Vallance, G., and Fortey, R.A. 1968. Ordovician succession in North Spitsbergen. Proc. Geol. Soc London 1648, 91-7.


Fortey,  R.A. Tristichograptus, a triserial graptolite from the Lower  Ordovician of Spitsbergen.  Palaeontology14 :  188-199,  pls 26 - 29.


Fortey,  R.A.  & Holdsworth,  B.K.  The oldest  well-preserved radiolaria. Boll. Soc. Paleont. Ital. 10 : 35-41, 2 pls.


Fortey,  R.A. & Bruton, D.L. Cambrian-Ordovician rocks adjacent to Hinlopenstretet, North Ny Friesland, Spitsbergen. Bull. geol.  Soc. Am. 84 : 2227 - 2242.


Fortey,  R.A.  The Ordovician  trilobites  of  Spitsbergen.  I. Olenidae. Skr. norsk Polarinst. 160: 1-81, 24 pls.

Fortey,  R.A.  A new pelagic trilobite from the  Ordovician  of Spitsbergen, Ireland and Utah. Palaeontology, 17 : 111 - 124, pls 13,14.


Fortey, R.A.  Early Ordovician  trilobite  communities.  Fossils Strata 4 : 331-352.

Fortey, R.A. and Owens, R.M. Proetida - a new order of trilobites Fossils Strata 4: 227-239.

Fortey,R.A.   The  Ordovician  trilobites  of  Spitsbergen.II. Asaphidae,   Nileidae,   Raphiophoridae  and   Telephinidae   of   the Valhallfonna Formation. Skr. norsk. Polarinst. 162: 1 - 125, 41 pls.


Fortey,R.A.   Correlation  of  shelly  and  graptolitic  early Ordovician  successions,   based  on  the  sequence  in   Spitsbergen. 263 - 280 in Bassett,M.G. (ed.) The Ordovician System.  University of Wales Press and National Museum of Wales, Cardiff.

Fortey,R.A.  & Clarkson,E.N.K.  The function of  the  glabellar `tubercle' in Nileus and other trilobites. Lethaia 9: 101 -106.

Fortey,R.A.  and  Whittaker,J.E.P. Janospira  -  an  Ordovician microfossil in search of a phylum. Lethaia 9: 397 - 403.

Fortey,R.A. & Rushton,A.W.A. Chelidonoceaphalus trilobite fauna from the Cambrian of Iran.  Bull. Br.  Mus.  nat.  Hist.  (Geol.) 27: 321 - 340, pls 8 -12.

Bockelie,T.  and Fortey,R.A. An  early  Ordovician  vertebrate. Nature 260 : 36 - 38.

Bockelie, T.G., Bruton, D.L. and R.A. Fortey. Research on the Ordovician rocks of North Ny Friesland, Spitsbergen. Arbok Norsk Polarinstitutt (for 1975), 214 - 7.

Morris,N.J.  and  Fortey,R.A.  Tironucula  gen.  nov.  and  its significance in bivalve evolution. J. Paleont. 50: 701 - 709, pl.1.


Fortey,R.A. & Morris,S.F. Variation in lens packing of Phacops (Trilobita) Geol. Mag. 114: 25 - 32.

Fortey,R.A.   &  Barnes,C.R.  Early  Ordovician  conodont  and trilobite  communities  of  Spitsbergen:  influence  on  biogeography. Alcheringa 1: 297 - 309.

Shaw,F.C. and Fortey,R.A. Middle Ordovician facies and trilobite faunas in N. America. Geol. Mag. 114: 409 - 443.


Fortey,R.A. & Owens,R.M. Early Ordovician (Arenig) stratigraphy and  faunas of the Carmarthen District,  south-west Wales.  Bull.  Br. Mus. nat. Hist. (Geol.) 30: 225 - 294, 11 pls.

Fortey, R.A. and Morris,S.F. Discovery of nauplius-like trilobite larvae. Palaeontology 21: 823 - 833, pl.94.

Cope,J.C.W., Fortey,R.A. & Owens,R.M. Newly discovered Tremadoc rocks  in  the  Camarthen District,  South Wales.  Geol.  Mag.  115: 195 -198.


Fortey,R.A.  Early  Ordovician  trilobites  from  the  Catoche Formation (St. George Group ), western Newfoundland. Bull. Geol. Surv. Can. 321: 61 - 114. 15 pls.

Fortey,R.A.  & Owens,R.M.  Enrollment in the classification  of trilobites. Lethaia 12: 219 - 226.

Rushton, A.W.A., Fortey, R.A. and Owens, R.M. Excavation of two candidate  sections for stratotypes in Wales by the Nature Conservancy Council. Earth Science Conservation : 16: 1 - 5.


Fortey, R.A.  The  Ordovician  trilobites  of  Spitsbergen.III. Remaining  trilobites  of  the  Valhallfonna  Formation.  Skr.  norsk. Polarinst. 171: 1 - 113, 25 pls.

Fortey, R.A.  The Ordovician of Spitsbergen and its relevance to the  base  of  the Middle Ordovician in North  America. 33 - 40,  In Wones,D.R. (ed.) The Caledonides in the U.S.A. IGCP Project 27, 329, Blacksburg, Va.

 Fortey, R.A.  Generic longevity in Lower Ordovician trilobites: relation to environment. Paleobiology 6: 24 - 31.

Fortey, R.A.    &    Skevington,D.    Correlation    of    the Cambrian-Ordovician  boundary  between Europe and North  America:  new data from western Newfoundland. Can. J. Earth. Sci. 17: 382 - 388.

Fortey, R.A. & Rushton,A.W.A. Acanthopleurella Groom 1902: origin and life habits of a miniature trilobite. Bull. Br.  Mus.  nat.  Hist. (Geol.) 33: 79 - 89

Fortey, R.A.  Basilicus tyrannus (Murchison) and  the  glabellar structure of asaphid trilobites. Bull. Br. Mus. nat. Hist.  (Geol.) 34: 225 - 264.


Fortey, R.A.  Prospectatrix  genatenta  (Stubblefield)  and  the trilobite superfamily Cyclopygacea. Geol. Mag. 118: 603 - 614.

