Dr Zerina Johanson

Biography

Tertiary Education

1998                    Ph.D School of Earth Sciences, Macquarie University, Sydney, Australia
                             Thesis title: Late Devonian fishes from New South Wales, with emphasis on the
                             Canowindra fauna 
1992                    M.Sc. Geology, University of Alberta, Edmonton, Alberta, Canada
                             Thesis title: A systematic revision of the North American Late Cretaceous
                             marsupial Alphadon
1989                    B.Sc. (Honours) Geology and Zoology, University of Alberta, Edmonton,
                             Alberta, Canada. Received APEGGA Medal in Geology, highest grade point 
                             average

 

Employment History

2007 - 2012       Research Curator, Department of Palaeontology, The Natural History Museum

2006 - 2007       Lecturer, School of Biological Sciences, Macquarie University, Sydney, Australia
2001 - 2005       Senior Fellow, Australian Research Council, Australian Museum, Sydney
1998 - 2000       Postdoctoral Research Fellow, Australian Research Council, Australian
                             Museum, Sydney
1994 - 2006       Palaeontologist associated with Gondwana Dreaming Pty, Canberra, involved in
                             palaeontological field excursions
1994                   Technical Officer (Palaeontology), Australian Museum
1992 - 1993       Research Assistant, Department of Geology, University of Alberta
1992 - 1993       Researcher (contract position), Archaeological Survey of Alberta
1989 - 1991       Teaching Assistant (Invertebrate Palaeontology, Vertebrate Palaeontology,
                             Introductory Geology), University of Alberta

Grants

Research

Current research

• Phylogenetic relationships among jawed vertebrates
• Evolution and development of the appendicular/axial skeleton of jawed vertebrates
• Comparative gene expression in axial and appendicular skeletons of the Australian lungfish Neoceratodus, focusing on the fin-limb transition and regionalisation of the vertebral column
• Evolution of the placoderm dentition and relationships of the Placodermi
• Evolution and development of the pufferfish dentition
• Evolution of neck musculature
• Evolution of the vertebrate dermoskeleton

Publications

Publications

2013

• ERICSSON, R., KNIGHT, R. and JOHANSON, Z. ( 2013 ) Evolution and development of the vertebrate neck Journal of Anatomy 222 : 67-78 .
• SMITH, M.M., JOHANSON, Z., UNDERWOOD, C. and DIEKWISCH, T. ( 2013 ) Pattern formation in development of chondrichthyan dentitions: a review of an evolutionary model. Historical Biology .

2012

• FRASER, G. J., BRITZ, R., HALL, A., JOHANSON, Z. and SMITH, M. M. ( 2012 ) Replacing the first generation dentition in pufferfish with a unique beak. Proceedings of the National Academy of Sciences (USA) 109(21) : 8179-8184 .
• FRIEDMAN, M. and JOHANSON, Z. ( 2012 ) †Opisthomyzon glaronensis (Wettstein, 1886) (Acanthomorpha: Echeneidae), a junior synonym of †Uropteryx elongatus Agassiz, 1844. Journal of Vertebrate Paleontology 32 : 1202-1206 .
• JOHANSON, Z., KEARSLEY, A., DEN BLAAUWEN, J., NEWMAN, M. and SMITH, M. M. ( 2012 ) Ontogenetic development of an exceptionally preserved Devonian cartilagenous skeleton. Journal of Experimental Zoology Part B, Molecular and Developmental Evolution 318 : 50-58 .
• RUECKLIN, M., DONOGHUE, P.C.J., JOHANSON, Z., TRINAJSTIC, K., MARONE, F. and  STAMPONI, M.   ( 2012 ) Development of teeth and jaws in the earliest jawed vertebrate Nature 491 : 748-751 .
• SMITH, M. M. and JOHANSON, Z. ( 2012 ) A molecular guide to evolution in the regulation of morphological pattern in development of the vertebrate dentition. Eds R. Asher and J. Mueller., In: From Clone to Bone: The Synergy of Morphological and Molecular Tools in Paleobiology . Cambridge University Press : Cambridge , 166-206 .
• TRINAJSTIC, K., LONG, J.A., JOHANSON, Z., YOUNG, G., and SENDEN, T. ( 2012 ) New morphological information on the ptyctodontid fishes (Placodermi, Ptyctodontida) from Western Australia. Journal of Vertebrate Paleontology 32 : 757-780 .

