Publications
1968
Vallance, G., and Fortey, R.A. 1968. Ordovician succession in North Spitsbergen. Proc. Geol. Soc London 1648, 91-7.
1971
Fortey, R.A. Tristichograptus, a triserial graptolite from the Lower Ordovician of Spitsbergen. Palaeontology, 14 : 188-199, pls 26 - 29.
1972
Fortey, R.A. & Holdsworth, B.K. The oldest well-preserved radiolaria. Boll. Soc. Paleont. Ital. 10 : 35-41, 2 pls.
1973
Fortey, R.A. & Bruton, D.L. Cambrian-Ordovician rocks adjacent to Hinlopenstretet, North Ny Friesland, Spitsbergen. Bull. geol. Soc. Am. 84 : 2227 - 2242.
1974
Fortey, R.A. The Ordovician trilobites of Spitsbergen. I. Olenidae. Skr. norsk Polarinst. 160: 1-81, 24 pls.
Fortey, R.A. A new pelagic trilobite from the Ordovician of Spitsbergen, Ireland and Utah. Palaeontology, 17 : 111 - 124, pls 13,14.
1975
Fortey, R.A. Early Ordovician trilobite communities. Fossils Strata 4 : 331-352.
Fortey, R.A. and Owens, R.M. Proetida - a new order of trilobites Fossils Strata 4: 227-239.
Fortey,R.A. The Ordovician trilobites of Spitsbergen.II. Asaphidae, Nileidae, Raphiophoridae and Telephinidae of the Valhallfonna Formation. Skr. norsk. Polarinst. 162: 1 - 125, 41 pls.
1976
Fortey,R.A. Correlation of shelly and graptolitic early Ordovician successions, based on the sequence in Spitsbergen. 263 - 280 in Bassett,M.G. (ed.) The Ordovician System. University of Wales Press and National Museum of Wales, Cardiff.
Fortey,R.A. & Clarkson,E.N.K. The function of the glabellar `tubercle' in Nileus and other trilobites. Lethaia 9: 101 -106.
Fortey,R.A. and Whittaker,J.E.P. Janospira - an Ordovician microfossil in search of a phylum. Lethaia 9: 397 - 403.
Fortey,R.A. & Rushton,A.W.A. Chelidonoceaphalus trilobite fauna from the Cambrian of Iran. Bull. Br. Mus. nat. Hist. (Geol.) 27: 321 - 340, pls 8 -12.
Bockelie,T. and Fortey,R.A. An early Ordovician vertebrate. Nature 260 : 36 - 38.
Bockelie, T.G., Bruton, D.L. and R.A. Fortey. Research on the Ordovician rocks of North Ny Friesland, Spitsbergen. Arbok Norsk Polarinstitutt (for 1975), 214 - 7.
Morris,N.J. and Fortey,R.A. Tironucula gen. nov. and its significance in bivalve evolution. J. Paleont. 50: 701 - 709, pl.1.
1977
Fortey,R.A. & Morris,S.F. Variation in lens packing of Phacops (Trilobita) Geol. Mag. 114: 25 - 32.
Fortey,R.A. & Barnes,C.R. Early Ordovician conodont and trilobite communities of Spitsbergen: influence on biogeography. Alcheringa 1: 297 - 309.
Shaw,F.C. and Fortey,R.A. Middle Ordovician facies and trilobite faunas in N. America. Geol. Mag. 114: 409 - 443.
1978
Fortey,R.A. & Owens,R.M. Early Ordovician (Arenig) stratigraphy and faunas of the Carmarthen District, south-west Wales. Bull. Br. Mus. nat. Hist. (Geol.) 30: 225 - 294, 11 pls.
Fortey, R.A. and Morris,S.F. Discovery of nauplius-like trilobite larvae. Palaeontology 21: 823 - 833, pl.94.
Cope,J.C.W., Fortey,R.A. & Owens,R.M. Newly discovered Tremadoc rocks in the Camarthen District, South Wales. Geol. Mag. 115: 195 -198.
1979
Fortey,R.A. Early Ordovician trilobites from the Catoche Formation (St. George Group ), western Newfoundland. Bull. Geol. Surv. Can. 321: 61 - 114. 15 pls.
Fortey,R.A. & Owens,R.M. Enrollment in the classification of trilobites. Lethaia 12: 219 - 226.
Rushton, A.W.A., Fortey, R.A. and Owens, R.M. Excavation of two candidate sections for stratotypes in Wales by the Nature Conservancy Council. Earth Science Conservation : 16: 1 - 5.
1980
Fortey, R.A. The Ordovician trilobites of Spitsbergen.III. Remaining trilobites of the Valhallfonna Formation. Skr. norsk. Polarinst. 171: 1 - 113, 25 pls.
Fortey, R.A. The Ordovician of Spitsbergen and its relevance to the base of the Middle Ordovician in North America. 33 - 40, In Wones,D.R. (ed.) The Caledonides in the U.S.A. IGCP Project 27, 329, Blacksburg, Va.
Fortey, R.A. Generic longevity in Lower Ordovician trilobites: relation to environment. Paleobiology 6: 24 - 31.
Fortey, R.A. & Skevington,D. Correlation of the Cambrian-Ordovician boundary between Europe and North America: new data from western Newfoundland. Can. J. Earth. Sci. 17: 382 - 388.
Fortey, R.A. & Rushton,A.W.A. Acanthopleurella Groom 1902: origin and life habits of a miniature trilobite. Bull. Br. Mus. nat. Hist. (Geol.) 33: 79 - 89
Fortey, R.A. Basilicus tyrannus (Murchison) and the glabellar structure of asaphid trilobites. Bull. Br. Mus. nat. Hist. (Geol.) 34: 225 - 264.
1981
Fortey, R.A. Prospectatrix genatenta (Stubblefield) and the trilobite superfamily Cyclopygacea. Geol. Mag. 118: 603 - 614.