Fortey, R.A. The Burgess Shale: a unique Cambrian fauna.  Nature 293: 189


Fortey, R.A.  &  Jefferies, R.P.S.  Fossils  and  phylogeny: a compromise approach. 197 - 234, In Joysey,K.A. & Friday,A.E.  (eds.) Problems of Phylogenetic reconstruction. Academic Press.

Fortey, R.A.,  Landing,E.  &  Skevington,D.  Cambrian-Ordovician boundary  sections  in  the  Cow  Head  Group,  western  Newfoundland. 95 - 129, In Basset, M.G. and Dean,W.T. (eds.) The Cambrian-Ordovician boundary. National Museum of Wales, Cardiff.

Fortey, R.A.  & Morris,S.F. The Ordovician trilobite  Neseuretus from Saudi Arabia and the palaeography of the Neseuretus fauna related to  Gondwana in the earlier Ordovician.  Bull.  Br.  Mus.  nat.  Hist. (Geol.) 36: 63 -75.

Fortey, R.A.   &  Peel,J.S.  Fossils  from  the  Poulsen  Cliff Formation,  Washington Land, western North Greenland.  Gronlands geol. Unders. 108: 24.

Fortey, R.A.  Fossils:  the key to the  past.  179  pp.  William Heinemann Ltd.,  London,  and British Museum (Natural History).  (U.S. edition Van Nostrand Rheinhold, New York)

Briggs, D.E.G. & Fortey, R.A. The cuticle of aglaspid arthropods: a  red herring in the early history of the vertebrates.  Lethaia 15: 25 - 29.

Owens, R.M. & Fortey, R.A. Arenig rocks of the Carmarthen-Llanarthney  district.  249 - 258,  In Bassett,M.G.  (ed.) Geological   Excursions  in  Dyfed,   south-west   Wales.   Geologists Association.

Owens, R.M., Fortey, R.A., Cope,J.C.W.,   Rushton,A.W.A.  & Bassett,M.G.  Tremadoc  faunas  from the  Carmarthen  district,  South Wales. Geol. Mag. 119:1 - 38, 8 pls.

Cocks, L.R.M. & Fortey, R.A.  Faunal  evidence  for   oceanic separations in the Palaeozoic of Britain. J. Geol. Soc. 139:465 - 478.

Cooper, R.A. & Fortey, R.A.  The  Ordovician  graptolites  of Spitsbergen. Bull. Br. Mus. nat. Hist. (Geol.) 36:157 - 302, 6 pls.


Fortey, R.A. Cambrian-Ordovician boundary trilobites from western Newfoundland   and  their  phylogenetic  significance.   Spec.   Pap. Palaeontology. 30: 179 - 211, 23-27.

Fortey, R.A.  &  Peel,J.S.  The  anomalous  bathyurid  trilobite Ceratopeltis and its homoemorphs. ibid: 51 - 57, pl.6.

Cooper, R.A.,  &  Fortey,R.A.  Development  of  the  graptolite rhabdosome. Alcheringa 7: 201 - 221.

Fortey, R.A.  Geometrical  constraints in  the  construction  of graptolite stipes. Paleobiology 9: 116 -125.


Fortey, R.A. Global Ordovician transgressions and regressions and their biological implications.37-50, In Bruton,D.L.  (ed.) Aspects of the Ordovician System. Universitetsforlaget, Oslo.

Fortey, R.A.  & Owens, R.M. A synopsis of the Arenig Series in South Wales. Proc. Geol. Assoc. 95, 389 - 90.

Whittington, H.B.,  Dean,W.T.,  Fortey,R.A.,   Rickards,R.B., Rushton,A.W.A. & Wright,A.D. Definition of the Tremadoc Series and the series of the Ordovician System in Britian. Geol. Mag. 121: 17 - 33.

Fortey,R.A. & Shergold,J.S. Early Ordovician trilobites from the Nora  Formation,  central  Australia.  Palaeontology  27: 315 - 366, pls.38 - 46.


Fortey,  R.A. Gradualism and punctuated equilibria as competing and complementary theories. Spec. Pap. Palaeontology 33: 17-28.

Fortey,  R.A.  Pelagic trilobites as an example of deducing the life habits of extinct arthropods. Trans. R. Soc. Edinb. 76 : 219-30.

Thomas, A.T. & Fortey, R.A. Trilobita, In, Murray,  J.W.  (ed.) Atlas of Invertebrate Macrofossils. Longman's, London.

Morris,  S.F.  & Fortey, R.A. Catalogue of the type and figured specimens  of  Trilobita  in the  British  Museum  (Natural  History). British Museum (Natural History), 183 pp, 3 pls.


Fortey, R.A. & Cooper, R.A. A phylogenetic classification of the graptoloids. Palaeontology 29 : 631-654, 11 figs.

Fortey,  R.A.  The type species  of  the  Ordovician  trilobite Symphysurus:  systematics,  functional morphology and terrace  ridges. Palaont. Z. 60, 255-275, 12 figs.

Fortey,  R.A.  &  Cocks,  L.R.M.  Introduction:   Fossils  and tectonics. J. geol. Soc. 143, 149-150.

Fortey,  R.A.  & Cocks, L.R.M.  Marginal faunal belts and their structural implications,  with examples from the Lower Palaeozoic.  J. geol. Soc. 143, 151-160, 7 figs.

Fortey,R.A. The course of evolution. Pp. 66-77, in Hallam, A.  & Berry,  R.J.  (eds)  The Collins  Encyclopedia  of  Animal  Evolution. Collins, London.

Zhou,  Zhiyi and Fortey, R.A.  Ordovician trilobites from North and Northeast China. Palaeontographica Abt. A 192, 157-210, 12 pls.

Fortey, R.A. Early Ordovician trilobites from the Wandel Valley Formation, eastern North Greenland. Rapp. Gronlands geol. Unders.132, 15-25.

Cocks,  L.R.M. & Fortey, R.A. New evidence on the South African Lower Palaeozoic: age and fossils reviewed. Geol. Mag. 123, 437-444, 1 fig.


