards

• Pakistan Journal of Marine Science, Editorial Advisor since 2000.
• Aquatic Invasions, Editorial Board since November 2005.
• Marine Biodiversity Records, Editor-in-Chief since January 2009.

Learned Societies

• Honorary Treasurer of the Ray Society 2006-present
• Council member of Tetrapods 
• Ray Society council member 2000
• European Governor, Crustacean Society. 1999 and 2000

Scientific conference committees

• 9^th Colloquium 2008, Crustacea Decapoda Mediterranea, Torino, Italia.
• 7^th Colloquium 1999, Crustacea Decapoda Mediterranea, Lisbon, Portugal.
• 6^th Colloquium 1996, Crustacea Decapoda Mediterranea, Florence, Italy.

Organisation of Conferences

• 6^th International Crustacean Congress, Glasgow, 2005, organiser of the 4^th Crustacean Larval Conference, Invasive Crustacea co-organiser with Liz Cook.
• 4^th International Crustacean Congress, Amsterdam, 1998, Invasive Crustacea, co-organiser with Pere Noël and Darryl Felder.
• 2nd European Crustacean Conference, Liege, Belgium, 1996. 1^st Larval Conference.
• Underwater Association, Annual Symposium convener 1981-86, London, UK

Invited speaker at conferences

• Advances in Decapod Crustacean Phylogenetics, San Antonio, Texas, 2008. Xanthid crab (Crustacea: Decapoda: Brachyura) classification and phylogeny: an appraisal using the setotaxy of first stage zoeas and with Jose Cuesta, The bearing of larval morphology on decapod phylogeny. Funded by NSF Tree of Life Programme.
• Shellfish Association of Great Britain, Annual Conference London, 2007, Chinese mitten crab and chips. Funded by Association.
• Symposium on Biodiversity, Holy Name University, Tagbilaran City, Bohol, Philippines. 2002. Funded. Invited speaker.
• Tweed Invasives 2, Kelso, Scotland, 2001, The Chinese Mitten Crab. Funded by Tweed Forum.
• An Introduction to Molluscs, Curation, Conservation and Uses, 2000, Oxford University Museum of Natural History. Invited speaker; Museum Glassware.

Contributions to Workshops

Benthic Marine Invertebrate Identification Workshop including free-living marine nematodes; polychaetes, peracarid Crustaceans and Decapoda. Department of BCB, University of Western Cape, Cape Town, South Africa: 2007. Prepared 5 talks for decapod workshop and presented a seminar on Thames mitten crabs. Workshop funded by The Royal Society.

PhDs examined

• Universitat de Barcelona, 2008
• University of Singapore, 2008
• University of Karachi, 2007

Societies

• Fellow of the Zoological Society of London
• Fellow of the Linnean Society of London
• Member of the Biological Society of Washington
• Member of the Crustacean Society
• Member of the Japanese Carcinological Society
• Member of the Brazilian Crustacean Society

Fieldwork

Diving qualifications:

• Commenced training, 10/10/72
• BSAC Snorkel Diver, 28/04/74
• BSAC 3rd Class Diver, 11/07/74
• BSAC 2nd Class Diver, 09/08/77
• BSAC Club Instructor, 10/12/77
• BSAC Advanced Instructor, 20/09/88
• Brevet de Moniteur International de Plongee, 1 star, M1 GB 425 issued BSAC London
• Brevet de Moniteur International de Plongee, 2 star, M2 GB 327 issued BSAC London
• Brevet International de Plongee Scientifique, PS GB O69 issued 15/07/81
• HSE Certificate of Competence, issued by the Secretary, British Museum (Natural History), 26/11/81
• Diver Training Certificate; PartIV/367/88, Appropriate training and/or experience verified by the Health and Safety Executive, 01/06/88
• Certificate of Diving First Aid, Certificate number 2993, Date of issue 1/10/91

Diving courses attended:

• Chartwork, 13/10/74, BSAC
• Boathandling, 23/2/76, BSAC
• Sub Aqua Bronze Medallion, 22/2/76, BSAC
• Compressor operations & Management, 22/4/78, BSAC
• Dry Suit Course, 2/85, BSAC
• VHF Radio, 20/2/86, Naval Club, London; RYA Day Skipper, 26/5/87, Kingsdale School, London
• Advanced Instructor Course, 4-5/7/87, BSAC, Fort, Bovisand
• First Aid for Divers; 7/2/88, BSAC
• Oxygen Administration, 8/5/88, B06/2, Fort Bovisand
• RYA Diver Cox'n, 13/5/89, BO9/7, Fort Bovisand

Diving experience:

• July 1973, Diving Expedition to Illes Medas, L'Estartit, Espana, Portsmouth Polytechnic
• 1974, Assisted Dr R.W. Ingle & Mr Wyn Thomas in Crayfish, Austropotamobuis palipes, mark and recapture experiments, River Darenth, Kent, England
• August 1974, Diving Expedition to Holy Island, West Coast of Scotland Assisting Dr R.W. Ingle in decapod studies
• August 1974, Diving Expedition to Tunisia Marine survey, Polytechnic of Central London
• BM(NH) field work, West Coast of Ireland, Falmouth, Brixham, Thames Estuary, Swanage
• April 1980, Sinai Peninsular, Red Sea Underwater Photography
• April 1985, Sinai Peninsular, Red Sea Underwater Photography
• July 1987, Ischia, Italy Decapod larval studies on Posidonia beds
• Sep.-Nov. 1987, BM(NH) Contract Work, North Norfolk Coast
• Nov. 1988-to date, BM(NH) Contract Work, North Norfolk Coast
• December 1988, Red Sea Underwater Photography
• December 1989, Cayman Brac Underwater Photography

Diving posts held on societies:

• West Wickham BSAC Branch 533 (Member since 1973, Instructor, Diving Officer Nov. 86-Nov. 88)
• Underwater Association (Council member 1979-1991, Symposium convener 1978-86)
• British Society of Underwater Photographers (Committee Member 1979-1981)

Photographic competitions:

Winner of BSOUP "On the Day" trophy, May 1989.

NHM diving contracts:

• Royal Society and Soviet Acadamy of Sciences; jointly with G. Boxshall; travel support grant for field work diving in Lake Biakal, 1990 1991& 1993.
• North Norfolk coast survey for Long Sewage outfall. 1998, 
• Environmental impact of the wrecking of the Million Hope on the coral reef at Al Fawz, South Sinai, Egypt. July 1996, Thanet subtidal survey, August 1997.
• Marlin on Sha’b Ashrâfi, North of Hurgharda, Gulf of Suez, Egypt, May 1998.
• Cliffe Pools Lagoon Invertebrate Survey, 1998. Fleet Lagoon, 1999.
• Normandy Farm and Acre Lagoons, Lymington and Portland Harbour, 2000

Land Rover Experience:

• Level I Familiarisation Course 13^th February 1996
• Level II Specialist Course 28^th July 1997
• Winching Course 16^th May 2002

Fieldwork:

Fundaçáo Universidade of Rio Grande, Departamento of Oceanografia, Laboratório de Crustáceos Decápodos, Brazil: Libinia spinosa with Danilo Calazans and Suelma S. Rodrigues, February - March 1996, Darwin Inititive.

European research project, INCO-DC no. IC18-CT96-0127, "Macrobenthos of Eastern African mangroves: life cycles and reproductive biology of exploited species" to work on Inhaca Island, Moçambique. Septemeber 1997. Collaboration with José Paula (Principle Investigator), IMAR-Guia Marine Lab, Faculty of Sciences of the University of Lisbon.

Diversity and distribution of marine benthic biotopes in the Laguna San Rafael National Park, Chile. Darwin Initiative Funding in Collaboration with Operation Raleigh. February 1999.

Coastal Biodiversity in Ranong, Thailand. Objective To develop and promote awareness and teaching of tropical marine biodiversity in Thailand, thus empowering the local community, students, conservationists to conserve fragile marine habitats, such as seagrass meadows and mangroves.

Project collaboration between Kasesart University, Bangkok, Wildlife Fund Thailand Kampaun Small Fisheries Co-operative and the NHM and PML. Project co-ordinator in Thailand: Dr Chittima Arythaka, Kasetsart University. Funding: Project supported By European Commission Directorate General for Development, with funds from the institutes concerned. 4-16 September 2000.

Publications

Full bibliography PDF (94.1 KB)

Most recent peer-reviewed publications

2011

Lai J, Mendoza J C, Guinot D, CLARK P F, Ng P K L (2011) Xanthidae MacLeay, 1838 (Decapoda: Brachyura: Xanthoidea) systematics: a multi-gene approach with support from adult and zoeal morphology. Zoologischer Anzeiger 250: 407-448.

CLARK P F, Ng. P K L (2011) A new species of Notonyx A. Milne-Edwards, 1873 (Crustacea: Decapoda: Brachyura: Goneplacidae) from the Philippines. Zootaxa 2982: 27-32.

CLARK P F (2011) The commercial exploitation of the Chinese mitten crab, Eriocheir sinensis in the River Thames, London: damned if we don’t and damned if we do. pp. 537-580. In: Galil B S, Clark P F, Carlton J D (Eds) In the Wrong Place – Alien Marine Crustaceans: Distribution, Biology and Impacts, Invading Nature – Springer Series in Invasion Ecology 6, Springer.