Fortey, R.A. The Burgess Shale: a unique Cambrian fauna. Nature 293: 189
1982
Fortey, R.A. & Jefferies, R.P.S. Fossils and phylogeny: a compromise approach. 197 - 234, In Joysey,K.A. & Friday,A.E. (eds.) Problems of Phylogenetic reconstruction. Academic Press.
Fortey, R.A., Landing,E. & Skevington,D. Cambrian-Ordovician boundary sections in the Cow Head Group, western Newfoundland. 95 - 129, In Basset, M.G. and Dean,W.T. (eds.) The Cambrian-Ordovician boundary. National Museum of Wales, Cardiff.
Fortey, R.A. & Morris,S.F. The Ordovician trilobite Neseuretus from Saudi Arabia and the palaeography of the Neseuretus fauna related to Gondwana in the earlier Ordovician. Bull. Br. Mus. nat. Hist. (Geol.) 36: 63 -75.
Fortey, R.A. & Peel,J.S. Fossils from the Poulsen Cliff Formation, Washington Land, western North Greenland. Gronlands geol. Unders. 108: 24.
Fortey, R.A. Fossils: the key to the past. 179 pp. William Heinemann Ltd., London, and British Museum (Natural History). (U.S. edition Van Nostrand Rheinhold, New York)
Briggs, D.E.G. & Fortey, R.A. The cuticle of aglaspid arthropods: a red herring in the early history of the vertebrates. Lethaia 15: 25 - 29.
Owens, R.M. & Fortey, R.A. Arenig rocks of the Carmarthen-Llanarthney district. 249 - 258, In Bassett,M.G. (ed.) Geological Excursions in Dyfed, south-west Wales. Geologists Association.
Owens, R.M., Fortey, R.A., Cope,J.C.W., Rushton,A.W.A. & Bassett,M.G. Tremadoc faunas from the Carmarthen district, South Wales. Geol. Mag. 119:1 - 38, 8 pls.
Cocks, L.R.M. & Fortey, R.A. Faunal evidence for oceanic separations in the Palaeozoic of Britain. J. Geol. Soc. 139:465 - 478.
Cooper, R.A. & Fortey, R.A. The Ordovician graptolites of Spitsbergen. Bull. Br. Mus. nat. Hist. (Geol.) 36:157 - 302, 6 pls.
1983
Fortey, R.A. Cambrian-Ordovician boundary trilobites from western Newfoundland and their phylogenetic significance. Spec. Pap. Palaeontology. 30: 179 - 211, 23-27.
Fortey, R.A. & Peel,J.S. The anomalous bathyurid trilobite Ceratopeltis and its homoemorphs. ibid: 51 - 57, pl.6.
Cooper, R.A., & Fortey,R.A. Development of the graptolite rhabdosome. Alcheringa 7: 201 - 221.
Fortey, R.A. Geometrical constraints in the construction of graptolite stipes. Paleobiology 9: 116 -125.
1984
Fortey, R.A. Global Ordovician transgressions and regressions and their biological implications.37-50, In Bruton,D.L. (ed.) Aspects of the Ordovician System. Universitetsforlaget, Oslo.
Fortey, R.A. & Owens, R.M. A synopsis of the Arenig Series in South Wales. Proc. Geol. Assoc. 95, 389 - 90.
Whittington, H.B., Dean,W.T., Fortey,R.A., Rickards,R.B., Rushton,A.W.A. & Wright,A.D. Definition of the Tremadoc Series and the series of the Ordovician System in Britian. Geol. Mag. 121: 17 - 33.
Fortey,R.A. & Shergold,J.S. Early Ordovician trilobites from the Nora Formation, central Australia. Palaeontology 27: 315 - 366, pls.38 - 46.
1985
Fortey, R.A. Gradualism and punctuated equilibria as competing and complementary theories. Spec. Pap. Palaeontology 33: 17-28.
Fortey, R.A. Pelagic trilobites as an example of deducing the life habits of extinct arthropods. Trans. R. Soc. Edinb. 76 : 219-30.
Thomas, A.T. & Fortey, R.A. Trilobita, In, Murray, J.W. (ed.) Atlas of Invertebrate Macrofossils. Longman's, London.
Morris, S.F. & Fortey, R.A. Catalogue of the type and figured specimens of Trilobita in the British Museum (Natural History). British Museum (Natural History), 183 pp, 3 pls.
1986
Fortey, R.A. & Cooper, R.A. A phylogenetic classification of the graptoloids. Palaeontology 29 : 631-654, 11 figs.
Fortey, R.A. The type species of the Ordovician trilobite Symphysurus: systematics, functional morphology and terrace ridges. Palaont. Z. 60, 255-275, 12 figs.
Fortey, R.A. & Cocks, L.R.M. Introduction: Fossils and tectonics. J. geol. Soc. 143, 149-150.
Fortey, R.A. & Cocks, L.R.M. Marginal faunal belts and their structural implications, with examples from the Lower Palaeozoic. J. geol. Soc. 143, 151-160, 7 figs.
Fortey,R.A. The course of evolution. Pp. 66-77, in Hallam, A. & Berry, R.J. (eds) The Collins Encyclopedia of Animal Evolution. Collins, London.
Zhou, Zhiyi and Fortey, R.A. Ordovician trilobites from North and Northeast China. Palaeontographica Abt. A 192, 157-210, 12 pls.
Fortey, R.A. Early Ordovician trilobites from the Wandel Valley Formation, eastern North Greenland. Rapp. Gronlands geol. Unders.132, 15-25.
Cocks, L.R.M. & Fortey, R.A. New evidence on the South African Lower Palaeozoic: age and fossils reviewed. Geol. Mag. 123, 437-444, 1 fig.
1987
Fortey, R.A. and Bell, A. Branching geometry and function of multiramous graptolites. Paleobiology 13, 1-19, 21 figs.
Fortey, R.A. and Owens, R.M. The Arenig Series in South Wales. Bull. Br. Mus. nat. Hist. (Geol.) 41 , 69-307, 135 figs.