Fortey,  R.A.  and Bell, A.  Branching geometry and function of multiramous graptolites. Paleobiology 13, 1-19, 21 figs.

Fortey,  R.A. and Owens, R.M. The Arenig Series in South Wales. Bull. Br. Mus. nat. Hist. (Geol.) 41 , 69-307, 135 figs.


Fortey,  R.A.  and Chatterton,  B.D.E.  Classification  of  the trilobite suborder Asaphina. Palaeontology 31, 165-222, pls 17-19.

Fortey,  R.A.  Seeing is believing:  gradualism and  punctuated equilibria in the fossil record. Sci. Prog. Oxf. 72, 1-19.

Fortey,  R.A.  and Cocks, L.R.M.  Arenig to  Llandovery  faunal distributions in the Caledonides.  In Harris,  A.  L.  & Fettes,  D.J. (eds) The Caledonian - Appalachian Orogen, 233-246.  Geol.  Soc. Spec. Pub. 38.

Fortey,  R.A. The Ordovician trilobite Hadrohybus Raymond 1925, and its family relationships. Postilla 202, 1-7.

Fortey, R.A. The international correlation of the lower part of the    Ordovician   and   how   it   should    influence    Laurentian chronostratigraphy. Bulletin N.Y.St. Mus. 462, 43 - 47.

Cocks,  L.R.M.  and Fortey, R.A.  Lower Palaeozoic  facies  and faunas around Gondwana. Geol. Soc. Spec. Pub. 37, 183-200.

Briggs, D.E.G., Fortey, R.A. & Clarkson, E.N.K.  Extinction and the fossil record of the arthropods. In Larwood, G.P. (ed.) Extinction and survival in the fossil record,  171 - 209.  Systematics  Association Special Volume No. 34. Clarendon Press, Oxford.


Fortey,   R.A.,   Owens,  R.M.,   and  Rushton,   A.W.A.   The palaeogeographic   position  of  the  Lake  District  in   the   early Ordovician. Geol. Mag.126, 9 - 17, 2 figs.

Fortey,  R.A.  and Whittington, H.B. The Trilobita as a natural group. Historical Biology 2, 125 -138.

Fortey,  R.A.  Genetics, Paleontology  and  Macroevolution,  by J.Levinton, essay review in Historical Biology 2, 179 -182.

R. A. Fortey. The collection connection (Review of Wonderful Life by S.J. Gould). Nature 342, 303.

Briggs,  D.E.G.  & Fortey,  R.A.  The early  radiation  of  the Arthropoda, Science, N.Y., 246, 241 - 243.

Fortey,  R.A.  There are extinctions and extinctions:  examples from the Lower Palaeozoic. Phil. Trans. Roy. Soc. ser. B 325, 327 - 355

Fortey,   R.A.  A  Devonian  trilobite  fauna  from  Thailand. Alcheringa 13, 257 - 267

Fortey,   R.A.  &  J.S.  Peel.  Stratigraphy  and  hystricurid trilobites  of  the  Christian Elv  Formation  (Lower  Ordovician)  of western North Greenland. Rapp. Groenlands geol. Unders 144, 5 - 15.


Fortey,  R.A. Overview: Analysis of taxonomy and phylogeny.  In Briggs,  D.E.G.  & Crowther,  P.R.  (eds) Palaeobiology:  a synthesis, 425-427. Blackwell, Oxford.

Fortey, R.A. & Cocks, L.R.M. Fossils and tectonics.  In Briggs, D.E.  G. and Crowther, P. R. (eds) Palaeobiology: a synthesis 482-492. Blackwell, Oxford.

Fortey,  R.A.   Ontogeny, hypostome  attachment  and  trilobite classification. Palaeontology 33 (3), 529-576.

Cocks,  L.R.M.  & Fortey, R.A.  Biogeography of Ordovician  and Silurian faunas, pp. 97-104, In McKerrow, W.S. & Scotese,  C.R.  (eds) Palaeozoic Palaeogeography and Biogeography. Geological Society Memoir No. 12.

Fortey,  R.A.  & Peel, J.S.  Early  Ordovician  trilobites  and molluscs from the Poulsen Cliff Formation,  Washington Land,  western North Greenland. Bull. geol. Soc. Denmark 38, 11-32.

Fortey,   R.A.  &  Owens,  R.M.   Arenig  biostratigraphy  and correlation in the Welsh Basin. J. Geol. Soc. London 147: 607-610.

Fortey,  R.A.,  Beckly, A.J.  & Rushton,  A.W.A.  International correlation  of  the base of the Llanvirn Series,  Ordovician  System. Newsletters in Stratigraphy 22: 119-142.

Fortey,  R.A.  & Owens, R.M. Trilobites. Chapter 5 in McNamara, K.J. (ed.) Evolutionary Trends, pp. 121 - 142. Belhaven Press, London.

Fortey, R.A. Trilobite evolution and systematics. Pp. 44-65,  In Arthropod  Paleobiology.  Short  courses in Paleontology  No.  3,  The Paleontological Society of America

Fortey,  R.A.  & Owens, R.M.  Evolutionary  radiations  in  the Trilobita.  Pp.  139 -164, In Taylor, P.D. & Larwood, G.P.  (eds) Major evolutionary  radiations  Systematics Association Special  Volume  42. Clarendon Press Oxford.


Cooper, R.A., Fortey, R.A. & Lindholm, K. Latitudinal and depth zonation of early Ordovician graptolites. Lethaia 24: 199 - 218.

Fortey,  R.A.  & Owens, R.M.  The  early  Ordovician  trilobite Beltella. Proc. Bristol nat. Hist. Soc. 49: 69 -79, 1 pl.

Fortey, R.A. Fossils: the key to the past. 2nd edition. 187 pp. Natural History Museum Publications and Harvard University Press (US). 

Fortey,  R.A.  & Owens, R.M. A trilobite fauna from the highest Shineton  Shales  in Shropshire,  and the correlation  of  the  latest Tremadoc.  Geol. Mag. 122: 437 - 464.

Fortey,  R.A.  & Wilmot, N.V.  Trilobite cuticle  thickness  in relation to palaeoenvironment. Pal. Zeitschr. 65: 141 -151.