2010

• JOHANSON, Z. ( 2010 ) Evolution of paired fins and the lateral somitic frontier. Journal of Experimental Zoology Part B, Molecular and Developmental Evolution 15 : 347-352 .
• JOHANSON, Z. and AHLBERG. P.E. ( 2010 ) Phylogeny of Lungfishes. Eds J. M. Jørgensen and J. M. P. Joss, In: The Biology of Lungfishes . Science Publishers, Inc. : Enfield, New Hampshire , 43-60 .
• JOHANSON, Z., KEARSLEY, A., DEN BLAAUWEN, J. NEWMAN, M., and SMITH, M. M. ( 2010 ) No bones about it: an enigmatic Devonian fossil reveals a new skeletal framework - a potential role of loss of gene regulation. Seminars in Cell and Developmental Biology 21 : 414-423 .
• SMITH, M. M. and JOHANSON, Z. ( 2010 ) The dipnoan dentition: a unique adaptation with a longstanding evolutionary record., In: The Biology of Lungfishes. Eds J. M. Jørgensen and J. M. P. Joss . Science Publishers, Inc. : Enfield, New Hampshire , 219-247 .

2009

• AHLBERG, P. E., TRINAJSTIC, K., JOHANSON, Z.,  and LONG, J. A. ( 2009 ) Pelvic claspers confirm chondrichthyan-like internal fertilisation in arthrodires. Nature 460 : 888-889 .
• CARR, R., JOHANSON, Z. and RITCHIE, A. ( 2009 ) The phyllolepid placoderm Cowralepis mclachlani: insights into the evolution of feeding mechanisms in jawed vertebrates. Journal of Morphology 270 : 775-804 . 10.1002/jmor.10719
• HODGKINSON, V. S., ERICSSON, R., JOHANSON, Z. and JOSS, J. M. P. ( 2009 ) The apical ectodermal ridge in the pectoral fin of the Australian Lungfish (Neoceratodus forsteri): keeping the fin to limb transition in the fold. Acta Zoologica 90 (Supplement 1) : 253-263 .
• JOHANSON, Z., CARR, R. and RITCHIE, A. ( 2009 ) Vertebral development in the gnathostome stem group (Placodermi): Fusion, gene misexpression, and homeotic transformations. International Journal of Developmental Biology 54 : 71-80 .
• JOHANSON, Z., ERICSSON, R., LONG, J. A., EVANS, F. and JOSS, J. M. P. ( 2009 ) Development of the axial skeleton and median fins in the Australian lungfish, Neoceratodus forsteri. The Open Zoology Journal 2 : 91-101 .
• LONG, J. A., TRINAJSTIC, K. and JOHANSON, Z. ( 2009 ) Embryos in Devonian arthrodiran fishes inform the origins of internal fertilization in vertebrates. Nature 457 : 1094-1095 .

2008

• JOHANSON, Z., TANAKA, M., CHAPLIN, N. and SMITH, M. M. ( 2008 ) Early Palaeozoic dentine and patterned scales in the embryonic catshark tail. Royal Society, Biology Letters 4: 87-90. .
• JOHANSON, Z., TRINAJSTIC, K., BRAZEAU, M. and SMITH, M. ( 2008 ) Placoderms with teeth that bite back. Journal of Vertebrate Paleontology 28: 97A .