1988
Fortey, R.A. and Chatterton, B.D.E. Classification of the trilobite suborder Asaphina. Palaeontology 31, 165-222, pls 17-19.
Fortey, R.A. Seeing is believing: gradualism and punctuated equilibria in the fossil record. Sci. Prog. Oxf. 72, 1-19.
Fortey, R.A. and Cocks, L.R.M. Arenig to Llandovery faunal distributions in the Caledonides. In Harris, A. L. & Fettes, D.J. (eds) The Caledonian - Appalachian Orogen, 233-246. Geol. Soc. Spec. Pub. 38.
Fortey, R.A. The Ordovician trilobite Hadrohybus Raymond 1925, and its family relationships. Postilla 202, 1-7.
Fortey, R.A. The international correlation of the lower part of the Ordovician and how it should influence Laurentian chronostratigraphy. Bulletin N.Y.St. Mus. 462, 43 - 47.
Cocks, L.R.M. and Fortey, R.A. Lower Palaeozoic facies and faunas around Gondwana. Geol. Soc. Spec. Pub. 37, 183-200.
Briggs, D.E.G., Fortey, R.A. & Clarkson, E.N.K. Extinction and the fossil record of the arthropods. In Larwood, G.P. (ed.) Extinction and survival in the fossil record, 171 - 209. Systematics Association Special Volume No. 34. Clarendon Press, Oxford.
1989
Fortey, R.A., Owens, R.M., and Rushton, A.W.A. The palaeogeographic position of the Lake District in the early Ordovician. Geol. Mag.126, 9 - 17, 2 figs.
Fortey, R.A. and Whittington, H.B. The Trilobita as a natural group. Historical Biology 2, 125 -138.
Fortey, R.A. Genetics, Paleontology and Macroevolution, by J.Levinton, essay review in Historical Biology 2, 179 -182.
R. A. Fortey. The collection connection (Review of Wonderful Life by S.J. Gould). Nature 342, 303.
Briggs, D.E.G. & Fortey, R.A. The early radiation of the Arthropoda, Science, N.Y., 246, 241 - 243.
Fortey, R.A. There are extinctions and extinctions: examples from the Lower Palaeozoic. Phil. Trans. Roy. Soc. ser. B 325, 327 - 355
Fortey, R.A. A Devonian trilobite fauna from Thailand. Alcheringa 13, 257 - 267
Fortey, R.A. & J.S. Peel. Stratigraphy and hystricurid trilobites of the Christian Elv Formation (Lower Ordovician) of western North Greenland. Rapp. Groenlands geol. Unders 144, 5 - 15.
1990
Fortey, R.A. Overview: Analysis of taxonomy and phylogeny. In Briggs, D.E.G. & Crowther, P.R. (eds) Palaeobiology: a synthesis, 425-427. Blackwell, Oxford.
Fortey, R.A. & Cocks, L.R.M. Fossils and tectonics. In Briggs, D.E. G. and Crowther, P. R. (eds) Palaeobiology: a synthesis 482-492. Blackwell, Oxford.
Fortey, R.A. Ontogeny, hypostome attachment and trilobite classification. Palaeontology 33 (3), 529-576.
Cocks, L.R.M. & Fortey, R.A. Biogeography of Ordovician and Silurian faunas, pp. 97-104, In McKerrow, W.S. & Scotese, C.R. (eds) Palaeozoic Palaeogeography and Biogeography. Geological Society Memoir No. 12.
Fortey, R.A. & Peel, J.S. Early Ordovician trilobites and molluscs from the Poulsen Cliff Formation, Washington Land, western North Greenland. Bull. geol. Soc. Denmark 38, 11-32.
Fortey, R.A. & Owens, R.M. Arenig biostratigraphy and correlation in the Welsh Basin. J. Geol. Soc. London 147: 607-610.
Fortey, R.A., Beckly, A.J. & Rushton, A.W.A. International correlation of the base of the Llanvirn Series, Ordovician System. Newsletters in Stratigraphy 22: 119-142.
Fortey, R.A. & Owens, R.M. Trilobites. Chapter 5 in McNamara, K.J. (ed.) Evolutionary Trends, pp. 121 - 142. Belhaven Press, London.
Fortey, R.A. Trilobite evolution and systematics. Pp. 44-65, In Arthropod Paleobiology. Short courses in Paleontology No. 3, The Paleontological Society of America
Fortey, R.A. & Owens, R.M. Evolutionary radiations in the Trilobita. Pp. 139 -164, In Taylor, P.D. & Larwood, G.P. (eds) Major evolutionary radiations Systematics Association Special Volume 42. Clarendon Press Oxford.
1991
Cooper, R.A., Fortey, R.A. & Lindholm, K. Latitudinal and depth zonation of early Ordovician graptolites. Lethaia 24: 199 - 218.
Fortey, R.A. & Owens, R.M. The early Ordovician trilobite Beltella. Proc. Bristol nat. Hist. Soc. 49: 69 -79, 1 pl.
Fortey, R.A. Fossils: the key to the past. 2nd edition. 187 pp. Natural History Museum Publications and Harvard University Press (US).
Fortey, R.A. & Owens, R.M. A trilobite fauna from the highest Shineton Shales in Shropshire, and the correlation of the latest Tremadoc. Geol. Mag. 122: 437 - 464.
Fortey, R.A. & Wilmot, N.V. Trilobite cuticle thickness in relation to palaeoenvironment. Pal. Zeitschr. 65: 141 -151.
Chapters 5 (S. Rigby and R.A. Fortey), 23 - 32, and 12 (R.A. Fortey, I. Strachan, R.B. Rickards), 69 -77, In Rickards, R.B. and Palmer, D. (Eds) Graptolites : Writing in the rocks. Boydell Press.