Chapters 5 (S.  Rigby and R.A.  Fortey),  23 - 32,  and 12  (R.A. Fortey,  I.  Strachan,  R.B. Rickards), 69 -77, In Rickards,  R.B.  and Palmer, D. (Eds) Graptolites : Writing in the rocks. Boydell Press.

Fortey,  R.A.,  Bassett, M.G., Harper,  D.A.T.,  Hughes,  R.A., Ingham, J.K., Molyneux, S.G., Owen, A.W., Owens, R.M., Rushton, A.W.A. & Sheldon,  P.R. Progress and problems in the selection of stratotypes for  the  bases of series in the Ordovician System of  the  historical type area in the U.K. Pap. Geol. Surv. Canada 90 - 9: 5  -25.

Fortey,  R.A. Wales and the Ordovician System. Geology Today 7: 136 -138.


Fortey, R.A. Taxonomy and extinction patterns. In, The Unity of Evolutionary  Biology.  Proceedings of the International  Congress  of Evolutionary & Systematic Biology, University of Maryland,  1990,  pp. 82 - 88.

Fortey, R.A. Ordovician trilobites from the Durness Group,  North-West Scotland, and their palaeobiogeography. Scot. J. Geol.  28: 115 - 121.

Fortey,  R.A.  & Mellish,  C.J.T.  Are some fossils better than others for inferring palaeogeography? Terra Nova 4: 210 - 216.

Fortey,  R.A.  South Britain, Tremadoc and Arenig (with  A.W.A. Rushton) and Llanvirn (with R.E. Bevins) in,  Atlas of Palaeogeography and lithofacies. (J.C.W. Cope, P.F. Rawson and J.K. Ingham [eds) Mem. Geological Society of London No. 13.

Fortey,  R.A.  & Cocks, L.R.M. The Ordovician paleogeography of the  North  Atlantic region as a test case in the use of  fossils  for continental reconstruction. Tectonophysics 206: 147 -158.

Briggs, D.E.G. and Fortey, R.A. The early Cambrian radiation of the arthropods,  In Lipps, J.H. & Signor, P.W. (eds). Origin and early evolution of the Metazoa, p. 335 - 373. Plenum Press, N.Y.

Briggs,  D.E.G.,  Fortey,  R.A.  and  Wills  M.   Morphological disparity in the Cambrian. Science N.Y. 256: 1670-1673.

Fortey,  R.A.  & Owens, R.M. The Habberley Formation:  youngest Tremadoc in the Welsh Borderlands. Geol. Mag. 129: 553 - 566.

Fortey,  R.A.,  Pankhurst,  R.J. &  Herve Ç,  F.  1992.  Devonian trilobites at Buill, Chile (42 S). Revista Geol. Chile 19: p. 133 -144.

Fortey,  R.A. & Owens, R.M. The trilobite Angelina unstretched. Geology Today 8: 219 - 221.

Briggs, D.E.G., Fortey, R.A. and Wills, M.A. Cambrian and Recent  morphological disparity: response to Foote and Gould, and Lees. Science 258: 1817 - 18.


Briggs,  D.E.G.,  Fortey,  R.A. and M.  Wills.  How big was the Cambrian  evolutionary  explosion?A  taxonomic  and   morphological comparison  of  Cambrian  and  Recent arthropods,  pp.  33  -  44  in, Evolutionary Patterns and Processes. Linnean Society of London.

Fortey,  R.A.  and Thomas,  R.E. The case of the  velvet  worm. Nature, 361, 205 - 6.

Fortey, R.A. Charles Lapworth and the biostratigraphic paradigm. Journal of the Geological Society 150: 209 - 218.

Fortey, R.A. The Hidden Landscape: a journey into the geological past. Jonathan Cape, London, 310 pp. (Reprinted as paperback, by Pimlico Press, 1994)

Fortey, R.A. in, Romano, M. and others:  Arthropoda (Trilobita) In, The Fossil Record 2 (M.J. Benton, ed.): 279 - 296. Chapman & Hall.


Fortey,  R.A.  Late Cambrian trilobites from the Sultanate of Oman. Neues Jahrbuch fur Geologie und Palaontologie 194: 25 - 53

Wills, M.A., Briggs, D.E.G. and Fortey, R.A. Disparity as an evolutionary index: a comparison of Cambrian and Recent  arthropods. Paleobiology 20: 93 -130.


Fortey R.A.  & Theron, J.  A new Ordovician arthropod Soomaspis and the agnostid problem. Palaeontology 37: 841 - 861.

Fortey R.A., Harper, D.A.T,. Ingham, J.K., Owen, A.W. and Rushton, A.W.A. A revision of Ordovician series and stages from the historical type area. Geological magazine 132: 15 - 30

Fortey, R.A. Trilobite life habits and Lower Palaeozoic continents. Geoscientist, 5, 16 -17.

Charles Lapworth and the biostratigraphic paradigm, in Le Bas, M.J. (ed.) Milestones in Geology. Memoir of the Geological Society 16, 93 -102.

Barnes, C..R., Fortey, R.A. and Williams, S.H. The Pattern of Global Bio-events during the Ordovician Period, in Walliser O. (ed.) Global Events and Event Stratigraphy, 113 - 139. Springer-Verlag, Berlin.

Wills, M.A., Briggs, D.E.G., Fortey, R.A., and Wilkinson, M. The significance of fossils in understanding arthropod evolution. Verhandlungen der Deutschen Zoologischen Gesellschaft, 88, 203 - 215.

Fortey, R.A. The Ordovician Series of the Historical Type Area: revision as a contribution to their utility in international correlation.  In Ordovician Odyssey: Short papers for the Seventh International Symposium on the Ordovician System, 11 - 14, Pacific Section SEPM Book 77.

Hughes N. C. and Fortey, R.A. Sexual dimorphism in trilobites, with an Ordovician case study. Ibid. 419 - 422.

Edgecombe, G.D. and Fortey, R.A. Brianurus new genus, a replacement name for Briania Chasen and Kloss, 1930 (Aves) [pro Briania Edgecombe 1994 (Trilobita) preoccupied). Journal of Paleontology 69 (3), 615.