2007

• CARR, R. K., JOHANSON, Z. and RITCHIE, A. ( 2007 ) Fossil evidence (Pisces; Placodermi) for the paired origin of basibranchials and their derivation from neural crest. 8th International Congress of Vertebrate Morphology, Abstracts: 19 .
• CARR, R. K., JOHANSON, Z. and RITCHIE, A. ( 2007 ) Ontogeny and phylogeny of the gnathostome branchial skeleton. 40th Anniversary Symposium on Early Vertebrates/Lower Vertebrates, Uppsala. Ichthyolith Issues Special Publication 19: 24-25. .
• CARR, R. K., JOHANSON, Z., RITCHIE, A., LELIÈVRE, H. and GOUJET, D. ( 2007 ) The contributions of placoderms to our understanding of the ontogeny and evolution of early gnathostomes. 55th Symposium of Vertebrate Palaeontology and Comparative Anatomy, Glasgow, Abstracts: 11. .
• ERICSSON, R., JOHANSON, Z., STOPPER, G. and JOSS, J. ( 2007 ) The function of sonic hedgehog in the development of the pectoral fin of the Australian lungfish, Neoceratodus forsteri. 8th International Congress of Vertebrate Morphology, Abstracts: 19. .
• ERICSSON, R., JOHANSON, Z., STOPPER, G. and JOSS, J. M. P ( 2007 ) The function and expression of sonic hedgehog in the development of the pectoral fin of the Australian lungfish Neoceratodus forsteri. Euro Evo Devo 2 (2nd Meeting of the European Society for Evolutionary Developmental Biology), Abstracts: 221 .
• HODGKINSON, V., JOHANSON, Z., ERICSSON, R. and JOSS, J. M. P. ( 2007 ) Apical ectodermal ridge (AER) development in the pectoral fin of the Australian lungfish (Neoceratodus forsteri). 8th International Congress of Vertebrate Morphology, Abstracts: 55. .
• HOLLAND, T., WARREN, A., JOHANSON, Z., LONG, J. A., PARKER, K. and GARVEY, J. ( 2007 ) A new species of Barameda (Rhizodontida) and heterochrony in the rhizodontid pectoral fin Journal of Vertebrate Paleontology 27 : 295-315 . 10.1671/0272-4634(2007)27[295:ANSOBR]2.0.CO;2
• JOHANSON, Z., CARR, R. and RITCHIE, A ( 2007 ) Segmental identity within the vertebral column: what can fossils tell us? 8th International Congress of Vertebrate Morphology, Abstracts: 61; 55th Symposium of Vertebrate Palaeontology and Comparative Anatomy, Glasgow, Abstracts: 20 .
• JOHANSON, Z., CARR, R. and RITCHIE, A. ( 2007 ) Segmental identity within the vertebral column: what can fossils tell us? 8th International Congress of Vertebrate Morphology, Abstracts: 61; 55th Symposium of Vertebrate Palaeontology and Comparative Anatomy, Glasgow, Abstracts: 20. .
• JOHANSON, Z., JOSS, J., BOISVERT, C. A., ERICSSON, R., SUTIJA, M. and AHLBERG, P. E. ( 2007 ) Fish fingers: digit homologues in sarcopterygian fish fins Journal of Experimental Zoology Part B, Molecular and Developmental Evolution 308B : 757-768 . 10.1002/jez.b.21197
• JOHANSON, Z., JOSS, J., BOISVERT, C., ERICSSON, R., SUTIJA, M. and AHLBERG, P. E. ( 2007 ) Fish fingers: digit homologues in sarcopterygian fish fins. 40th Anniversary Symposium on Early Vertebrates/Lower Vertebrates, Uppsala. Ichthyolith Issues Special Publication 19: 49-50 .
• JOHANSON, Z., LONG, J. A., TALENT, J. A., JANVIER, P. and WARREN, J. W. ( 2007 ) New onychodontiform (Osteichthyes: Sarcopterygii) from the Lower Devonian of Victoria, Australia Journal of Paleontology 81 : 1034-1046 . 10.1666/pleo05-023.1
• JOHANSON, Z., SMITH M.M. and JOSS. J.M. ( 2007 ) Early scale development in Heterodontus (Heterodontiformes; Chondrichthyes): a novel chondrichthyan scale pattern Acta Zoologica 88 : 249-256 . 10.1111/j.1463-6395.2007.00276.x
• JOSS, J. M. and JOHANSON, Z. ( 2007 ) Is Palaeospondylus gunni a fossil larval lungfish? Insights from Neoceratodus forsteri development Journal of Experimental Zoology Part B, Molecular and Developmental Evolution 306 : 163-171 . 10.1002/jez.b.21125
• JOSS, J., JOHANSON, Z., HODGKINSON, V., ERICSSON, R., SUTIJA, M., STOPPER, G., COLE, N. and SORDINO, P. ( 2007 ) Development of paired fins of a lungfish, Neoceratodus forsteri informs the fin-to-limb transition in the evolution of tetrapods. 40th Anniversary Symposium on Early Vertebrates/Lower Vertebrates, Uppsala. Ichthyolith Issues Special Publication 19: 51-52 .
• SMITH, M. M., JOHANSON, Z. and JOSS, J. ( 2007 ) Filling a phylogenetic gap in teeth at the margins of lungfish dentitions. 40th Anniversary Symposium on Early Vertebrates/Lower Vertebrates, Uppsala. Ichthyolith Issues Special Publication 19: 82-83 .
• SMITH, M. M., JOHANSON, Z., OKABA, M. and JOSS, J. M. P. ( 2007 ) Building the marginal dentition of lungfish in a stereotypic osteichthyan pattern. 8th International Congress of Vertebrate Morphology, Abstracts: 119 .