Fortey, R.A., Bassett, M.G., Harper, D.A.T., Hughes, R.A., Ingham, J.K., Molyneux, S.G., Owen, A.W., Owens, R.M., Rushton, A.W.A. & Sheldon, P.R. Progress and problems in the selection of stratotypes for the bases of series in the Ordovician System of the historical type area in the U.K. Pap. Geol. Surv. Canada 90 - 9: 5 -25.
Fortey, R.A. Wales and the Ordovician System. Geology Today 7: 136 -138.
1992
Fortey, R.A. Taxonomy and extinction patterns. In, The Unity of Evolutionary Biology. Proceedings of the International Congress of Evolutionary & Systematic Biology, University of Maryland, 1990, pp. 82 - 88.
Fortey, R.A. Ordovician trilobites from the Durness Group, North-West Scotland, and their palaeobiogeography. Scot. J. Geol. 28: 115 - 121.
Fortey, R.A. & Mellish, C.J.T. Are some fossils better than others for inferring palaeogeography? Terra Nova 4: 210 - 216.
Fortey, R.A. South Britain, Tremadoc and Arenig (with A.W.A. Rushton) and Llanvirn (with R.E. Bevins) in, Atlas of Palaeogeography and lithofacies. (J.C.W. Cope, P.F. Rawson and J.K. Ingham [eds) Mem. Geological Society of London No. 13.
Fortey, R.A. & Cocks, L.R.M. The Ordovician paleogeography of the North Atlantic region as a test case in the use of fossils for continental reconstruction. Tectonophysics 206: 147 -158.
Briggs, D.E.G. and Fortey, R.A. The early Cambrian radiation of the arthropods, In Lipps, J.H. & Signor, P.W. (eds). Origin and early evolution of the Metazoa, p. 335 - 373. Plenum Press, N.Y.
Briggs, D.E.G., Fortey, R.A. and Wills M. Morphological disparity in the Cambrian. Science N.Y. 256: 1670-1673.
Fortey, R.A. & Owens, R.M. The Habberley Formation: youngest Tremadoc in the Welsh Borderlands. Geol. Mag. 129: 553 - 566.
Fortey, R.A., Pankhurst, R.J. & Herve Ç, F. 1992. Devonian trilobites at Buill, Chile (42 S). Revista Geol. Chile 19: p. 133 -144.
Fortey, R.A. & Owens, R.M. The trilobite Angelina unstretched. Geology Today 8: 219 - 221.
Briggs, D.E.G., Fortey, R.A. and Wills, M.A. Cambrian and Recent morphological disparity: response to Foote and Gould, and Lees. Science 258: 1817 - 18.
1993
Briggs, D.E.G., Fortey, R.A. and M. Wills. How big was the Cambrian evolutionary explosion?A taxonomic and morphological comparison of Cambrian and Recent arthropods, pp. 33 - 44 in, Evolutionary Patterns and Processes. Linnean Society of London.
Fortey, R.A. and Thomas, R.E. The case of the velvet worm. Nature, 361, 205 - 6.
Fortey, R.A. Charles Lapworth and the biostratigraphic paradigm. Journal of the Geological Society 150: 209 - 218.
Fortey, R.A. The Hidden Landscape: a journey into the geological past. Jonathan Cape, London, 310 pp. (Reprinted as paperback, by Pimlico Press, 1994)
Fortey, R.A. in, Romano, M. and others: Arthropoda (Trilobita) In, The Fossil Record 2 (M.J. Benton, ed.): 279 - 296. Chapman & Hall.
1994
Fortey, R.A. Late Cambrian trilobites from the Sultanate of Oman. Neues Jahrbuch fur Geologie und Palaontologie 194: 25 - 53
Wills, M.A., Briggs, D.E.G. and Fortey, R.A. Disparity as an evolutionary index: a comparison of Cambrian and Recent arthropods. Paleobiology 20: 93 -130.
1995
Fortey R.A. & Theron, J. A new Ordovician arthropod Soomaspis and the agnostid problem. Palaeontology 37: 841 - 861.
Fortey R.A., Harper, D.A.T,. Ingham, J.K., Owen, A.W. and Rushton, A.W.A. A revision of Ordovician series and stages from the historical type area. Geological magazine 132: 15 - 30
Fortey, R.A. Trilobite life habits and Lower Palaeozoic continents. Geoscientist, 5, 16 -17.
Charles Lapworth and the biostratigraphic paradigm, in Le Bas, M.J. (ed.) Milestones in Geology. Memoir of the Geological Society 16, 93 -102.
Barnes, C..R., Fortey, R.A. and Williams, S.H. The Pattern of Global Bio-events during the Ordovician Period, in Walliser O. (ed.) Global Events and Event Stratigraphy, 113 - 139. Springer-Verlag, Berlin.
Wills, M.A., Briggs, D.E.G., Fortey, R.A., and Wilkinson, M. The significance of fossils in understanding arthropod evolution. Verhandlungen der Deutschen Zoologischen Gesellschaft, 88, 203 - 215.
Fortey, R.A. The Ordovician Series of the Historical Type Area: revision as a contribution to their utility in international correlation. In Ordovician Odyssey: Short papers for the Seventh International Symposium on the Ordovician System, 11 - 14, Pacific Section SEPM Book 77.
Hughes N. C. and Fortey, R.A. Sexual dimorphism in trilobites, with an Ordovician case study. Ibid. 419 - 422.
Edgecombe, G.D. and Fortey, R.A. Brianurus new genus, a replacement name for Briania Chasen and Kloss, 1930 (Aves) [pro Briania Edgecombe 1994 (Trilobita) preoccupied). Journal of Paleontology 69 (3), 615.
1996
Fortey, R.A. and Droser, M.L. A trilobite fauna from the base of the Middle Ordovician, Basin ranges, western USA Journal of Paleontology, 70, 73 - 99.
Droser, M.L., Fortey, R.A. and Xing, L. The Ordovician Radiation. American Scientist 84: 122 - 131 (and cover).
Fortey, R.A., Briggs, D.E.G. and M. A. Wills. The Cambrian evolutionary 'explosion': decoupling cladogenesis from morphological disparity. Biological Journal of the Linnean Society 57: 13 - 33.
