Dr Simon Loader

Dr Simon Loader

Principal Curator in Charge, Vertebrates

Department: Life Sciences
Division: LS Vertebrates Division
Phone: 0207942 6459

Summary

I am a Principal Curator of Vertebrates at the Natural History Museum. My research encompasses Africa as my geographical area of research, and vertebrates as my group of expertise. Using both Africa and vertebrates applying a range of approaches in systematics, I investigate the current patterns and processes behind the maintenance and formation of biodiversity.

Qualifications

Degrees

PhD, University of Glasgow, UK, 2000 - 2005

MSc in Advanced methods in Taxonomy and Biodiversity, Imperial College, UK, 1998 - 1999

BSc in Zoology, University of Leeds, Leeds, 1995 - 1998

Employment history

Academic

Senior Lecturer in Zoology, University of Roehampton, London Borough of Wandsworth, Life Sciences, United Kingdom, 2015 - 2017

Lecturer, University of Basel, Institute of Biogeography, Switzerland, 2007 - 2015

Non-academic

IUCN SSC Amphibian Specialist Group Regional Chair for East Africa, IUCN, 2012 - ongoing

Fellowships

UKPSF and Fellowship of the Higher Education Authority,

Honorary Fellow, London, United Kingdom,

Committees

Chair, Amphibian Specialist Group Regional Chair for East Africa, IUCN, 2008 - on going.

Memberships

Council member, 2013 - 2016.

Editorships

Associate editor, Bulletin- British Herpetological Society, 2011 - on going.

Publications

Bittencourt-silva GB, Lawson LP, Tolley KA, Portik DM, Barratt CD, Nagel P, Loader SP (2017) Impact of species delimitation and sampling on niche models and phylogeographical inference: A case study of the East African reed frog Hyperolius substriatus Ahl, 1931. Molecular Phylogenetics and Evolution, 114 : 261 - 270. doi: 10.1016/j.ympev.2017.06.022

Wilkinson MARK, Malonza PATRICKK, Campbell PATRICK, Loader SIMONP (2017) A new species of Boulengerula Tornier, 1896 (Amphibia: Gymnophiona: Herpelidae) from Kenya and the “rediscovery” of Boulengerula denhardti. Zootaxa, 4286 (4) : 525 - 525. doi: 10.11646/zootaxa.4286.4.5

Liedtke HC, Müller H, Hafner J, Penner J, Gower DJ, Mazuch T, Rödel M-O, Loader SP (2017) Terrestrial reproduction as an adaptation to steep terrain in African toads. Proceedings of the Royal Society B: Biological Sciences, 284 (1851) : 20162598 - 20162598. doi: 10.1098/rspb.2016.2598

Barratt CD, Lawson LP, Bittencourt-silva GB, Doggart N, Morgan-brown T, Nagel P, Loader SP (2017) A new, narrowly distributed, and critically endangered species of spiny-throated reed frog (Anura: Hyperoliidae) from a highly threatened coastal forest reserve in Tanzania. HERPETOLOGICAL JOURNAL, 27 (1) : 13 - 24.

Liedtke HC, Müller H, Rödel M-O, Menegon M, Gonwouo LN, Barej MF, Gvoždík V, Schmitz A, Channing A, Nagel P, Loader SP (2016) No ecological opportunity signal on a continental scale? Diversification and life-history evolution of African true toads (Anura: Bufonidae). Evolution, 70 (8) : 1717 - 1733. doi: 10.1111/evo.12985

Geiser MF, Hagmann R, Nagel P, Loader SP (2016) A first broad-scale molecular phylogeny of Prionoceridae (Coleoptera: Cleroidea) provides insight into taxonomy, biogeography and life history evolution. ARTHROPOD SYSTEMATICS & PHYLOGENY, 74 (1) : 3 - 21.

Bittencourt-silva GB, Conradie W, Siu-ting K, Tolley KA, Channing A, Cunningham M, Farooq HM, Menegon M, Loader SP (2016) The phylogenetic position and diversity of the enigmatic mongrel frog Nothophryne Poynton, 1963 (Amphibia, Anura). Molecular Phylogenetics and Evolution, 99 : 89 - 102. doi: 10.1016/j.ympev.2016.03.021

Beck J, Liedtke HC, Widler S, Altermatt F, Loader SP, Hagmann R, Lang S, Fiedler K (2016) Patterns or mechanisms? Bergmann’s and Rapoport’s rule in moths along an elevational gradient. Community Ecology, 17 (2) : 137 - 148. doi: 10.1556/168.2016.17.2.2

Conradie W, Bittencourt-silva G, Engelbrecht HM, Loader SP, Menegon M, Nanvonamuquitxo C, Scott M, Tolley KA (2016) Exploration into the hidden world of Mozambique’s sky island forests: new discoveries of reptiles and amphibians. Zoosystematics and Evolution, 92 (2) : 163 - 180. doi: 10.3897/zse.92.9948

Conradie W, Bittencourt-silva GB, Loader SP, Menegon M, Nanvonamuquitxo C, Kotzé A, Dalton DL, Engelbrecht HM, Tolley KA (2016) Batrachochytrium dendrobatidis Survey of Amphibians in the Northern Mozambique “Sky Islands” and Low-lying Areas. Herpetological Review, 47 (1) : 42 - 46.

Gereau RE, Cumberlidge N, Hemp C, Hochkirch A, Jones T, Kariuki M, Lange CN, Loader SP, Malonza PK, Menegon M (2016) Globally threatened biodiversity of the Eastern Arc mountains and coastal forests of Kenya and Tanzania. Journal of East African Natural History, 105 (1) : 115 - 201.

Hertach T, Puissant S, Gogala M, Trilar T, Hagmann R, Baur H, Kunz G, Wade EJ, Loader SP, Simon C, Nagel P (2016) Complex within a Complex: Integrative Taxonomy Reveals Hidden Diversity in Cicadetta brevipennis (Hemiptera: Cicadidae) and Unexpected Relationships with a Song Divergent Relative. PLOS ONE, 11 (11) : e0165562 - e0165562. doi: 10.1371/journal.pone.0165562

Poynton JC, Loader SP, Conradie W, Rödel M-O, Liedtke HC (2016) Designation and description of a neotype of Sclerophrys maculata (Hallowell, 1854), and reinstatement of S. pusilla (Mertens, 1937) (Amphibia: Anura: Bufonidae).. Zootaxa, 4098 (1) : 73 - 94. doi: 10.11646/zootaxa.4098.1.3

Lawson LP, Bates JM, Menegon M, Loader SP (2015) Divergence at the edges: peripatric isolation in the montane spiny throated reed frog complex. BMC Evolutionary Biology, 15 (1) : doi: 10.1186/s12862-015-0384-3

Loader SP, Lawson LP, Portik DM, Menegon M (2015) Three new species of spiny throated reed frogs (Anura: Hyperoliidae) from evergreen forests of Tanzania. BMC Research Notes, 8 (1) : doi: 10.1186/s13104-015-1050-y

Loader SP (2015) The distribution of Mertensophryne cf. anotis with a new record in Northern Mozambique. Herpetology notes, 8 : 305 - 307.