Fortey, R.A. and Droser, M.L. A trilobite fauna from the base of the Middle Ordovician, Basin ranges,  western USA Journal of Paleontology, 70, 73 - 99.

Droser, M.L., Fortey, R.A. and Xing, L.  The Ordovician Radiation. American Scientist 84: 122 - 131 (and cover).

Fortey, R.A., Briggs, D.E.G. and M. A. Wills. The Cambrian  evolutionary 'explosion': decoupling  cladogenesis from morphological disparity. Biological Journal of the Linnean Society 57: 13 - 33.


Fortey, R.A., Briggs, D.E.G. and Wills, M.A. The Cambrian evolutionary 'explosion' recalibrated. Bioessays 19:  429 - 434.

Fortey, Richard. “Life: an unauthorised biography” Harper Collins, London 400pp

Fortey, R.A. & Owens, R.M. Bubble-headed trilobites, and a new olenid example. Palaeontology 40: 451 - 460.

Fortey, R.A.  Late Ordovician trilobites from southern Thailand. Palaeontology 40: 397 - 450.

Landing, E., Bowring, S.A., Fortey, R.A., and Davidek, K.L. U-Pb date from Avalonian Cape Breton Island and geochonologic calibration of the Early Ordovician. Canadian Journal of Earth Sciences 34: 724 - 730.

Cocks, L.R.M. and Fortey, R.A. A new Hirnantia fauna from Thailand and the biogeography of the latest Ordovician of South-east Asia. Geobios M.S. No. 20, 117 - 126.

Fortey R.A. Classification, in Treatise on Invertebrate Paleontology. Part  O (Arthropoda 1, revised), 289 - 302.

Fortey, R.A. & Owens, R.M. Evolutionary History, Ibid. 249 - 288.

Fortey, R.A. & Thomas, R.H. (eds) Arthropod Relationships. Chapman & Hall, London, 383 pp. Systematics Association Special Volume Number 55.

Wills, M.A., Briggs, D.E.G. and Fortey, R.A. Evolutionary correlates of arthropod tagmosis: scrambled legs, 57 - 65 in Arthropod Relationships

Austin, J.J., Ross, A.J., Smith, A,. B.,  Fortey, R.A. & Thomas, R.H. Problems of reproducibility - does geologically ancient DNA survive in amber-preserved insects? Proceedings of the Royal Society of London 264: 467 - 474.

Fortey R.A. & Seilacher, A. The trace fossil Cruziana semiplicata and the trilobite that made it. Lethaia 30:105 -112.

Adrain, J.M. and Fortey, R.A. Ordovician trilobites from the Tourmakeady Limestone, western Ireland. Bulletin of the British Museum (Natural History) Geology series, 53:79 - 115.


McCormick, T. and Fortey, R.A.  Independent testing of a paleobiological hypothesis: the optical design of two Ordovician pelagic trilobites reveals their  relative paleobathymetry. Paleobiology 24: 235 - 253.

L.R.M. Cocks and Fortey, R.A.  The Lower Palaeozoic margins of Baltica. Geol. Foren. Forh. 120. 173 - 179.

Wills, M.A., Briggs, D.E.G., Fortey, R.A, et al. An arthropod phylogeny based on fossil and Recent taxa. In Edgecombe, G.E. (ed.)  Arthropod fossils and phylogeny, 33 -106. Columbia University Press.

Cooper, A. and Fortey, R.A. Evolutionary explosions and the phylogenetic fuse. Trends in Ecology and Evolution. 13(4):151 - 156.

Richard Fortey LIFE: a natural history of the first 4000 million years of life on earth (reprinted from Life: An Unauthorised biography) Alfred Knopf, New York.

Fortey, R.A. and Cocks L.R.M. Lower Palaeozoic palaeogeography of Thailand: a review In, Hall, R and Holloway J.D. (eds). Biogeography and Geological Evolution  of Southeast Asia, 43 - 57. Backhuys, Leiden.

Adrain,  J.M., Fortey, R.A. and Westrop, S.R. Post-Cambrian trilobite diversity and Evolutionary Faunas. Science 280:1922 - 5.

Fortey, R.A. and Hughes, N.C. Brood Pouches in  trilobites. Journal  of Paleontology 72: 638 - 649.

Davidek, K. Landing, E, Bowring, S.A., Westrop, S.R., Rushton, A.W.A., Fortey, R.A. and Adrain J.M. New uppermost Cambrian U-Pb date from Avalonian Wales and age of the Cambrian Ordovician boundary. Geological Magazine 135: 305 - 9.

Bromham, L., Rambaut, A, and Fortey, R. et al. Testing the Cambrian explosion hypothesis by using a molecular dating technique. PNAS 95:12386 - 9.

Austin, J.J., Smith, A.B., Fortey, R.A. and Thomas, R.H. Ancient DNA  from amber inclusions: a review of the evidence. Ancient biomolecules 2: 167 -176.


Fortey, R.A. & Owens, R.M.  The Trilobite exoskeleton, in, Savazzi, E. (ed.) Functional morphology of the invertebrate exoskeleton, 537 - 562. Wiley, New York.

Fortey, R.A. and Droser, M.L. Trilobites from the base of the type Whiterockian (Middle Ordovician) in Nevada. Journal of Paleontology, 73: 182 - 201.

McCormick, T., and Fortey, R. A. The most widely distributed trilobite species: Ordovician Carolinites genacinaca Journal of Paleontology. 73: 202 - 218.

Fortey, R.A. & Owens, R.M. Feeding habits in trilobites. Palaeontology  42 (3): 429 - 465.

Fortey, R A. Olenid trilobites as chemoautotrophic symbionts. Acta Universitatis Carolinae 43: 355 - 6.

Patterson, C. (late) & Fortey R.A. Thomas Stanley Westoll. Biographical Memoirs of the Fellows of the Royal Society, 45: 531 - 546.

Fortey, R.A. Colin Patterson. Biographical Memoirs of the Fellows of the Royal Society, 45:365 - 37.