1997
Fortey, R.A., Briggs, D.E.G. and Wills, M.A. The Cambrian evolutionary 'explosion' recalibrated. Bioessays 19: 429 - 434.
Fortey, Richard. “Life: an unauthorised biography” Harper Collins, London 400pp
Fortey, R.A. & Owens, R.M. Bubble-headed trilobites, and a new olenid example. Palaeontology 40: 451 - 460.
Fortey, R.A. Late Ordovician trilobites from southern Thailand. Palaeontology 40: 397 - 450.
Landing, E., Bowring, S.A., Fortey, R.A., and Davidek, K.L. U-Pb date from Avalonian Cape Breton Island and geochonologic calibration of the Early Ordovician. Canadian Journal of Earth Sciences 34: 724 - 730.
Cocks, L.R.M. and Fortey, R.A. A new Hirnantia fauna from Thailand and the biogeography of the latest Ordovician of South-east Asia. Geobios M.S. No. 20, 117 - 126.
Fortey R.A. Classification, in Treatise on Invertebrate Paleontology. Part O (Arthropoda 1, revised), 289 - 302.
Fortey, R.A. & Owens, R.M. Evolutionary History, Ibid. 249 - 288.
Fortey, R.A. & Thomas, R.H. (eds) Arthropod Relationships. Chapman & Hall, London, 383 pp. Systematics Association Special Volume Number 55.
Wills, M.A., Briggs, D.E.G. and Fortey, R.A. Evolutionary correlates of arthropod tagmosis: scrambled legs, 57 - 65 in Arthropod Relationships
Austin, J.J., Ross, A.J., Smith, A,. B., Fortey, R.A. & Thomas, R.H. Problems of reproducibility - does geologically ancient DNA survive in amber-preserved insects? Proceedings of the Royal Society of London B 264: 467 - 474.
Fortey R.A. & Seilacher, A. The trace fossil Cruziana semiplicata and the trilobite that made it. Lethaia 30:105 -112.
Adrain, J.M. and Fortey, R.A. Ordovician trilobites from the Tourmakeady Limestone, western Ireland. Bulletin of the British Museum (Natural History) Geology series, 53:79 - 115.
1998
McCormick, T. and Fortey, R.A. Independent testing of a paleobiological hypothesis: the optical design of two Ordovician pelagic trilobites reveals their relative paleobathymetry. Paleobiology 24: 235 - 253.
L.R.M. Cocks and Fortey, R.A. The Lower Palaeozoic margins of Baltica. Geol. Foren. Forh. 120. 173 - 179.
Wills, M.A., Briggs, D.E.G., Fortey, R.A, et al. An arthropod phylogeny based on fossil and Recent taxa. In Edgecombe, G.E. (ed.) Arthropod fossils and phylogeny, 33 -106. Columbia University Press.
Cooper, A. and Fortey, R.A. Evolutionary explosions and the phylogenetic fuse. Trends in Ecology and Evolution. 13(4):151 - 156.
Richard Fortey LIFE: a natural history of the first 4000 million years of life on earth (reprinted from Life: An Unauthorised biography) Alfred Knopf, New York.
Fortey, R.A. and Cocks L.R.M. Lower Palaeozoic palaeogeography of Thailand: a review In, Hall, R and Holloway J.D. (eds). Biogeography and Geological Evolution of Southeast Asia, 43 - 57. Backhuys, Leiden.
Adrain, J.M., Fortey, R.A. and Westrop, S.R. Post-Cambrian trilobite diversity and Evolutionary Faunas. Science 280:1922 - 5.
Fortey, R.A. and Hughes, N.C. Brood Pouches in trilobites. Journal of Paleontology 72: 638 - 649.
Davidek, K. Landing, E, Bowring, S.A., Westrop, S.R., Rushton, A.W.A., Fortey, R.A. and Adrain J.M. New uppermost Cambrian U-Pb date from Avalonian Wales and age of the Cambrian Ordovician boundary. Geological Magazine 135: 305 - 9.
Bromham, L., Rambaut, A, and Fortey, R. et al. Testing the Cambrian explosion hypothesis by using a molecular dating technique. PNAS 95:12386 - 9.
Austin, J.J., Smith, A.B., Fortey, R.A. and Thomas, R.H. Ancient DNA from amber inclusions: a review of the evidence. Ancient biomolecules 2: 167 -176.
1999
Fortey, R.A. & Owens, R.M. The Trilobite exoskeleton, in, Savazzi, E. (ed.) Functional morphology of the invertebrate exoskeleton, 537 - 562. Wiley, New York.
Fortey, R.A. and Droser, M.L. Trilobites from the base of the type Whiterockian (Middle Ordovician) in Nevada. Journal of Paleontology, 73: 182 - 201.
McCormick, T., and Fortey, R. A. The most widely distributed trilobite species: Ordovician Carolinites genacinaca Journal of Paleontology. 73: 202 - 218.
Fortey, R.A. & Owens, R.M. Feeding habits in trilobites. Palaeontology 42 (3): 429 - 465.
Fortey, R A. Olenid trilobites as chemoautotrophic symbionts. Acta Universitatis Carolinae 43: 355 - 6.
Patterson, C. (late) & Fortey R.A. Thomas Stanley Westoll. Biographical Memoirs of the Fellows of the Royal Society, 45: 531 - 546.
Fortey, R.A. Colin Patterson. Biographical Memoirs of the Fellows of the Royal Society, 45:365 - 37.
2000
Fortey , R.A. Olenid trilobites: the oldest chemoautotrophic symbionts? Proceedings of the National Academy of Sciences, Washington, 97(12) 6574-8.
Fortey, R.A., Harper, D. A. T., Ingham, J.K., Owen, A.W., Parkes, M.A., Rushton, A.W.A. & N.H. Woodcock. A revised correlation of Ordovician Rocks in the British Isles. Geological Society, Special report No. 24, 83 pp, 34 figs.