Loader SP (2015) Stormy waters: Review of The Monkey's Voyage, by Alan de Queiroz, Basic Books, pp. 368. :

Menegon M, Loader SP, Davenport TRB, Howell KM, Tilbury CR, Machaga S, Tolley KA (2015) A new species of Chameleon (Sauria: Chamaeleonidae: Kinyongia) highlights the biological affinities between the Southern Highlands and Eastern Arc Mountains of Tanzania. ACTA HERPETOLOGICA, 10 (2) : 111 - 120.

Penske S, Gvozdik V, Menegon M, Loader SP, Muller H (2015) Description of the tadpole of Leptopelis cf. grandiceps (Amphibia: Anura: Arthroleptidae) from the Uluguru Mountains, Tanzania. HERPETOLOGICAL JOURNAL, 25 (1) : 61 - 64.

Menegon M, Loader SP, Marsden SJ, Branch WR, Davenport TRB, Ursenbacher S (2014) The genus Atheris (Serpentes: Viperidae) in East Africa: Phylogeny and the role of rifting and climate in shaping the current pattern of species diversity. Molecular Phylogenetics and Evolution, 79 : 12 - 22. doi: 10.1016/j.ympev.2014.06.007

Loader SP, Sara CeccarelliF, Menegon M, Howell KM, Kassahun R, Mengistu AA, Saber SA, Gebresenbet F, De SáR, Davenport TRB, Larson JG, Müller H, Wilkinson M, Gower DJ (2014) Persistence and stability of Eastern Afromontane forests: evidence from brevicipitid frogs. Journal of Biogeography, 41 (9) : 1781 - 1792. doi: 10.1111/jbi.12331

Gvoždík V, Tillack F, Menegon M, Loader SP (2014) The status of Leptopelis barbouri Ahl, 1929 and eleven other nomina of the current tree-frog genus Leptopelis (Arthroleptidae) described from East Africa, with a redescription of Leptopelis grandiceps Ahl, 1929.. Zootaxa, 3793 : 165 - 187. doi: 10.11646/zootaxa.3793.1.8

Ceccarelli FS, Menegon M, Tolley KA, Tilbury CR, Gower DJ, Laserna MH, Kasahun R, Rodriguez-prieto A, Hagmann R, Loader SP (2014) Evolutionary relationships, species delimitation and biogeography of Eastern Afromontane horned chameleons (Chamaeleonidae: Trioceros). Molecular Phylogenetics and Evolution, 80 : 125 - 136. doi: 10.1016/j.ympev.2014.07.023

Liedtke HC, Hügli D, Dehling JM, Pupin F, Menegon M, Plumptre AJ, Kujirakwinja D, Loader SP (2014) One or two species? On the case of Hyperolius discodactylus Ahl, 1931 and H. alticola Ahl, 1931 (Anura: Hyperoliidae).. Zootaxa, 3768 : 253 - 290. doi: 10.11646/zootaxa.3768.3.2

Barej MF, Schmitz A, Günther R, Loader SP, Mahlow K, Rödel M-O (2014) The first endemic West African vertebrate family – a new anuran family highlighting the uniqueness of the Upper Guinean biodiversity hotspot. Frontiers in Zoology, 11 (1) : 8 - 8. doi: 10.1186/1742-9994-11-8

Barej MF, Rödel M-O, Loader SP, Menegon M, Gonwouo NL, Penner J, Gvoždík V, Günther R, Bell RC, Nagel P, Schmitz A (2014) Light shines through the spindrift – Phylogeny of African torrent frogs (Amphibia, Anura, Petropedetidae). Molecular Phylogenetics and Evolution, 71 : 261 - 273. doi: 10.1016/j.ympev.2013.11.001

San MauroD, Gower DJ, Müller H, Loader SP, Zardoya R, Nussbaum RA, Wilkinson M (2014) Life-history evolution and mitogenomic phylogeny of caecilian amphibians. Molecular Phylogenetics and Evolution, 73 : 177 - 189. doi: 10.1016/j.ympev.2014.01.009

Barratt CD, Tonelli E, Menegon M, Doggart N, Bowkett A, Harris WE, Howell K, Ngalason W, Loader SP (2014) Fragmented habitats and species: the challenges of Amphibian conservation in Tanzania today. Froglog,

Bwong BA, Malonza PK, Wasonga DV, Nyamache JO, Nagel P, Loader SP (2014) At a Biogeographical Crossroads: Amphibian Paradise in Shimba Hills of Kenya. Froglog, 72 - 74.

Ceccarelli FS, Menegon M, Tolley KA, Tilbury CR, Gower DJ, Laserna MH, Kasahun R, Rodriguez-prieto A, Hagmann R, Loader SP (2014) Evolutionary relationships, species delimitation and biogeography. null,

Liedtke HC, Müller H, Hafner J, Nagel P, Loader SP (2014) Interspecific patterns for egg and clutch sizes of African Bufonidae (Amphibia: Anura). Zoologischer Anzeiger - A Journal of Comparative Zoology, 253 (4) : 309 - 315. doi: 10.1016/j.jcz.2014.02.003

Mapouyat L, Hirschfeld M, Rödel M, Liedtke HC, Loader SP, Gonwouo LN, Dahmen M, Doherty-bone TM, Barej MF (2014) The tadpoles of nine Cameroonian Leptodactylodon species (Amphibia, Anura, Arthroleptidae). Zootaxa, 3765 (1) : 029 - 053.

Mapouyat L, Hirschfeld M, Roedel M-O, Liedtke HC, Loader SP, Gonwouo LN, Dahmen M, Doherty-bone TM, Barej MF (2014) The tadpoles of nine Cameroonian Leptodactylodon species (Amphibia, Anura, Arthroleptidae). ZOOTAXA, 3765 (1) : 29 - 53.