Allen C J, CLARK P F, Paterson G L J, Hawkins L E, Hauton C and Aryuthaka C (2011) New record of Dotilla intermedia De Man 1888 [Brachyura: Ocypodidae] from Thailand. Marine Biological Records 4: 1-6.

Froglia C and CLARK P F (2011) The forgotten Narrative of H.M.S. Challenger and the implications for decapod nomenclature. Zootaxa 2788: 45-56.

Palero F, Guerao G, CLARK P F and Abelló P 2011. Scyllarus arctus (Crustacea: Decapoda: Scyllaridae) final stage phyllosoma identified by DNA analysis. Journal of Marine Biological Association 91: 485-492.

2010

Fujita Y and CLARK P F (2010) The larval development of Chirostylus stellaris Osawa, 2007 (Crustacea: Anomura: Galatheoidea: Chirostylidae) described from laboratory reared material. Crustacean Research 39: 55-66.

CLARK P F and Ng P K L (2010) A description of the first zoeal stage morphology of Domecia glabra Alcock, 1899 (Crustacea: Brachyura, Domeciidae) with implications for the systematics of the Trapezioidea. Proceedings of the Biological Society of Washington 123: 258-273.

Richard J and CLARK P F (2010) A new species of Caridina H. Milne Edwards, 1837 (Decapoda, Caridea, Atyidae) from the Betsiboka River basin, Madagascar. Pp. 637-644. In: Fransen C, de Grave S and Ng P K L (Eds) Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustaceana Monographs 14: xii, 754 pp.

Ng P K L and CLARK P F (2010) The identity of Euryplax bevisi Stebbing, 1921 (Decapoda, Brachyura). Pp. 573-584. In: Fransen C, de Grave S and Ng P K L (Eds) Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustaceana Monographs 14: xii, 754 pp.

Low M E Y and CLARK P F (2010) Kraussia Dana, 1852 (Crustacea: Decapoda: Brachyura), the senior homonym of Kraussia Davidson, 1852 (Brachiopoda: Terebratulida: Terebratellidina): confirmation and authorship of the replacement name Kraussina Suess, 1859. Zootaxa 2628: 66-68.

Cumberlidge N and CLARK P F (2010) Two new species of freshwater crabs from Mount Elgon, East Africa (Brachyura: Potamoidea: Potamonautidae) and a redescription of Potamonautes niloticus (A. Milne-Edwards, 1887) with notes on their natural history and onchocerciasis. Journal of Natural History 44: 1807-1829.

Ng P K L, CLARK P F and Cuesta J A (2010) Establishment of a new subfamily for Shenius anomalus (Shen, 1935) (Crustacea: Decapoda: Brachyura: Dotillidae). Journal of Natural History 44: 1531-1553.

Allen C J, Paterson G L J, Hawkins L E, Hauton C, CLARK P F, Aryuthaka C (2010) Zonation on sandy tropical beaches: a case study using Dotilla intermedia i(Brachyura; Ocypodidae). Marine Ecology Progress Series 408: 97-107.

Ng P K L and CLARK P F (2010) On two new species of Notonyx A. Milne-Edwards, 1873 (Crustacea: Decapoda: Brachyura: Goneplacidae) from the Philippines and Fiji. Zootaxa 2509: 30–38.

Ng P K L, CLARK P F and Tan G S (2010) A new genus for Cleistostoma edwardsii Macleay, 1838 (Ocypodoidea: Camptandriidae) from South Africa. Pp. 251-260. In: Castro P, Davie P J F, Ng P K L and Richer de Forges B (Eds) Studies on Brachyura: a homage to Danièle Guinot. Crustacean Monographs 11: i-x, 1-366.

Cumberlidge N and CLARK P F (2010) A redescription of Potamonautes loveni (Colosi, 1924), a medically important freshwater crab from western Kenya and eastern Uganda, east Africa (Brachyura: Potamoidea: Potamonautidae). Pp. 61-74. In: Castro P, Davie P J F, Ng P K L and Richer de Forges B (Eds) Studies on Brachyura: a homage to Danièle Guinot. Crustacean Monographs 11: i-x, 1-366.

Palero F, Hall S, CLARK P F, Johnston D, Mackenzie-Dodds J and Thatje S (2010) DNA extraction from formalin-fixed tissue: New light from the Deep Sea. Biotechniques. Scientia Marina 74: 465-470.

Palero F, Guerao G, CLARK P F and Abelló P (2010) Final-stage phyllosoma of Palinustus A. Milne Edwards, 1880 (Crustacea: Decapoda: Achelata: Palinuridae) – The first complete description. Zootaxa 2403: 42-58.