R. A. Fortey. “Trilobite! Eye Witness to Evolution.” HarperCollins, London. 269 pp.
Edgecombe, G.E. & Fortey, R.A. Silurian trilobites from the El Carmen Formation, Bolivia. Senckenbergiana lethaia 79, 329-55.
Fortey R. A. Crystal Eyes. Natural History 109, 68-73.
Landing, E, Bowring, S.A., Davidek, K.L., Rushton, A.W.A., Fortey, R.A. & Wimbledon, W.A.P. Cambrian-Ordovician boundary age and duration of the lowest Ordovician Tremadoc Series based on U-Pb zircon dates from Avalonian Wales. Geological Magazine 137, 485-494.
Wills, M A and Fortey R A. The shape of life: how much is written in stone? Bioessays 22:1142-1152.
2001
Owens, R.M., Lane, P.D., Thomas A.T, Fortey , R.A. et al. South Wales and the Welsh Borderland. Field Guide for the Third International Symposium on Trilobites. Palaeontological Association, London.
Zhang Y.-D. & Fortey, R. A. The proximal development and thecal structure of the Ordovician graptolites Tylograptus and Sinograptus. Palaeontology 44, 553-573.
Forey, P.L. & Fortey, R.A. Fossils in the reconstruction of phylogeny. 515-9, in Briggs, D.E.G. and Crowther, P.R. (eds) Palaeobiology II. Blackwells, Oxford.
Fortey, R.A. The Cambrian explosion exploded? Science, N.Y. 293, 438-9.
Bettley, R., Fortey, R.A. and Siveter, D.J.. High resolution correlation of Anglo-Welsh Middle to Upper Ordovician sequences and its relevance to international chronostratigraphy. Journal of the Geological Society, London, 158, 937-52.
Adrain. J.M., Westrop, S.R. Landing, E. and Fortey, R.A. Systematics of the Ordovician trilobites Ischyrotoma and Dimeropygiella with species from the type Ibexian area, western USA. Journal of Paleontology 75, 947-971.
R. A. Fortey. Trilobite systematics: the last 75 years. Journal of Paleontology 75, 1141-51.
Siveter, D.J., Waloszek, D., Williams, M. and Fortey, R.A. Crustaceans and the Cambrian explosion. Science, N Y 294, 2047.
2002
McCormick, T. and R.A.Fortey. The Ordovician trilobite Carolinites, a test case for microevolution in a macrofossil lineage. Palaeontology, 45: 229 - 257.
R. A Fortey. Retrospective. Stephen Jay Gould (1941-2002). Science, N.Y. 296:1984.
Rushton, A. W. A., Cocks, L.R.M. and Fortey, R. A. Upper Cambrian trilobites and brachiopods from Severnaya Zemlya, Arctic Russia, and their implications for correlation and biogeography. Geological magazine 139: 281 - 290.
Lees, D C., Fortey, R.A and Cocks, L.R.M. Quantifying paleogeography using biogeography: a test case for the Ordovician and Silurian of Avalonia based on brachiopods and trilobites. Paleobiology 28: 343 - 363.
Fortey, R A. Fossils: the key to the Past. 3rd Edition, 232 pp. Natural History Museum, London, and Smithsonian Institution, Washington.
Cocks, L. R.M and Fortey, R.A. The palaeogeographical significance of the latest Ordovician fauna from the Pangsha Pye Formation of Burma. Special Papers in Palaeontology 67: 57 - 76.
2003.
Fortey, R. A. and Cocks, L. R. M. Palaeontological evidence bearing on global Ordovician–Silurian continental reconstructions, Earth Science Reviews, 61: 245 - 307.
Vandenbrouke, T., Fortey, R.A., Siveter, D.J. and Rickards, R.B. Chitinozoans from key sections in the Upper Ordovician Series: new GSSPs and classical British sections 151-2, in, Ordovician from the Andes. Instituto Superior de Correlacion Geologica Insugeo. Serie Correlacion Geologica 17.
Cocks, L.R. M. and Fortey, R.A. Central Asian terranes and biogeography in the Ordovician, 511 - 6, in, Ordovician from the Andes. Instituto Superior de Correlacion Geologica Insugeo. Serie Correlacion Geologica 17: 511 - 515.
Fortey, R.A. and B.D.E.Chatterton. A Devonian trilobite with an eye shade. Science 301, 1689.
Fortey, R.A. and Rushton, A.W.A. A new aglaspidid arthropod from the Lower Ordovician of Wales. Palaeontology 46:1031 -1038.
Fortey, R. A. Strangelove versus Calculus: the two faces of Science, pp. 66-69 in Words and Pictures: explaining science through art and writing. Cumbria Institute of the Arts.
Lane, P.D. Fortey, R. A. and Siveter, D.J. (eds). Trilobites and their relatives. Special Papers in Palaeontolog,y 70: 397(and Introduction, by Fortey, Siveter and Lane, p. 7-8).
Fortey, R. A. Foreword, ix-x, in Hou X-G et al. The Cambrian Fossils of Chengjiang, China. Blackwells, Oxford.
Fortey, R.A., Jackson J. and Strugnell, J. Phylogenetic fuses and evolutionary ‘explosions: conflicting evidence and critical tests. In Donoghue, P.C. J. and Smith M. P. (eds) Telling the evolutionary time, pp. 41-65. CRC Press.
Fortey, Richard, Foreward to Frontiers 03: Science and technology 2002-2003. (edited by Tim Radford). Atlantic Books, 128 pp.
2004
Richard Fortey “The Earth: an intimate history.” HarperCollins. 501 pp. [Published in the USA under the title “Earth an intimate history” by Knopf]
Forey, P.L., Fortey, R.A., Kenrick, P. and Smith, A.B. Taxonomy and fossils: a critical appraisal. Philosophical Transactions of the Royal Society, Series B 359, 639 - 653.