2006

• AHLBERG, P. E., SMITH, M. M. and JOHANSON, Z. ( 2006 ) Developmental plasticity and extreme disparity in early dipnoan (lungfish) dentitions Evolution and Development 8 : 331-349 . 10.1111/j.1525-142X.2006.00106.x
• JOHANSON, Z. ( 2006 ) Origin of mammals: molecules, morphology and a petrosal or two. In: J. R. Merrick, M. Archer, G. Hickey & M. S. Y. Lee (eds), In: Evolution and Zoogeography of Australasian Vertebrates . AUSCIPUB Pty. Ltd : Sydney , 491-522 .
• JOHANSON, Z., LONG, J. A., TALENT, J. A., JANVIER, P. and WARREN, J. W. ( 2006 ) Oldest coelacanth-from the Early Devonian of Australia. Royal Society, Biology Letters 2: 443-446 .

2005

• GARVEY, J. M., JOHANSON, Z. and WARREN, A. ( 2005 ) Redescription of the pectoral fin and vertebral column of the rhizodontid fish Barameda decipiens from the Lower Carboniferous of Australia Journal of Vertebrate Paleontology 25 : 8-18 . 10.1671/0272-4634(2005)025[0008:ROTPFA]2.0.CO;2
• JOHANSON, Z., and SMITH, M. M ( 2005 ) Origin and evolution of gnathostome dentitions: a question of teeth and pharyngeal denticles in placoderms Biological Reviews 80 : 303-345 . 10.1017/S1464793104006682
• JOHANSON, Z., BURROW, C., WARREN, A. and GARVEY, J. ( 2005 ) Homology of the fin lepidotrichia in osteichthyan fishes Lethaia 38 : 27-36 . 10.1080/00241160510013141
• JOHANSON, Z., LONG, J. A., TALENT, J. A., JANVIER, P. and WARREN, J. W. ( 2005 ) Diverse new fish fauna from the Lower Devonian of Victoria, Australia. 10th Conference on Australasian Vertebrate Evolution, Palaeontology & Systematics, Programme and Abstracts: 43 .
• JOHANSON, Z., SUTIJA, M. and JOSS, J. M. P. ( 2005 ) Regionalization of the axial skeleton in the lungfish Neoceratodus forsteri (Dipnoi) Journal of Experimental Zoology Part B, Molecular and Developmental Evolution 304 : 229-237 .
• PARKER, K., WARREN, A. and JOHANSON, Z. ( 2005 ) Strepsodus (Rhizodontida; Sarcopterygii) pectoral elements from the Lower Carboniferous Ducabrook Formation, Queensland, Australia Journal of Vertebrate Paleontology 25 : 46-62 . 10.1671/0272-4634(2005)025[0046:SRSPEF]2.0.CO;2
• SMITH, M. and JOHANSON, Z. ( 2005 ) Dynamics of tooth addition and dental tissues in placoderm dentitions. 10th Conference on Australasian Vertebrate Evolution, Palaeontology & Systematics, Programme and Abstracts: 70 .

2004

• JOHANSON, Z. ( 2004 ) Late Devonian sarcopterygian fishes from eastern Gondwana (Australia and Antarctica) and their importance in phylogeny and biogeography. In: G. Arratia, R. Cloutier & M. V. H. Wilson (eds), In: Recent Advances in the Origin and Early Radiation of Vertebrates . Verlag Dr. F. Pfeil : München , 285-306 .
• JOHANSON, Z. and SMITH, M. M. ( 2004 ) Denticles and teeth at the origin of jaws. 7th International Congress of Vertebrate Morphology. Journal of Morphology 260: 302-303 .
• JOHANSON, Z., JANVIER, P., LONG, J. A., TALENT, J. A. and WARREN, J. ( 2004 ) New fishes from the Devonian of Victoria, Australia. 10th International Symposium on Early Vertebrates/Lower Vertebrates Gramado, Brazil. .
• JOHANSON, Z., JOSS, J. and SUTIJA, M., ( 2004 ) Scapulocoracoid homology among bony fishes based on morphology and development of the Queensland lungfish, Neoceratodus forsteri (Sarcoptergygii). 7th International Congress of Vertebrate Morphology. Journal of Morphology 260: 303. .
• JOHANSON, Z., JOSS, J. M. P and WOOD, D. ( 2004 ) Scapulocoracoid development in the Queensland lungfish Neoceratodus forsteri (Dipnoi; Sarcopterygii) Zoology 107 : 93-110 . 10.1016/j.zool.2004.01.001
• SMITH, M. M., JOHANSON, Z. and LELIEVRE, H. ( 2004 ) Teeth and denticles in arthrodire dentitions, an invasive and pervasive solution. 10th International Symposium on Early Vertebrates/Lower Vertebrates Gramado, Brazil .