Richard J and CLARK P F (2010) Caridina H. Milne Edwards, 1837 (Crustacea: Decapoda: Caridea: Atyoidea: Atyidae) – freshwater shrimps from eastern and southern Africa. pp. 305–337. In: De Grave S and Fransen C H J M (Eds) Contributions to shrimp taxonomy. Zootaxa 2372: 1-414.

Palero F, Guerao G, CLARK P F and Abelló P (2010) Final-stage phyllosoma of Palinustus A. Milne Edwards, 1880 (Crustacea: Decapoda: Achelata: Palinuridae) – The first complete description. Zootaxa 2403: 42-58.

Richard J and CLARK P F (2010) Caridina H. Milne Edwards, 1837 (Crustacea: Decapoda: Caridea: Atyoidea: Atyidae) – freshwater shrimps from eastern and southern Africa. pp. 305–337. In: De Grave S and Fransen C H J M (Eds) Contributions to shrimp taxonomy. Zootaxa 2372: 1-414.

2009

Rose M D, CLARK P F, Rainbow P S, Mortimer D N, Averns J M, Smith F, Fernandes A R and Panton S (2009) Chinese mitten crabs in European rivers: contamination with dioxins, PCBs, PBBs and PBDEs and implications for human consumption. Organohalogen Compounds 71: 901-904.

CLARK P F (2009) The bearing of larval morphology on brachyuran phylogeny. In: Martin J W, Crandall K A and Darryl F L (Eds) Decapod Crustacean Phylogenetics. Taylor & Francis London, Crustaceans Issues 18: 221-241.

CLARK P F, Mortimer D, Law R J, Averns J, Cohen B, Wood D, Rose M, Fernandes A and Rainbow P S (2009) Dioxin and PCB contamination in Thames Chinese mitten crabs: implications for human consumption as a control mechanism for an alien invader. Environmental Science and Technology 43: 1624-1629.

Huys R and CLARK P F (2009) Case 3467. Zosimidae Seifried, 2003 (Crustacea, Copepoda, Harpacticoida): proposed emendation of spelling to Zosimeidae to remove homonymy with Zosiminae Alcock, 1898 (Crustacea, Decapoda, Xanthidae). The Bulletin of Zoological Nomenclature 66: 24-29.

Naruse T and CLARK P F (2009) Establishment of a new genus for Asthenognathus gallardoi Serène & Soh, 1976 within Gaeticinae Davie & N.K. Ng, 2007 (Crustacea: Decapoda: Brachyura: Varunidae). Zootaxa 1987: 61-68.

Palero F, Guerao G, CLARK P F and Abelló P (2009) The true identities of the slipper lobsters Nisto laevis and Nisto asper (Crustacea: Decapoda: Scyllaridae) verified by DNA analysis. Invertebrate Systematics 23: 77-85.

Richard J and CLARK P F (2009) African Caridina (Crustacea: Decapoda: Caridea: Atyidae): redescriptions of C. africana Kingsley, 1882, C. togoensis Hilgendorf, 1893, C. natalensis (Bouvier, 1925) and C. roubaudi (Bouvier, 1925) with descriptions of 14 new species. Zootaxa 1995: 1-75.

2008

CLARK P F and Guerao G (2008) A description of Calocarcinus africanus Calman, 1909 (Brachyura, Xanthoidea) first zoeal stage morphology with implications for Trapeziidae systematics. Proceedings of the Biological Society of Washington 121: 475-500.

CLARK P F, Ng P K L, Fransen C H J M, Mclaughlin P A, Dworschak P C and Baba K (2008) A checklist of Crustacea Decapoda collected from Conic Island Cave and adjacent areas of Hong Kong. Journal of Natural History 42: 913-926.

CLARK P F and Ng P K L (2008) The lecithotrophic zoea of Chirostylus ortmanni Miyake & Baba, 1968 (Crustacea: Anomura: Galatheoidea: Chirostylidae) described from laboratory hatched material. Raffles Bulletin of Zoology 56: 85-94.

Ng P K L and CLARK P F (2008) A new species of Notonyx A. Milne-Edwards, 1873 (Crustacea: Decapoda: Brachyura: Goneplacoidea) from Indonesia. Zootaxa 1897: 20-26.

Ng P K L and CLARK P F (2008) A revision of Latopilumnus Türkay & Schuhmacher, 1985, and Aniptumnus Ng, 2002 (Crustacea: Decapoda: Brachyura: Pilumnidae). Journal of Natural History 42: 885-912.

Palero F, Guerao G and CLARK P F (2008) Palinustus mossambicus Barnard, 1926 (Crustacea: Decapoda: Achelata: Palinuridae); morphology of the puerulus stage. Zootaxa 1857: 44–54.