Fortey , R.A. Olenid trilobites: the oldest chemoautotrophic symbionts? Proceedings of the National Academy of Sciences, Washington, 97(12) 6574-8.

 Fortey, R.A., Harper, D. A. T., Ingham, J.K., Owen, A.W., Parkes, M.A., Rushton, A.W.A. & N.H. Woodcock. A revised correlation of Ordovician Rocks in the British Isles. Geological Society, Special report No. 24, 83 pp, 34 figs.

 R. A. Fortey. “Trilobite! Eye Witness to Evolution.” HarperCollins, London. 269 pp.

 Edgecombe, G.E. & Fortey, R.A. Silurian trilobites from the El Carmen Formation, Bolivia. Senckenbergiana lethaia 79, 329-55.

 Fortey R. A. Crystal Eyes. Natural History 109, 68-73.

 Landing, E, Bowring, S.A.,  Davidek, K.L., Rushton, A.W.A., Fortey, R.A. & Wimbledon, W.A.P. Cambrian-Ordovician boundary age and duration of the lowest Ordovician Tremadoc Series based on U-Pb zircon dates from Avalonian Wales. Geological Magazine 137, 485-494.

Wills, M A and Fortey R A. The shape of life: how much is written in stone? Bioessays 22:1142-1152.


Owens, R.M., Lane, P.D., Thomas A.T, Fortey , R.A.  et al. South Wales and the Welsh Borderland. Field Guide for the Third International Symposium on Trilobites. Palaeontological Association, London.

Zhang Y.-D. & Fortey, R. A. The proximal development and thecal structure of the Ordovician graptolites Tylograptus and Sinograptus. Palaeontology 44, 553-573.

Forey, P.L. & Fortey, R.A. Fossils in the reconstruction of phylogeny. 515-9, in Briggs, D.E.G. and Crowther, P.R. (eds) Palaeobiology II. Blackwells, Oxford.

Fortey, R.A. The Cambrian explosion exploded? Science, N.Y. 293, 438-9.

Bettley, R., Fortey, R.A. and Siveter, D.J.. High resolution correlation of Anglo-Welsh Middle to Upper Ordovician sequences and its relevance to international chronostratigraphy.  Journal of the Geological Society, London, 158, 937-52.

Adrain. J.M., Westrop, S.R. Landing, E. and Fortey, R.A. Systematics of the Ordovician trilobites Ischyrotoma and Dimeropygiella with species from the type Ibexian area, western USA. Journal of Paleontology  75, 947-971.

R. A. Fortey. Trilobite systematics: the last 75 years. Journal of Paleontology 75, 1141-51.

Siveter, D.J., Waloszek, D., Williams, M. and Fortey, R.A. Crustaceans and the Cambrian explosion. Science, N Y 294, 2047.


McCormick, T. and R.A.Fortey. The Ordovician trilobite Carolinites, a test case for microevolution in a macrofossil lineage. Palaeontology, 45: 229 - 257.

R. A Fortey. Retrospective. Stephen Jay Gould (1941-2002). Science, N.Y. 296:1984.

 Rushton, A. W. A., Cocks, L.R.M. and Fortey, R. A. Upper Cambrian trilobites  and  brachiopods from Severnaya Zemlya, Arctic Russia, and their implications for correlation and biogeography. Geological magazine 139: 281 - 290.

 Lees, D C.,  Fortey, R.A and Cocks, L.R.M. Quantifying paleogeography using biogeography: a test case for the Ordovician and Silurian of Avalonia based on brachiopods and trilobites. Paleobiology 28: 343 - 363.

 Fortey, R A. Fossils: the key to the Past. 3rd Edition, 232 pp. Natural History Museum, London, and Smithsonian Institution, Washington.

 Cocks, L. R.M and Fortey, R.A. The palaeogeographical significance of the latest Ordovician fauna from the Pangsha Pye Formation of Burma. Special Papers in Palaeontology 67: 57 - 76.



Fortey, R. A. and Cocks, L. R. M. Palaeontological evidence bearing on global Ordovician–Silurian continental reconstructions, Earth Science Reviews, 61: 245 - 307.

Vandenbrouke, T., Fortey, R.A., Siveter, D.J. and Rickards, R.B. Chitinozoans from key sections in the Upper Ordovician Series: new GSSPs and classical British sections 151-2, in, Ordovician from the Andes. Instituto Superior de Correlacion Geologica Insugeo. Serie Correlacion Geologica 17.

 Cocks, L.R. M. and Fortey, R.A. Central Asian terranes and biogeography in the Ordovician, 511 - 6, in, Ordovician from the Andes. Instituto Superior de Correlacion Geologica Insugeo. Serie Correlacion Geologica 17: 511 - 515.

 Fortey, R.A. and B.D.E.Chatterton. A Devonian trilobite with an eye shade. Science 301, 1689.

 Fortey, R.A. and Rushton, A.W.A. A new aglaspidid arthropod from the Lower Ordovician of Wales. Palaeontology 46:1031 -1038.

 Fortey, R. A. Strangelove versus Calculus: the two faces of Science, pp. 66-69 in Words and Pictures: explaining science through art and writing. Cumbria Institute of the Arts.

 Lane, P.D. Fortey, R. A. and Siveter, D.J. (eds). Trilobites and their relatives. Special Papers in Palaeontolog,y 70: 397(and Introduction, by Fortey, Siveter and Lane, p. 7-8).

 Fortey, R. A. Foreword, ix-x, in Hou X-G et al. The Cambrian Fossils of Chengjiang, China.  Blackwells, Oxford.

 Fortey, R.A., Jackson J. and Strugnell, J. Phylogenetic fuses and evolutionary ‘explosions: conflicting evidence and critical tests. In  Donoghue, P.C. J. and Smith M. P. (eds) Telling the evolutionary time, pp. 41-65. CRC Press.

Fortey, Richard, Foreward to Frontiers 03: Science and technology 2002-2003. (edited by Tim Radford). Atlantic Books, 128 pp.