Adrain, J.M. Edgecombe, G.D., Fortey, R.A, et al. Trilobites, pp. 231 - 254, in Webby, B.D., Paris, F., Droser, M.L. and Percival, I. (eds) The great Ordovician biodiversification event. Columbia University Press. 484 pp.
Fortey R, A. The lifestyles of the trilobites. American Scientist 92: 446 - 453.
2005
Cocks, L.R. M., Fortey, R.A and C.P. Lee. A review of Lower and Middle Palaeozoic biostratigraphy in west peninsular Malaysia and southern Thailand in its context within the Sibumasu Terrane. Journal of Asian Earth Sciences 24: 703 - 717.
Fortey, R.A. Ordovician, in Selley, R. Cocks, L.R.M. and Plimer, I. (eds) Encyclopedia of Geology, 4:175 - 284.
Cotton, T.J. and Fortey, R.A. Comparative morphology and relationships of the Agnostida, in Koenemann, S. and Jenner, R.A. (eds) Crustacea and arthropod relationships. Crustacean Issues 16: 95 -136.
Fortey, R.A. and Cocks, L.R.M. Late Ordovician global warming – the Boda event. Geology 33, 405 - 408.
Knell, R and Fortey R.A. Trilobite spines and beetle horns: sexual selection in the Palaeozoic? Biology Letters 1(2):196 - 9.
Briggs, D.E.G. and Fortey, R.A. Wonderful strife: systematics, stem groups and the phylogenetic signal of the Cambrian radiation. In Macroevolution: diversity, disparity, contingency. Supplement to Paleobiology 31: 94 -112.
Fortey, R.A. Opipeuterella, a replacement name for the trilobite Opipeuter Fortey, 1974, preoccupied. Journal of Paleontology 79: 989.
Pollitt, J.R., Fortey, R.A. and Wills, M.A. Systematics of the trilobite families Lichidae Hawle & Corda 1847 and Lichkephalidae Tripp, 1957; the application of Bayesian inference to morphological data. Journal of Systematic Palaeontology 3: 225 - 241.
Fortey, R.A., Zhang Yuandong and Mellish, C.J.T. The relationships of biserial graptolites. Palaeontology 48: 1241 - 1272.
Fortey, R.A., and Cocks, L.R.M. Fossil evidence supports Lower Palaeozoic oceanic separations. Courier Forsch.-Inst. Senckenberg 254: 465 - 472.
2006
Cronier, C and Fortey, R.A. Morphology and ontogeny of an early Devonian phacopid with reduced sight from southern Thailand. Journal of Paleontology 89: 529 - 536.
Yuan, W.-W., Fortey, R.A. and Turvey, S.T. 2006. Ontogeny and relationships of the trilobite Pseudopetigurus Prantl & Pribyl. Palaeontology 49: 537 - 546.
Fortey R. A. .A new deep water Upper Ordovician (Caradocian) trilobite fauna from South –West Wales. Geological Journal 41: 243 - 253.
Chatterton, B.D.E., Fortey, R.A., Brett. K., Gibb, S. and McKellar, R. Trilobites from the upper Lower to Middle Devonian Timrhanrhart Formation, Jebel Gara el Zguilma, southern Morocco. Palaeontographica Canadiana 25, 177 pp.
2007
Foreword. Whatever is under the earth Gordon Herries Davies, Geological Society Publishing House.
Siveter, D.J., Fortey, R.A., Sutton, M. Briggs, D.E.G., and Siveter, D. A Silurian ‘marrellomorph’ arthropod. Proceedings of the Royal Society B, 274: 2223 - 9.
Forward into the past. Geoscientist 13: 121 - 4.
An autobiography of the frozen, 42 - 3 In Findings on ice. Pars Foundation.
Fortey R.A., and Rushton A.W.A. 2007. The mid-Cambrian trilobite Beishanella and the limits of convergent evolution. Memoirs of the Australasian Association of Palaeontologists 34: 391 - 9.
2008
Fortey, R. A. Dry Store Room Number 1. Harper Collins, London. 332 pp. (Published in USA by Knopf)
Chatterton, B. D. E. & Fortey, R.A. Linear clusters of articulated trilobites from Lower Ordovician (Arenig) strata at Bini Tin Zoulin North of Zagora, southern Morocco, in Rabano, I, Gozalo, R & Garcia –Bellido, D (eds) Advances in trilobite research, Publicaciones del Instituto Geologico y Minero de Espana. Cuarernos del Museo Geominero 9:73 -78.
Siveter, D.J., Briggs, D.E.G., Siveter, D.J., Sutton, M.D. and Fortey, R.A. Horns, eggs and legs: exceptionally preserved new arthropods from the Hereford (Silurian) Lagerstatte. Ibid, 371 - 4.
Fortey, R.A. In retrospect: Liebniz’s Protogaea. Nature 455 (4): 35.
Fortey, R.A. Cambrian evolutionary explosion. In, Benton, M. (ed.) The seventy great mysteries of the natural world. Thames and Hudson.
2009
Vandenbroucke, T. R.A., Fortey, R.A.., Verniers, J. and Ancilletta, A. A modern assessment of Ordovician chitinozoans from the Shelve and Caradoc areas, Shropshire, and their significance for correlation. Geological Magazine, 146: 216 - 236
Fortey, R.A.. Predicciones geologicas para el futuro del humanidad In, Punset,. E. (ed) El impacto de la ciencia en la sociedad 2008: Por que somos como somos, 93 - 104. Fundacion Banco Santander.
Fortey, R.A. Biogeography and evolution in the Lower Palaeozoic. 60-72, in Sepkoski, D. and Ruse, M. (eds) The Paleobiological Revolution: essays on the growth of modern paleontology. University of Chicago Press.
Fortey, R.A., A new giant asaphid trilobite from the Lower Ordovician of Morocco. Memoirs of the Association of Australasian Palaeontologists 37: 9 -16.
Fortey, R.A. & Rushton, A.W.A., The Ordovician aglaspidid arthropod Tremaglaspis reconsidered. Memoirs of the Association of Australasian Palaeontologists 37:17 - 23.