Siu-ting K, Gower DJ, Pisani D, Kassahun R, Gebresenbet F, Menegon M, Mengistu AA, Saber SA, De SáR, Wilkinson M, Loader SP (2014) Evolutionary relationships of the Critically Endangered frog Ericabatrachus baleensis Largen, 1991 with notes on incorporating previously unsampled taxa into large-scale phylogenetic analyses. BMC Evolutionary Biology, 14 (1) : 44 - 44. doi: 10.1186/1471-2148-14-44

Muller H, Liedtke HC, Menegon M, Beck J, Ballesteros-mejia L, Nagel P, Loader SP (2013) Forests as promoters of terrestrial life-history strategies in East African amphibians. Biology Letters, 9 (3) : 20121146 - 20121146. doi: 10.1098/rsbl.2012.1146

Gower DJ, Doherty-bone T, Loader SP, Wilkinson M, Kouete MT, Tapley B, Orton F, Daniel OZ, Wynne F, Flach E, Müller H, Menegon M, Stephen I, Browne RK, Fisher MC, Cunningham AA, Garner TWJ (2013) Batrachochytrium dendrobatidis Infection and Lethal Chytridiomycosis in Caecilian Amphibians (Gymnophiona). EcoHealth, 10 (2) : 173 - 183. doi: 10.1007/s10393-013-0831-9

Loader SP, Ceccarelli FS, Wilkinson M, Menegon M, Mariaux J, De SáRO, Howell KM, Gower DJ (2013) Species boundaries and biogeography of East African torrent frogs of the genus Petropedetes (Amphibia: Anura: Petropeditidae). African Journal of Herpetology, 62 (1) : 40 - 48. doi: 10.1080/21564574.2013.781549

Wilkinson M, Gower DJ, Loader SP, Müller H (2013) A visceral synapomorphy of Scolecomorphus Boulenger, 1883 (Amphibia: Gymnophiona: Scolecomorphidae). African Journal of Herpetology, 62 (1) : 21 - 27. doi: 10.1080/21564574.2013.781548

Doherty-bone TM, Gonwouo NL, Hirschfeld M, Ohst T, Weldon C, Perkins M, Kouete MT, Browne RK, Loader SP, Gower DJ, Wilkinson MW, Rödel MO, Penner J, Barej MF, Schmitz A, Plötner J, Cunningham AA (2013) Batrachochytrium dendrobatidis in amphibians of Cameroon, including first records for caecilians. Diseases of Aquatic Organisms, 102 (3) : 187 - 194. doi: 10.3354/dao02557

Gower DJ, Aberra RK, Schwaller S, Largen MJ, Collen B, Spawls S, Menegon M, Zimkus BM, De SáR, Mengistu AA, Gebresenbet F, Moore RD, Saber SA, Loader SP (2013) Long-term data for endemic frog genera reveal potential conservation crisis in the Bale Mountains, Ethiopia. Oryx, 47 (01) : 59 - 69. doi: 10.1017/S0030605311001426

Loader SP, Menegon M, Howell KM, Müller H (2013) A celebration of the works of John Charles Poynton. African Journal of Herpetology, 62 (1) : 1 - 4. doi: 10.1080/21564574.2013.793215

Mengistu AA, Nagel P, Getahun A, Saber SA, Loader SP (2013) Updated review of amphibian diversity, distribution and conservation in Ethiopia. Ethiop J Biol Sci, 12 (1) : 81 - 116.

Gower DJ, Doherty-bone TM, Aberra RK, Mengistu A, Schwaller S, Menegon M, De SaR, Saber SA, Cunningham AA, Loader SP (2012) High prevalence of the amphibian chytrid fungus (Batrachochytrium dendrobatidis) across multiple taxa and localities in the highlands of Ethiopia. HERPETOLOGICAL JOURNAL, 22 (4) : 225 - 233.

Barratt CD, Horsburgh GJ, Dawson DA, Gower DJ, Wilkinson M, Loader SP, Jehle R (2012) Characterisation of nine microsatellite loci in the caecilian amphibian Boulengerula uluguruensis (Gymnophiona), and their cross-species utility in three congeneric species. Conservation Genetics Resources, 4 (2) : 225 - 229. doi: 10.1007/s12686-011-9512-6

De SáRO, Streicher JW, Sekonyela R, Forlani MC, Loader SP, Greenbaum E, Richards S, Haddad CFB (2012) Molecular phylogeny of microhylid frogs (Anura: Microhylidae) with emphasis on relationships among New World genera.. BMC evolutionary biology, 12 : doi: 10.1186/1471-2148-12-241

Hirschfeld M, Barej MF, Loader SP, Roedel M-O (2012) Description of two Werneria tadpoles from Cameroon (Amphibia: Anura: Bufonidae). ZOOTAXA, 65 - 68.

Zimkus BM, Lawson L, Loader SP, Hanken J (2012) Terrestrialization, miniaturization and rates of diversification in African puddle frogs (Anura: Phrynobatrachidae).. PloS one, 7 (4) : doi: 10.1371/journal.pone.0035118

Menegon M, Gower DJ, Loader SP (2011) A remarkable new species of Callulina (Amphibia: Anura: Brevicipitidae) with massive, boldly coloured limb glands. ZOOTAXA, 15 - 26.

Gower DJ, Papadopoulou A, Doherty-bone TM, Pupin F, San MauroD, Loader SP, Wilkinson M (2011) The systematics of Boulengerula fischeri (Amphibia: Gymnophiona: Caeciliidae) based on morphological and molecular data. ZOOTAXA, 14 - 24.

Loader SP, Poynton JC, Lawson LP, Blackburn DC, Menegon M (2011) Herpetofauna of Montane Areas of Tanzania. 3. Amphibian Diversity in the Northwestern Eastern Arc Mountains, with the Description of a New Species of Arthroleptis (Anura: Arthroleptidae). Fieldiana Life and Earth Sciences, 4 : 90 - 102. doi: 10.3158/2158-5520-4.1.90

Loader SP, Wilkinson M, Cotton JA, Measey GJ, Menegon M, Howell KM, Mueller H, Gower DJ (2011) Molecular phylogenetics of Boulengerula (Amphibia: Gymnophiona: Caeciliidae) and implications for taxonomy, biogeography and conservation. HERPETOLOGICAL JOURNAL, 21 (1) : 5 - 16.

Menegon M, Bracebridge C, Owen N, Loader SP (2011) Herpetofauna of Montane Areas of Tanzania. 4. Amphibians and Reptiles of Mahenge Mountains, with Comments on Biogeography, Diversity, and Conservation. Fieldiana Life and Earth Sciences, 4 : 103 - 111. doi: 10.3158/2158-5520-4.1.103

Loader SP, Gower DJ, Mueller H, Menegon M (2010) Two new species of Callulina (Amphibia: Anura: Brevicipitidae) from the Nguru Mountains, Tanzania. ZOOTAXA, 26 - 42.