2003

• JOHANSON, Z. ( 2003 ) The clavobranchialis musculature in sarcopterygian fishes, and contribution to osteichthyan feeding and respiration Contributions to Zoology 72 : 17-37 . Contributions to Zoology
• JOHANSON, Z. ( 2003 ) Placoderm branchial and hypobranchial muscles and origins in jawed vertebrates Journal of Vertebrate Paleontology 23 : 735-749 . 10.1671/2
• JOHANSON, Z. and SMITH, M. M. ( 2003 ) Placoderm fishes, pharyngeal denticles and the vertebrate dentition Journal of Morphology 257 : 289-307 . 10.1002/jmor.10124
• JOHANSON, Z., AHLBERG, P. E. and RITCHIE, A. ( 2003 ) The braincase and palate of the tetrapodomorph sarcopterygian Mandageria fairfaxi: morphological variability near the fish-tetrapod transition Palaeontology 46 : 271-293 . 10.1111/1475-4983.00298
• SMITH, M. M. and JOHANSON, Z. ( 2003 ) Separate evolutionary origins of teeth within jawed vertebrates Science 299 : 1235-1236 . 10.1126/science.1079623
• SMITH, M. M. and JOHANSON, Z. ( 2003 ) Response to Comment on "Separate Evolutionary Origins of Teeth from Evidence in Fossil Jawed Vertebrates" Science 300 : 1661 . 10.1126/science.1084686

2002

• JOHANSON, Z ( 2002 ) Late Devonian sarcopterygian fishes from Eastern Gondwana (Australia and Antarctica) and their importance in phylogeny and biogeography. Journal of Vertebrate Paleontology 22: 72A .
• JOHANSON, Z. ( 2002 ) Vascularization of the osteostracan and antiarch (Placodermi) pectoral fin: similarities, and implications for placoderm relationships Lethaia 35 : 169-186 . 10.1080/002411602320184024
• JOHANSON, Z. and SMITH, M. M. ( 2002 ) Placoderm fishes and the vertebrate oropharyngeal dentition. First International Palaeontological Congress, Sydney 2002. Geological Society of Australia, Abstracts 68: 84 .
• SMITH, M. M. and JOHANSON, Z. ( 2002 ) Dentition characters as a contribution to gnathostome and placoderm phylogeny. First International Palaeontological Congress, Sydney 2002. Geological Society of Australia, Abstracts 68: 145 .

2001

• AHLBERG, P. E., JOHANSON, Z. and DAESCHLER, E. B. ( 2001 ) The Late Devonian lungfish Soederberghia (Sarcopterygii, Dipnoi) from Australia and North America, and its biogeographical implications Journal of Vertebrate Paleontology 21 : 1-12 . 10.1671/0272-4634(2001)021[0001:TLDLSS]2.0.CO;2
• JOHANSON, Z. and AHLBERG, P. E. ( 2001 ) Devonian rhizodontids (Sarcopterygii; Tetrapodomorpha) from East Gondwana Transactions of the Royal Society of Edinburgh: Earth Sciences 92 : 43-74 . 10.1017/S0263593300000043

2000

• JOHANSON, Z. and RITCHIE, A. ( 2000 ) A new Late Famennian lungfish from New South Wales, Australia, and its bearing on Australian-Asian terrane relations Alcheringa 24 : 99-118 . 10.1080/03115510008619527
• JOHANSON, Z. and RITCHIE, A. ( 2000 ) Rhipidistians (Sarcopterygii) from the Hunter Siltstone (Late Famennian) near Grenfell, NSW, Australia Mitteilungen aus dem Museum für Naturkunde in Berlin, Geowissenschaftliche Reihe 3 : 111-136 . 10.1002/mmng.20000030107
• JOHANSON, Z. and RITCHIE, A. ( 2000 ) Sarcopterygian fishes from the Hunter Siltstone (Late Fammenian) near Grenfell, NSW, Australia. Where Worlds Collide: Faunal and floral migrations and evolution in SE Asia-Australasia, University of New England, Abstracts: 37 .
• JOHANSON, Z. and RITCHIE, A. ( 2000 ) Sarcopterygian fishes from the Hunter Siltstone (Late Fammenian) near Grenfell, NSW, Australia. Palaeontology Down Under 2000. Geological Society of Australia, Abstracts 61: 44 .
• JOHANSON, Z., TURNER, S., and WARREN, A. ( 2000 ) First East Gondwana record of Strepsodus, from the Early Carboniferous Ducabrook Formation, Central Queensland Geodiversitas 22 : 161-169 .