Cocks, L.R.M. and Fortey, R.A. Avalonia: a long-lived terrane in the Lower Palaeozoic? In Bassett, M.G. (ed.) Early Palaeozoic Peri-Gondwana: new insights from tectonics and biogeography. Geological Society of London, Special Publications 325, 3 - 21.
Owens, R. M. and Fortey, R.A. Silicified Upper Ordovician trilobites from Pai-khoi, Arctic Russia. Palaeontology 52, 1209 -1220.
Fortey, R.A. Fossils: the key to the past (Fourth edition) Natural History Museum London (In USA as Fossils: the history of life, Sterling 256 pp).
2010
Fortey, R.A.Trilobites of the genus Dikelokephalina from Ordovician Gondwana and Avalonia. Geological Journal (Online October 2010) Geological Journal 46: 405 - 415 (Sept 2011)
Fortey, R. A.. Charles Darwin: the scientist as hero. Contributions to Science 5(2), 183- 191. Institute of Catalan Studies.
Fortey, R.A. Archives of life: science and collections, p. 183 - 201. In Bryson, W. (ed) Seeing further (History of The Royal Society) Harper Collins.
Fortey R.A. A naturalist becomes a scientist, in, Sabbagh, K. (ed.) A Book of Kings Third Millennium.
Fortey,R.A. New Introduction. Pp. vii- xi The Earth: an intimate history The Folio Society, 416 pp.
2011
Fortey, R.A., Heward, A. P., Miller, C.G. Sedimentary facies and trilobite and conodont faunas of the Ordovician Rann Formation, Ras Al Khaimah, United Arab Emirates. Geoarabia, 16. 127 - 152.
Fortey, R.A.. Lyell and deep time. Geoscientist 21, 12 -17.
Landing E. and Fortey, R.A. Tremadocian (Lower Ordovician) sea level changes and biotas on the Avalon microcontinent. Journal of Paleontology 85, 678 - 694.
Fortey, Richard . Survivors: the animals and plants that time has left behind. Harper Press, 336 pp.
Extended reviews for literary magazines include:
Surfing the lava (Review of Volcanoes, by R. Fisher et al.) London Reviw of Books October 1997.
Shock Lobsters. (Review of The Crucible of Creation by S Conway Morris). London Review of Books. October 1998.
Most curious of seas. (Review of Noah’s Flood: The new scientific discoveries about the event that changed History by W Ryan and W Pitman) LRB July 1999
Did the self-made man fake it with Bohemian fossils? (Review of The Deprat Affair by R Osborne) LRB November 1999
No Such thing as a fish (Review of Deep Time, by Henry Gee) LRB July 2000
They might be giants (Review of The first fossil hunters, by A Mayor) LRB November 2000
Prophet of the rocks (Review of The map that changed the world by S Winchester) LRB August 2001
America’s immigrants (The eternal frontier, by T Flannery) Lead review, Times Lit. Supp. August 2001.
Down to the last flea (Review of Mammoth, by R Stone) LRB May 2002
Everything but the glue (Review of Victorian Sensation by J Secord) LRB August 2002.
Archaeology is rubbish. (Review of After the Ice: a global human history by A Mithen) LRB Dec 2003.
Decline and Fall (Review of Collapse, by Jared Diamond) The Lancet Feb 26 2005, p. 745.
Magma’s memorabilia. (Review of the Legacy of Dr Johnston-Lavis Collection, UCL). Times Lit. Suppl. Jan. 6 2006.
Natural progression. (Review of Charles Darwin, Geologist by Sandra Herbert) Times Literary Supplement, 13 Jan 2006.
Hammers for pipes. (Review of Bursting the Limits of Time by Martin Rudwick) London Review of Books, 9 February 2006
Review of Coral, by Steve Jones, for the Guardian, Saturday Review, April 2007.
Earth's Green heroes Nature 449 284-5 (Review of Oliver Morton’s Eating the Sun), 2007
In Good Measure (Review of A Natural History of Time, Pascal Richet) Times Literary Supplement, January pp. 3-5. 2008.
At nightmare’s edge (Review of The Earth on Show by Ralph O’Connor & Worlds before Adam by Martin Rudwick) Times Literary Supplement October 26-27 2008.
Diets of Worms (Review of Darwin’s Lost World by Martin Brasier) Times Literary Supplement 2009
Tasty butterflies (Review of Bugs and the Victorians by J.F.M. Clark) London Review of Books 24 September 2009 26-27
Sea Slugs, wombats, microbes (Review of The Species Seekers by Richard Conniff)
London Review of Books, 11 April 2011
Literary introductions
Introduction to the Folio Society’s reprint of The Third Policeman, by Flann O’Brien (2006)
Foreword to the Folio Society’s reprint of Life: an unauthorised biography (2007)
Mycological notes and papers
1998. Fortey, R.A. Battarraea in Oxfordshire. Mycologist 12, 159-69
1999. Fortey, R.A. Profiles of fungi. Cephalotheca suphurea. Mycologist 13, 146
2000. Fortey R.A. Old churchyards as fungus conservation areas. Field Mycology 1
2004. Fortey R.A. and Skeates, S.J. Pitptorus quercinus Field Mycology 5
2004. Fortey R A. Psilocybe aurantiaca – and a case of mistaken identity. Field Mycology 5, 77-80.
2005. Fortey, R.A. Fungi as art – Lorenzen ceramics. Field Mycology 6
2007Fortey R.A. & Ryvarden L. Ceriporiopsis herbicola nov. sp. (Polyporacea, Basidiomycota) Synopsis fungorum 23 13-15
Electronic
Fortey, R.A. 2009. Autumn Foray Report, Loxton, Somerset, October
2008. Mycologist News 2009 (2) 19-21
Fortey, R.A. 2010. Autumn Foray Report, Leiston, Suffolk, November 2009. Mycologist News 2010 (2), 5-8.