Loader SP, Gower DJ, Ngalason W, Menegon M (2010) Three new species of Callulina (Amphibia: Anura: Brevicipitidae) highlight local endemism and conservation plight of Africa's Eastern Arc forests. ZOOLOGICAL JOURNAL OF THE LINNEAN SOCIETY, 160 (3) : 496 - 514. doi: 10.1111/j.1096-3642.2010.00652.x

Van BocxlaerI, Loader SP, Roelants K, Biju SD, Menegon M, Bossuyt F (2010) Gradual Adaptation Toward a Range-Expansion Phenotype Initiated the Global Radiation of Toads. Science, 327 (5966) : 679 - 682. doi: 10.1126/science.1181707

Loader S (2010) Amphibians as indicators of change in Ethiopian highlands. null,

Biju SD, Van BocxlaerI, Giri VB, Loader SP, Bossuyt F (2009) Two new endemic genera and a new species of toad (Anura: Bufonidae) from the Western Ghats of India. BMC Research Notes, 2 (1) : 241 - 241. doi: 10.1186/1756-0500-2-241

Loader SP, Poynton JC, Davenport TRB, Roedel M-O (2009) Re-description of the type series of Nectophrynoides viviparus (Bufonidae), with a taxonomic reassessment. ZOOTAXA, 41 - 50.

Van BocxlaerI, Biju SD, Loader SP, Bossuyt F (2009) Toad radiation reveals into-India dispersal as a source of endemism in the Western Ghats-Sri Lanka biodiversity hotspot. BMC Evolutionary Biology, 9 (1) : 131 - 131. doi: 10.1186/1471-2148-9-131

Müller HENDRIK, Wilkinson MARK, Loader SIMONP, Wirkner CHRISTIANS, Gower DAVIDJ (2009) Morphology and function of the head in foetal and juvenile Scolecomorphus kirkii (Amphibia: Gymnophiona: Scolecomorphidae). Biological Journal of the Linnean Society, 96 (3) : 491 - 504. doi: 10.1111/j.1095-8312.2008.01152.x

Loader SP, Measey GJ, De SaRO, Malonza PK (2009) A new brevicipitid species (Brevicipitidae: Callulina) from the fragmented forests of the Taita Hills, Kenya. ZOOTAXA, 55 - 68.

Loader SP, Menegon M, Müller H, Gower DJ, Wilkinson M, Howell KM, Orton F (2009) Remarkable amphibian diversity in the South Nguru Mountains, Tanzania. Froglog, 8 - 10.

Menegon M, Burgess N, Loader SP, Doggart N, Owen N (2009) The South Nguru Mountains: a new Jewel in the Eastern Arc crown. Oryx, 43 : 174 - 176.

Poynton J, Menegon M, Loader SP (2008) A new giant species of Arthroleptis (Amphibia: Anura) from the forests of the Nguru Mountains, Tanzania. AFRICAN JOURNAL OF HERPETOLOGY, 57 (2) : 63 - 74.

Poynton JC, Loader SP (2008) Clinal Variation and Its Taxonomic Consequences in the Common Tanzanian Forest Frog, Arthroleptis affinis. Copeia, 2008 (3) : 517 - 526. doi: 10.1643/CH-07-057

Andreone F, Channing A, Drewes R, Gerlach J, Glaw F, Howell K, Largen M, Loader S, Lötters S, Minter L (2008) . AMPHIBIANS OF THE AFROTROPICAL REALM. null,

Loader SP, Pisani D, Cotton JA, Gower DJ, Day JJ, Wilkinson M (2007) Relative time scales reveal multiple origins of parallel disjunct distributions of African caecilian amphibians. Biology Letters, 3 (5) : 505 - 508. doi: 10.1098/rsbl.2007.0266

Müller H, Loader SP, Ngalason W, Howell KM, Gower DJ (2007) Reproduction in Brevicipitid Frogs (Amphibia: Anura: Brevicipitidae)—Evidence from Probreviceps M. Macrodactylus. Copeia, 2007 (3) : 726 - 733. doi: 10.1643/0045-8511(2007)2007[726:RIBFAA]2.0.CO;2

Poynton JC, Loader SP, Sherratt E, Clarke BT (2007) Amphibian Diversity in East African Biodiversity Hotspots: Altitudinal and latitudinal Patterns. Biodiversity and Conservation, 16 (4) : 1103 - 1118. doi: 10.1007/s10531-006-9074-1

Roelants K, Gower DJ, Wilkinson M, Loader SP, Biju SD, Guillaume K, Moriau L, Bossuyt F (2007) Global patterns of diversification in the history of modern amphibians. Proceedings of the National Academy of Sciences, 104 (3) : 887 - 892. doi: 10.1073/pnas.0608378104

Burgess ND, Butynski TM, Cordeiro NJ, Doggart NH, Fjeldså J, Howell KM, Kilahama FB, Loader SP, Lovett JC, Mbilinyi B, Menegon M, Moyer DC, Nashanda E, Perkin A, Rovero F, Stanley WT, Stuart SN (2007) The biological importance of the Eastern Arc Mountains of Tanzania and Kenya. Biological Conservation, 134 (2) : 209 - 231. doi: 10.1016/j.biocon.2006.08.015

Menegon M, Salvidio S, Ngalason W, Loader SP (2007) A new dwarf forest toad (Amphibia : Bufonidae : Nectophrynoides) from the Ukaguru Mountains, Tanzania. ZOOTAXA, 31 - 40.

Loader SP, Channing A, Menegon M, Davenport TRB (2006) A new species of Probreviceps (Amphibia : Anura) from the Eastern Arc Mountains, Tanzania. ZOOTAXA, 45 - 60.

Jones DT, Loader SP, Gower DJ (2006) Trophic ecology of East African caecilians (Amphibia: Gymnophiona), and their impact on forest soil invertebrates. Journal of Zoology, 269 (1) : 117 - 126. doi: 10.1111/j.1469-7998.2006.00045.x

Loader SP (2005) Systematics and biogeography of amphibians of the African Eastern Arc mountains. null,

Muller H, Measey GJ, Loader SP, Malonza PK (2005) A new species of Boulengerula tornier (Amphibia : Gymnophiona : Caeciliidae) from an isolated mountain block of the Taita Hills, Kenya. ZOOTAXA, 37 - 50.

Menegon M, Salvidio S, Loader SP (2004) Five new species of Nectophrynoides Noble 1926 (Amphibia Anura Bufonidae) from the Eastern Arc Mountains, Tanzania. Tropical Zoology, 17 (1) : 97 - 121. doi: 10.1080/03946975.2004.10531201

De SáRO, Loader SP, Channing A (2004) A New Species of Callulina (Anura: Microhylidae) from the West Usambara Mountains, Tanzania. Journal of Herpetology, 38 (2) : 219 - 224. doi: 10.1670/209-03A

Gower DJ, Rasmussen JB, Loader SP, Wilkinson M (2004) The caecilian amphibian Scolecomorphus kirkii Boulenger as prey of the burrowing asp Atractaspis aterrima Gunther: trophic relationships of fossorial vertebrates. African Journal of Ecology, 42 (2) : 83 - 87. doi: 10.1111/j.1365-2028.2004.00495.x

Loader SP, Poynton JC, Mariaux J (2004) Herpetofauna of Mahenge Mountain, Tanzania: a window on African biogeography. AFRICAN ZOOLOGY, 39 (1) : 71 - 76.