Richard Fortey “The Earth: an intimate history.” HarperCollins. 501 pp. [Published in the USA under the title “Earth an intimate history” by Knopf]

Forey, P.L., Fortey, R.A., Kenrick, P. and Smith, A.B. Taxonomy and  fossils: a critical appraisal. Philosophical Transactions of the Royal Society, Series B 359, 639 - 653.

Adrain, J.M. Edgecombe, G.D., Fortey, R.A, et al. Trilobites, pp. 231 - 254, in Webby, B.D., Paris, F., Droser, M.L. and Percival, I. (eds) The great Ordovician biodiversification event. Columbia University Press. 484 pp.

Fortey R, A. The lifestyles of the trilobites. American Scientist 92: 446 - 453.


Cocks, L.R. M., Fortey, R.A and C.P. Lee. A review of Lower and Middle Palaeozoic biostratigraphy  in west peninsular Malaysia and southern Thailand in its context within the Sibumasu Terrane. Journal of Asian Earth Sciences 24: 703 - 717.

Fortey, R.A. Ordovician, in Selley, R. Cocks, L.R.M. and Plimer, I. (eds) Encyclopedia of Geology, 4:175 - 284.

Cotton, T.J. and Fortey, R.A. Comparative morphology and relationships of the Agnostida, in Koenemann, S. and Jenner, R.A. (eds) Crustacea and arthropod relationships. Crustacean Issues 16: 95 -136.

Fortey, R.A. and Cocks, L.R.M. Late Ordovician global warming – the Boda event. Geology  33, 405 - 408.

Knell, R and Fortey R.A. Trilobite spines and beetle horns: sexual selection in the Palaeozoic? Biology Letters 1(2):196 - 9.

Briggs, D.E.G. and Fortey, R.A. Wonderful strife: systematics, stem groups and the phylogenetic signal of the Cambrian radiation.  In Macroevolution: diversity, disparity, contingency. Supplement to Paleobiology 31: 94 -112.

Fortey, R.A. Opipeuterella, a replacement name for the trilobite Opipeuter Fortey, 1974, preoccupied. Journal of Paleontology 79: 989.

Pollitt, J.R., Fortey, R.A. and Wills, M.A. Systematics of the trilobite families Lichidae Hawle & Corda 1847 and Lichkephalidae Tripp, 1957; the application of Bayesian inference to morphological data. Journal of Systematic Palaeontology 3: 225 - 241.

Fortey, R.A., Zhang Yuandong and Mellish, C.J.T. The relationships of biserial graptolites. Palaeontology 48: 1241 - 1272.

Fortey, R.A., and Cocks, L.R.M.  Fossil evidence supports Lower Palaeozoic oceanic separations. Courier Forsch.-Inst. Senckenberg 254: 465 - 472.


Cronier, C and Fortey, R.A. Morphology and ontogeny of an early Devonian phacopid with reduced sight from southern Thailand.  Journal of Paleontology 89: 529 - 536.

Yuan, W.-W., Fortey, R.A. and Turvey, S.T. 2006. Ontogeny and relationships of the trilobite Pseudopetigurus Prantl & Pribyl. Palaeontology 49: 537 - 546.

Fortey R. A. .A new deep water Upper Ordovician (Caradocian) trilobite fauna from South –West Wales. Geological Journal 41: 243 - 253.

Chatterton, B.D.E., Fortey, R.A., Brett. K., Gibb, S. and McKellar, R. Trilobites from the upper Lower to Middle Devonian Timrhanrhart Formation, Jebel Gara el Zguilma, southern Morocco. Palaeontographica Canadiana 25, 177 pp.


Foreword. Whatever is under the earth Gordon Herries Davies, Geological Society Publishing House.

Siveter, D.J., Fortey, R.A., Sutton, M. Briggs, D.E.G., and Siveter, D. A Silurian ‘marrellomorph’ arthropod. Proceedings of the Royal Society B, 274: 2223 - 9.

Forward into the past. Geoscientist 13: 121 - 4.

An autobiography of the frozen, 42 - 3 In Findings on ice. Pars Foundation.

Fortey R.A., and Rushton A.W.A. 2007. The mid-Cambrian trilobite Beishanella and the limits of convergent evolution. Memoirs of the Australasian Association of Palaeontologists 34: 391 - 9.


Fortey, R. A. Dry Store Room Number 1. Harper Collins, London. 332 pp. (Published in USA by Knopf)

Chatterton, B. D. E. & Fortey, R.A. Linear clusters of articulated trilobites from Lower Ordovician (Arenig) strata at Bini Tin Zoulin North of Zagora, southern  Morocco, in Rabano, I, Gozalo, R & Garcia –Bellido, D (eds) Advances in trilobite research, Publicaciones del Instituto Geologico y Minero de Espana. Cuarernos del Museo Geominero 9:73 -78.

Siveter, D.J., Briggs, D.E.G., Siveter, D.J., Sutton, M.D. and Fortey, R.A. Horns, eggs and legs: exceptionally preserved new arthropods from the Hereford (Silurian) Lagerstatte. Ibid, 371 - 4.

Fortey, R.A. In retrospect: Liebniz’s Protogaea. Nature 455 (4): 35.

Fortey, R.A. Cambrian evolutionary explosion. In, Benton, M. (ed.) The seventy great mysteries of the natural world. Thames and Hudson.


Vandenbroucke, T. R.A.,  Fortey, R.A.., Verniers, J. and Ancilletta, A. A modern assessment of Ordovician chitinozoans from the Shelve and Caradoc areas, Shropshire, and their significance for correlation. Geological Magazine, 146: 216 - 236

Fortey, R.A.. Predicciones geologicas para el futuro del humanidad In, Punset,. E. (ed) El impacto de la  ciencia en la sociedad 2008: Por que somos como somos, 93 - 104. Fundacion Banco Santander.

Fortey, R.A. Biogeography and evolution in the Lower Palaeozoic. 60-72, in Sepkoski, D. and Ruse, M. (eds) The Paleobiological Revolution: essays on the growth of modern paleontology. University of Chicago Press.

Fortey, R.A., A new giant asaphid trilobite from the Lower Ordovician of Morocco. Memoirs of the Association of Australasian Palaeontologists 37: 9 -16.