1999

• JOHANSON, Z. and YOUNG, G. C. ( 1999 ) New Bothriolepis (Antiarchi; Placodermi) from the Braidwood region, New South Wales, Australia (Middle-Late Devonian) Records of the Western Australian Museum Supplement 57 : 55-76 .

1998

• AHLBERG, P. E. and JOHANSON, Z. ( 1998 ) Osteolepiforms and the ancestry of tetrapods Nature 395 : 792-794 . 10.1038/27421
• JOHANSON, Z. ( 1998 ) The Upper Devonian fish Bothriolepis (Placodermi: Antiarchi) from near Canowindra, New South Wales, Australia Records of the Australian Museum 50 : 315-348 .
• JOHANSON, Z. and AHLBERG, P. E. ( 1998 ) A complete primitive rhizodont from Australia Nature 394 : 569-573 . 10.1038/29058

1997

• AHLBERG, P. E. and JOHANSON, Z. ( 1997 ) The second tristichopterid (Sarcopterygii, Osteolepiformes) from the Upper Devonian of Canowindra, New South Wales, Australia Journal of Vertebrate Paleontology 17 : 653-673 . Journal of Vertebrate Paleontology
• AHLBERG, P. E. and JOHANSON, Z. ( 1997 ) The interrelationships of the Osteolepiformes. Journal of Vertebrate Paleontology 17: 28A .
• JOHANSON, Z. ( 1997 ) New Bothriolepis and Remigolepis (Placodermi; Antiarchi) from the Hunter Siltstone (Upper Famennian) near Grenfell, NSW Alcheringa 21 : 197-217 . 10.1080/03115519708619173
• JOHANSON, Z. and AHLBERG, P. E. ( 1997 ) New tristichopterid (Osteolepiformes; Sarcopterygii) from the Mandagery Sandstone (Famennian) near Canowindra, N.S.W., Australia Transactions of the Royal Society of Edinburgh: Earth Sciences 88 : 39-53 .

1996

• JOHANSON, Z. ( 1996 ) Revision of the North American Late Cretaceous North American marsupial genus Alphadon Palaeontographica A 242 : 127-184 . Palaeontographica A
• JOHANSON, Z. ( 1996 ) New marsupial from the Fort Union Formation, Swain Quarry, Wyoming Journal of Paleontology 70 : 1023-1031 .

1995

• JOHANSON, Z. ( 1995 ) New information concerning Albertatherium Fox, 1971 Journal of Vertebrate Paleontology 14 : 595-602 . Journal of Vertebrate Paleontology
• JOHANSON, Z. ( 1995 ) New information on jaw elements of Remigolepis (Placodermi; Antiarchi), from Canowindra, N.S.W., Australia. (Upper Devonian) Geobios, Mémoire spécial 19 : 103-108 . 10.1016/S0016-6995(95)80095-6

1994

• CHATTERTON, B. D. E., JOHANSON, Z. and SUTHERLAND, G. ( 1994 ) The digestive system of Trilobita - alimentary structures in Pterocephalia Journal of Paleontology 68 : 294-304 . Journal of Paleontology
• CIFELLI, R. C. and JOHANSON, Z ( 1994 ) New marsupial from Utah Journal of Vertebrate Paleontology 14 : 292-295 . Journal of Vertebrate Paleontology

1993

• JOHANSON, Z. ( 1993 ) A revision of the Late Cretaceous (Campanian) marsupial Iqualadelphis lactea Fox, 1987 Journal of Vertebrate Paleontology 13 : 373-377 . Journal of Vertebrate Paleontology

1992

• JOHANSON, Z. ( 1992 ) New information concerning the Late Cretaceous marsupial genus Albertatherium Fox, 1971. Journal of Vertebrate Paleontology 12: 35A .

1991

• JOHANSON, Z. ( 1991 ) A revision of Alphadon marshi Simpson, 1927 and Alphadon wilsoni Lillegraven, 1969 (Marsupialia). Journal of Vertebrate Paleontology 11: 38A .