Cohen BL, Wilkinson M, Loader S, Howell K, De SáRO, Gower D, Clarke BT, Doggart N, Rödel MO (2004) Phylogenetic relationships of African Microhylid frogs inferred from DNA sequences of mitochondrial 12S and 16S ribosomal rRNA genes. Org, Divers Evol 4: 227-235. null,

Gower DJ, Loader SP, Moncrieff CB, Wilkinson M (2004) Niche separation and comparative abundance of Boulengerula boulengeri and Scolecomorphus vittatus (Amphibia: Gymnophiona) in an East Usambara forest, Tanzania. African Journal of Herpetology, 53 (2) : 183 - 190. doi: 10.1080/21564574.2004.9635510

Gower DJ, Loader SP, Wilkinson M (2004) Assessing the conservation status of soil‐dwelling vertebrates: Insights from the rediscovery of Typhlops uluguruensis (Reptilia: Serpentes: Typhlopidae). Systematics and Biodiversity, 2 (1) : 79 - 82. doi: 10.1017/S1477200004001343

Wilkinson M, Loader SP, Müller H, Gower DJ (2004) Taxonomic status and phylogenetic relationships of Boulengerula denhardtiNieden, 1912 (Amphibia, Gymnophiona, Caeciliidae). Zoosystematics and Evolution, 80 (1) : 41 - 51.

Loader SP, Wilkinson M, Gower DJ, Msuya CA (2003) A remarkable young Scolecomorphus vittatus (Amphibia: Gymnophiona: Scolecomorphidae) from the North Pare Mountains, Tanzania. Journal of Zoology, 259 (1) : 93 - 101. doi: 10.1017/S0952836902003060

Wilkinson M, Loader SP, Gower DJ, Sheps JA, Cohen BL (2003) Phylogenetic relationships of African caecilians (Amphibia: Gymnophiona): Insights from mitochondrial rRNA gene sequences. African Journal of Herpetology, 52 (2) : 83 - 92. doi: 10.1080/21564574.2003.9635483

Gower DJ, Kupfer A, Oommen OV, Himstedt W, Nussbaum RA, Loader SP, Presswell B, Muller H, Krishna SB, Boistel R, Wilkinson M (2002) A molecular phylogeny of ichthyophiid caecilians (Amphibia: Gymnophiona: Ichthyophiidae): out of India or out of South East Asia?. Proceedings of the Royal Society B: Biological Sciences, 269 (1500) : 1563 - 1569. doi: 10.1098/rspb.2002.2050

Wilkinson M, Loader S, Msuya CA, Gower D (2002) A remarkable young Scolecomorphus vittatus (Amphibia: Gymnophiona Scolecomorphidae) from the North Pare Mountains, Tanzania. null,

Krause J, Loader SP, Kirkman E, Ruxton GD (1999) Refuge use by fish as a function of body weight changes. acta ethologica, 2 (1) : 29 - 34. doi: 10.1007/PL00012229

Krause J, Loader SP, Mcdermott J, Ruxton GD (1998) Refuge use by fish as a function of body length-related metabolic expenditure and predation risks. Proceedings of the Royal Society B: Biological Sciences, 265 (1413) : 2373 - 2379. doi: 10.1098/rspb.1998.0586

Barratt CD, Bwong BA, Onstein RE, Rosauer DF, Menegon M, Doggart N, Nagel P, Kissling WD, Loader SP (2017) Environmental correlates of phylogenetic endemism in amphibians and the conservation of refugia in the Coastal Forests of Eastern Africa. Diversity and Distributions, 23 (8) : 875 - 887. doi: 10.1111/ddi.12582

Barratt CD, Lawson LP, Bittencourt-silva GB, Doggart N, Morgan-brown T, Nagel P, Loader SP (null) Herpetological Journal FULL PAPER. null,

Mengistu AA, Loader SP (null) Amphibian Conservation in the Ethiopian Highlands: The Case of Endemic Tree Frogs (Leptopelis, Arthroleptidae) and Ridged Frogs (Ptychadena, Ptychadenidae). AMPHIBIAN DIVERSITY, DISTRIBUTION AND CONSERVATION IN THE ETHIOPIAN HIGHLANDS,

Mengistu AA, Nagel P, Getahun A, Saber SA, Loader SP (null) Current Status of Amphibian Diversity, Distribution and Conservation in Ethiopia: with emphasis on Leptopelis and Ptychadena (Anura). AMPHIBIAN DIVERSITY, DISTRIBUTION AND CONSERVATION IN THE ETHIOPIAN HIGHLANDS,

Introduction

Summary

I am a Principal Curator of Vertebrates at the Natural History Museum. My research encompasses Africa as my geographical area of research, and vertebrates as my group of expertise. Using both Africa and vertebrates applying a range of approaches in systematics, I investigate the current patterns and processes behind the maintenance and formation of biodiversity.

Qualifications

Degrees

PhD, University of Glasgow, UK, 2000 - 2005

MSc in Advanced methods in Taxonomy and Biodiversity, Imperial College, UK, 1998 - 1999

BSc in Zoology, University of Leeds, Leeds, 1995 - 1998

Employment history

Academic

Senior Lecturer in Zoology, University of Roehampton, London Borough of Wandsworth, Life Sciences, United Kingdom, 2015 - 2017

Lecturer, University of Basel, Institute of Biogeography, Switzerland, 2007 - 2015

Non-academic

IUCN SSC Amphibian Specialist Group Regional Chair for East Africa, IUCN, 2012 - ongoing

Professional activities

Fellowships

UKPSF and Fellowship of the Higher Education Authority,

Honorary Fellow, London, United Kingdom,

Committees

Chair, Amphibian Specialist Group Regional Chair for East Africa, IUCN, 2008 - on going.

Memberships

Council member, 2013 - 2016.

Editorships

Associate editor, Bulletin- British Herpetological Society, 2011 - on going.