Fortey, R.A. & Rushton, A.W.A., The Ordovician aglaspidid arthropod Tremaglaspis reconsidered. Memoirs of the Association of Australasian Palaeontologists 37:17 - 23.

Cocks, L.R.M. and Fortey, R.A. Avalonia: a long-lived terrane in the Lower Palaeozoic? In Bassett, M.G. (ed.) Early Palaeozoic Peri-Gondwana: new insights from tectonics and biogeography. Geological Society of London, Special Publications 325, 3 - 21.

Owens, R. M. and Fortey, R.A. Silicified Upper Ordovician trilobites from Pai-khoi, Arctic Russia. Palaeontology 52, 1209 -1220.

Fortey, R.A. Fossils: the key to the past (Fourth edition) Natural History Museum London (In USA as Fossils: the history of life, Sterling 256 pp).


Fortey, R.A.Trilobites of the genus Dikelokephalina from Ordovician Gondwana and Avalonia. Geological Journal (Online October 2010) Geological Journal 46: 405 - 415 (Sept 2011)

Fortey, R. A.. Charles Darwin: the scientist as hero. Contributions to Science 5(2), 183- 191. Institute of Catalan Studies.

 Fortey, R.A. Archives of life: science and collections, p. 183 - 201. In Bryson, W. (ed) Seeing further (History of The Royal Society) Harper Collins.

Fortey R.A. A naturalist becomes a scientist, in, Sabbagh, K. (ed.) A Book of Kings Third Millennium.

Fortey,R.A.  New Introduction. Pp. vii- xi The Earth: an intimate history The Folio Society, 416 pp.


Fortey, R.A., Heward, A. P., Miller, C.G. Sedimentary facies and trilobite and conodont faunas of the Ordovician Rann Formation, Ras Al Khaimah, United Arab Emirates. Geoarabia, 16. 127 - 152.

Fortey, R.A.. Lyell and deep time. Geoscientist 21, 12 -17.

Landing E. and Fortey, R.A. Tremadocian (Lower Ordovician) sea level changes and biotas on the Avalon microcontinent. Journal of Paleontology 85, 678 - 694.

Fortey, Richard . Survivors: the animals and plants that time has left behind. Harper Press, 336 pp.


Extended reviews for literary magazines include:

Surfing the lava (Review of Volcanoes, by R. Fisher et al.) London Reviw of Books October 1997.

Shock Lobsters. (Review of The Crucible of Creation by S Conway Morris). London Review of Books. October 1998.

Most curious of seas. (Review of Noah’s Flood: The new scientific discoveries about the event that changed History by W Ryan and W Pitman) LRB July 1999

Did the self-made man fake it with Bohemian fossils? (Review of The Deprat Affair by R Osborne) LRB November 1999

No Such thing as a fish (Review of Deep Time, by Henry Gee)  LRB July 2000

They might be giants (Review of The first fossil  hunters, by A Mayor) LRB November 2000

Prophet of the rocks (Review of The map that changed the world by S Winchester) LRB August 2001

America’s immigrants (The eternal frontier, by T Flannery) Lead review, Times Lit. Supp. August 2001.

Down to the last flea (Review of Mammoth, by R Stone) LRB May 2002

Everything but the glue  (Review of Victorian Sensation by J Secord)  LRB August 2002.

Archaeology is rubbish. (Review of After the Ice: a global human history  by A Mithen) LRB Dec 2003.

Decline and Fall (Review of Collapse, by Jared Diamond) The Lancet  Feb 26 2005, p. 745.

Magma’s memorabilia. (Review  of the Legacy of Dr Johnston-Lavis Collection, UCL). Times Lit. Suppl. Jan. 6 2006.

Natural progression. (Review of Charles Darwin, Geologist by Sandra Herbert) Times Literary Supplement, 13 Jan 2006.

Hammers for pipes. (Review of Bursting the Limits of Time by Martin Rudwick) London Review of Books, 9 February 2006

Review of Coral, by Steve Jones, for the Guardian, Saturday Review, April 2007.

Earth's Green heroes Nature  449 284-5 (Review of Oliver Morton’s Eating the Sun), 2007

In Good Measure (Review of A Natural History of Time, Pascal Richet) Times Literary Supplement, January pp. 3-5. 2008.

At nightmare’s edge (Review of The Earth on Show by Ralph O’Connor & Worlds before Adam by Martin Rudwick) Times Literary Supplement October 26-27 2008.

Diets of Worms (Review of Darwin’s Lost World by Martin Brasier) Times Literary Supplement 2009

Tasty butterflies (Review of Bugs and the Victorians by J.F.M. Clark)  London Review of Books 24 September 2009 26-27

Sea Slugs, wombats, microbes (Review of The Species Seekers by Richard Conniff)

London Review of Books, 11 April 2011

Literary  introductions

Introduction to the Folio Society’s reprint of The Third Policeman, by Flann O’Brien (2006)

Foreword to the Folio Society’s reprint of Life: an unauthorised biography (2007)

Mycological notes and papers

1998. Fortey, R.A. Battarraea in Oxfordshire. Mycologist 12, 159-69

1999. Fortey, R.A. Profiles of fungi. Cephalotheca suphurea. Mycologist 13, 146

2000. Fortey R.A. Old churchyards as fungus conservation areas. Field Mycology  1

2004. Fortey R.A. and Skeates, S.J. Pitptorus quercinus Field Mycology 5

2004. Fortey R A. Psilocybe aurantiaca – and a case of mistaken identity. Field Mycology 5, 77-80.

2005. Fortey, R.A. Fungi as art – Lorenzen ceramics. Field Mycology 6

2007Fortey R.A. & Ryvarden L. Ceriporiopsis herbicola nov. sp. (Polyporacea, Basidiomycota) Synopsis fungorum 23 13-15




Fortey, R.A.  2009. Autumn Foray Report, Loxton, Somerset, October 

2008. Mycologist News 2009 (2) 19-21

Fortey, R.A. 2010. Autumn Foray Report, Leiston, Suffolk, November 2009. Mycologist News 2010 (2), 5-8.