Publications

Publications

Bittencourt-silva GB, Lawson LP, Tolley KA, Portik DM, Barratt CD, Nagel P, Loader SP (2017) Impact of species delimitation and sampling on niche models and phylogeographical inference: A case study of the East African reed frog Hyperolius substriatus Ahl, 1931. Molecular Phylogenetics and Evolution, 114 : 261 - 270. doi: 10.1016/j.ympev.2017.06.022

Wilkinson MARK, Malonza PATRICKK, Campbell PATRICK, Loader SIMONP (2017) A new species of Boulengerula Tornier, 1896 (Amphibia: Gymnophiona: Herpelidae) from Kenya and the “rediscovery” of Boulengerula denhardti. Zootaxa, 4286 (4) : 525 - 525. doi: 10.11646/zootaxa.4286.4.5

Liedtke HC, Müller H, Hafner J, Penner J, Gower DJ, Mazuch T, Rödel M-O, Loader SP (2017) Terrestrial reproduction as an adaptation to steep terrain in African toads. Proceedings of the Royal Society B: Biological Sciences, 284 (1851) : 20162598 - 20162598. doi: 10.1098/rspb.2016.2598

Barratt CD, Lawson LP, Bittencourt-silva GB, Doggart N, Morgan-brown T, Nagel P, Loader SP (2017) A new, narrowly distributed, and critically endangered species of spiny-throated reed frog (Anura: Hyperoliidae) from a highly threatened coastal forest reserve in Tanzania. HERPETOLOGICAL JOURNAL, 27 (1) : 13 - 24.

Liedtke HC, Müller H, Rödel M-O, Menegon M, Gonwouo LN, Barej MF, Gvoždík V, Schmitz A, Channing A, Nagel P, Loader SP (2016) No ecological opportunity signal on a continental scale? Diversification and life-history evolution of African true toads (Anura: Bufonidae). Evolution, 70 (8) : 1717 - 1733. doi: 10.1111/evo.12985

Geiser MF, Hagmann R, Nagel P, Loader SP (2016) A first broad-scale molecular phylogeny of Prionoceridae (Coleoptera: Cleroidea) provides insight into taxonomy, biogeography and life history evolution. ARTHROPOD SYSTEMATICS & PHYLOGENY, 74 (1) : 3 - 21.

Bittencourt-silva GB, Conradie W, Siu-ting K, Tolley KA, Channing A, Cunningham M, Farooq HM, Menegon M, Loader SP (2016) The phylogenetic position and diversity of the enigmatic mongrel frog Nothophryne Poynton, 1963 (Amphibia, Anura). Molecular Phylogenetics and Evolution, 99 : 89 - 102. doi: 10.1016/j.ympev.2016.03.021

Beck J, Liedtke HC, Widler S, Altermatt F, Loader SP, Hagmann R, Lang S, Fiedler K (2016) Patterns or mechanisms? Bergmann’s and Rapoport’s rule in moths along an elevational gradient. Community Ecology, 17 (2) : 137 - 148. doi: 10.1556/168.2016.17.2.2

Conradie W, Bittencourt-silva G, Engelbrecht HM, Loader SP, Menegon M, Nanvonamuquitxo C, Scott M, Tolley KA (2016) Exploration into the hidden world of Mozambique’s sky island forests: new discoveries of reptiles and amphibians. Zoosystematics and Evolution, 92 (2) : 163 - 180. doi: 10.3897/zse.92.9948

Conradie W, Bittencourt-silva GB, Loader SP, Menegon M, Nanvonamuquitxo C, Kotzé A, Dalton DL, Engelbrecht HM, Tolley KA (2016) Batrachochytrium dendrobatidis Survey of Amphibians in the Northern Mozambique “Sky Islands” and Low-lying Areas. Herpetological Review, 47 (1) : 42 - 46.

Gereau RE, Cumberlidge N, Hemp C, Hochkirch A, Jones T, Kariuki M, Lange CN, Loader SP, Malonza PK, Menegon M (2016) Globally threatened biodiversity of the Eastern Arc mountains and coastal forests of Kenya and Tanzania. Journal of East African Natural History, 105 (1) : 115 - 201.

Hertach T, Puissant S, Gogala M, Trilar T, Hagmann R, Baur H, Kunz G, Wade EJ, Loader SP, Simon C, Nagel P (2016) Complex within a Complex: Integrative Taxonomy Reveals Hidden Diversity in Cicadetta brevipennis (Hemiptera: Cicadidae) and Unexpected Relationships with a Song Divergent Relative. PLOS ONE, 11 (11) : e0165562 - e0165562. doi: 10.1371/journal.pone.0165562

Poynton JC, Loader SP, Conradie W, Rödel M-O, Liedtke HC (2016) Designation and description of a neotype of Sclerophrys maculata (Hallowell, 1854), and reinstatement of S. pusilla (Mertens, 1937) (Amphibia: Anura: Bufonidae).. Zootaxa, 4098 (1) : 73 - 94. doi: 10.11646/zootaxa.4098.1.3

Lawson LP, Bates JM, Menegon M, Loader SP (2015) Divergence at the edges: peripatric isolation in the montane spiny throated reed frog complex. BMC Evolutionary Biology, 15 (1) : doi: 10.1186/s12862-015-0384-3

Loader SP, Lawson LP, Portik DM, Menegon M (2015) Three new species of spiny throated reed frogs (Anura: Hyperoliidae) from evergreen forests of Tanzania. BMC Research Notes, 8 (1) : doi: 10.1186/s13104-015-1050-y

Loader SP (2015) The distribution of Mertensophryne cf. anotis with a new record in Northern Mozambique. Herpetology notes, 8 : 305 - 307.

Loader SP (2015) Stormy waters: Review of The Monkey's Voyage, by Alan de Queiroz, Basic Books, pp. 368. :

Menegon M, Loader SP, Davenport TRB, Howell KM, Tilbury CR, Machaga S, Tolley KA (2015) A new species of Chameleon (Sauria: Chamaeleonidae: Kinyongia) highlights the biological affinities between the Southern Highlands and Eastern Arc Mountains of Tanzania. ACTA HERPETOLOGICA, 10 (2) : 111 - 120.

Penske S, Gvozdik V, Menegon M, Loader SP, Muller H (2015) Description of the tadpole of Leptopelis cf. grandiceps (Amphibia: Anura: Arthroleptidae) from the Uluguru Mountains, Tanzania. HERPETOLOGICAL JOURNAL, 25 (1) : 61 - 64.

Menegon M, Loader SP, Marsden SJ, Branch WR, Davenport TRB, Ursenbacher S (2014) The genus Atheris (Serpentes: Viperidae) in East Africa: Phylogeny and the role of rifting and climate in shaping the current pattern of species diversity. Molecular Phylogenetics and Evolution, 79 : 12 - 22. doi: 10.1016/j.ympev.2014.06.007

Loader SP, Sara CeccarelliF, Menegon M, Howell KM, Kassahun R, Mengistu AA, Saber SA, Gebresenbet F, De SáR, Davenport TRB, Larson JG, Müller H, Wilkinson M, Gower DJ (2014) Persistence and stability of Eastern Afromontane forests: evidence from brevicipitid frogs. Journal of Biogeography, 41 (9) : 1781 - 1792. doi: 10.1111/jbi.12331

Gvoždík V, Tillack F, Menegon M, Loader SP (2014) The status of Leptopelis barbouri Ahl, 1929 and eleven other nomina of the current tree-frog genus Leptopelis (Arthroleptidae) described from East Africa, with a redescription of Leptopelis grandiceps Ahl, 1929.. Zootaxa, 3793 : 165 - 187. doi: 10.11646/zootaxa.3793.1.8

Ceccarelli FS, Menegon M, Tolley KA, Tilbury CR, Gower DJ, Laserna MH, Kasahun R, Rodriguez-prieto A, Hagmann R, Loader SP (2014) Evolutionary relationships, species delimitation and biogeography of Eastern Afromontane horned chameleons (Chamaeleonidae: Trioceros). Molecular Phylogenetics and Evolution, 80 : 125 - 136. doi: 10.1016/j.ympev.2014.07.023

Liedtke HC, Hügli D, Dehling JM, Pupin F, Menegon M, Plumptre AJ, Kujirakwinja D, Loader SP (2014) One or two species? On the case of Hyperolius discodactylus Ahl, 1931 and H. alticola Ahl, 1931 (Anura: Hyperoliidae).. Zootaxa, 3768 : 253 - 290. doi: 10.11646/zootaxa.3768.3.2

Barej MF, Schmitz A, Günther R, Loader SP, Mahlow K, Rödel M-O (2014) The first endemic West African vertebrate family – a new anuran family highlighting the uniqueness of the Upper Guinean biodiversity hotspot. Frontiers in Zoology, 11 (1) : 8 - 8. doi: 10.1186/1742-9994-11-8

Barej MF, Rödel M-O, Loader SP, Menegon M, Gonwouo NL, Penner J, Gvoždík V, Günther R, Bell RC, Nagel P, Schmitz A (2014) Light shines through the spindrift – Phylogeny of African torrent frogs (Amphibia, Anura, Petropedetidae). Molecular Phylogenetics and Evolution, 71 : 261 - 273. doi: 10.1016/j.ympev.2013.11.001

San MauroD, Gower DJ, Müller H, Loader SP, Zardoya R, Nussbaum RA, Wilkinson M (2014) Life-history evolution and mitogenomic phylogeny of caecilian amphibians. Molecular Phylogenetics and Evolution, 73 : 177 - 189. doi: 10.1016/j.ympev.2014.01.009

Barratt CD, Tonelli E, Menegon M, Doggart N, Bowkett A, Harris WE, Howell K, Ngalason W, Loader SP (2014) Fragmented habitats and species: the challenges of Amphibian conservation in Tanzania today. Froglog,

Bwong BA, Malonza PK, Wasonga DV, Nyamache JO, Nagel P, Loader SP (2014) At a Biogeographical Crossroads: Amphibian Paradise in Shimba Hills of Kenya. Froglog, 72 - 74.

Ceccarelli FS, Menegon M, Tolley KA, Tilbury CR, Gower DJ, Laserna MH, Kasahun R, Rodriguez-prieto A, Hagmann R, Loader SP (2014) Evolutionary relationships, species delimitation and biogeography. null,

Liedtke HC, Müller H, Hafner J, Nagel P, Loader SP (2014) Interspecific patterns for egg and clutch sizes of African Bufonidae (Amphibia: Anura). Zoologischer Anzeiger - A Journal of Comparative Zoology, 253 (4) : 309 - 315. doi: 10.1016/j.jcz.2014.02.003

Mapouyat L, Hirschfeld M, Rödel M, Liedtke HC, Loader SP, Gonwouo LN, Dahmen M, Doherty-bone TM, Barej MF (2014) The tadpoles of nine Cameroonian Leptodactylodon species (Amphibia, Anura, Arthroleptidae). Zootaxa, 3765 (1) : 029 - 053.

Mapouyat L, Hirschfeld M, Roedel M-O, Liedtke HC, Loader SP, Gonwouo LN, Dahmen M, Doherty-bone TM, Barej MF (2014) The tadpoles of nine Cameroonian Leptodactylodon species (Amphibia, Anura, Arthroleptidae). ZOOTAXA, 3765 (1) : 29 - 53.

Siu-ting K, Gower DJ, Pisani D, Kassahun R, Gebresenbet F, Menegon M, Mengistu AA, Saber SA, De SáR, Wilkinson M, Loader SP (2014) Evolutionary relationships of the Critically Endangered frog Ericabatrachus baleensis Largen, 1991 with notes on incorporating previously unsampled taxa into large-scale phylogenetic analyses. BMC Evolutionary Biology, 14 (1) : 44 - 44. doi: 10.1186/1471-2148-14-44

Muller H, Liedtke HC, Menegon M, Beck J, Ballesteros-mejia L, Nagel P, Loader SP (2013) Forests as promoters of terrestrial life-history strategies in East African amphibians. Biology Letters, 9 (3) : 20121146 - 20121146. doi: 10.1098/rsbl.2012.1146

Gower DJ, Doherty-bone T, Loader SP, Wilkinson M, Kouete MT, Tapley B, Orton F, Daniel OZ, Wynne F, Flach E, Müller H, Menegon M, Stephen I, Browne RK, Fisher MC, Cunningham AA, Garner TWJ (2013) Batrachochytrium dendrobatidis Infection and Lethal Chytridiomycosis in Caecilian Amphibians (Gymnophiona). EcoHealth, 10 (2) : 173 - 183. doi: 10.1007/s10393-013-0831-9

Loader SP, Ceccarelli FS, Wilkinson M, Menegon M, Mariaux J, De SáRO, Howell KM, Gower DJ (2013) Species boundaries and biogeography of East African torrent frogs of the genus Petropedetes (Amphibia: Anura: Petropeditidae). African Journal of Herpetology, 62 (1) : 40 - 48. doi: 10.1080/21564574.2013.781549

Wilkinson M, Gower DJ, Loader SP, Müller H (2013) A visceral synapomorphy of Scolecomorphus Boulenger, 1883 (Amphibia: Gymnophiona: Scolecomorphidae). African Journal of Herpetology, 62 (1) : 21 - 27. doi: 10.1080/21564574.2013.781548

Doherty-bone TM, Gonwouo NL, Hirschfeld M, Ohst T, Weldon C, Perkins M, Kouete MT, Browne RK, Loader SP, Gower DJ, Wilkinson MW, Rödel MO, Penner J, Barej MF, Schmitz A, Plötner J, Cunningham AA (2013) Batrachochytrium dendrobatidis in amphibians of Cameroon, including first records for caecilians. Diseases of Aquatic Organisms, 102 (3) : 187 - 194. doi: 10.3354/dao02557

Gower DJ, Aberra RK, Schwaller S, Largen MJ, Collen B, Spawls S, Menegon M, Zimkus BM, De SáR, Mengistu AA, Gebresenbet F, Moore RD, Saber SA, Loader SP (2013) Long-term data for endemic frog genera reveal potential conservation crisis in the Bale Mountains, Ethiopia. Oryx, 47 (01) : 59 - 69. doi: 10.1017/S0030605311001426

Loader SP, Menegon M, Howell KM, Müller H (2013) A celebration of the works of John Charles Poynton. African Journal of Herpetology, 62 (1) : 1 - 4. doi: 10.1080/21564574.2013.793215

Mengistu AA, Nagel P, Getahun A, Saber SA, Loader SP (2013) Updated review of amphibian diversity, distribution and conservation in Ethiopia. Ethiop J Biol Sci, 12 (1) : 81 - 116.

Gower DJ, Doherty-bone TM, Aberra RK, Mengistu A, Schwaller S, Menegon M, De SaR, Saber SA, Cunningham AA, Loader SP (2012) High prevalence of the amphibian chytrid fungus (Batrachochytrium dendrobatidis) across multiple taxa and localities in the highlands of Ethiopia. HERPETOLOGICAL JOURNAL, 22 (4) : 225 - 233.

Barratt CD, Horsburgh GJ, Dawson DA, Gower DJ, Wilkinson M, Loader SP, Jehle R (2012) Characterisation of nine microsatellite loci in the caecilian amphibian Boulengerula uluguruensis (Gymnophiona), and their cross-species utility in three congeneric species. Conservation Genetics Resources, 4 (2) : 225 - 229. doi: 10.1007/s12686-011-9512-6

De SáRO, Streicher JW, Sekonyela R, Forlani MC, Loader SP, Greenbaum E, Richards S, Haddad CFB (2012) Molecular phylogeny of microhylid frogs (Anura: Microhylidae) with emphasis on relationships among New World genera.. BMC evolutionary biology, 12 : doi: 10.1186/1471-2148-12-241

Hirschfeld M, Barej MF, Loader SP, Roedel M-O (2012) Description of two Werneria tadpoles from Cameroon (Amphibia: Anura: Bufonidae). ZOOTAXA, 65 - 68.

Zimkus BM, Lawson L, Loader SP, Hanken J (2012) Terrestrialization, miniaturization and rates of diversification in African puddle frogs (Anura: Phrynobatrachidae).. PloS one, 7 (4) : doi: 10.1371/journal.pone.0035118

Menegon M, Gower DJ, Loader SP (2011) A remarkable new species of Callulina (Amphibia: Anura: Brevicipitidae) with massive, boldly coloured limb glands. ZOOTAXA, 15 - 26.

Gower DJ, Papadopoulou A, Doherty-bone TM, Pupin F, San MauroD, Loader SP, Wilkinson M (2011) The systematics of Boulengerula fischeri (Amphibia: Gymnophiona: Caeciliidae) based on morphological and molecular data. ZOOTAXA, 14 - 24.

Loader SP, Poynton JC, Lawson LP, Blackburn DC, Menegon M (2011) Herpetofauna of Montane Areas of Tanzania. 3. Amphibian Diversity in the Northwestern Eastern Arc Mountains, with the Description of a New Species of Arthroleptis (Anura: Arthroleptidae). Fieldiana Life and Earth Sciences, 4 : 90 - 102. doi: 10.3158/2158-5520-4.1.90

Loader SP, Wilkinson M, Cotton JA, Measey GJ, Menegon M, Howell KM, Mueller H, Gower DJ (2011) Molecular phylogenetics of Boulengerula (Amphibia: Gymnophiona: Caeciliidae) and implications for taxonomy, biogeography and conservation. HERPETOLOGICAL JOURNAL, 21 (1) : 5 - 16.

Menegon M, Bracebridge C, Owen N, Loader SP (2011) Herpetofauna of Montane Areas of Tanzania. 4. Amphibians and Reptiles of Mahenge Mountains, with Comments on Biogeography, Diversity, and Conservation. Fieldiana Life and Earth Sciences, 4 : 103 - 111. doi: 10.3158/2158-5520-4.1.103

Loader SP, Gower DJ, Mueller H, Menegon M (2010) Two new species of Callulina (Amphibia: Anura: Brevicipitidae) from the Nguru Mountains, Tanzania. ZOOTAXA, 26 - 42.

Loader SP, Gower DJ, Ngalason W, Menegon M (2010) Three new species of Callulina (Amphibia: Anura: Brevicipitidae) highlight local endemism and conservation plight of Africa's Eastern Arc forests. ZOOLOGICAL JOURNAL OF THE LINNEAN SOCIETY, 160 (3) : 496 - 514. doi: 10.1111/j.1096-3642.2010.00652.x

Van BocxlaerI, Loader SP, Roelants K, Biju SD, Menegon M, Bossuyt F (2010) Gradual Adaptation Toward a Range-Expansion Phenotype Initiated the Global Radiation of Toads. Science, 327 (5966) : 679 - 682. doi: 10.1126/science.1181707

Loader S (2010) Amphibians as indicators of change in Ethiopian highlands. null,

Biju SD, Van BocxlaerI, Giri VB, Loader SP, Bossuyt F (2009) Two new endemic genera and a new species of toad (Anura: Bufonidae) from the Western Ghats of India. BMC Research Notes, 2 (1) : 241 - 241. doi: 10.1186/1756-0500-2-241

Loader SP, Poynton JC, Davenport TRB, Roedel M-O (2009) Re-description of the type series of Nectophrynoides viviparus (Bufonidae), with a taxonomic reassessment. ZOOTAXA, 41 - 50.

Van BocxlaerI, Biju SD, Loader SP, Bossuyt F (2009) Toad radiation reveals into-India dispersal as a source of endemism in the Western Ghats-Sri Lanka biodiversity hotspot. BMC Evolutionary Biology, 9 (1) : 131 - 131. doi: 10.1186/1471-2148-9-131

Müller HENDRIK, Wilkinson MARK, Loader SIMONP, Wirkner CHRISTIANS, Gower DAVIDJ (2009) Morphology and function of the head in foetal and juvenile Scolecomorphus kirkii (Amphibia: Gymnophiona: Scolecomorphidae). Biological Journal of the Linnean Society, 96 (3) : 491 - 504. doi: 10.1111/j.1095-8312.2008.01152.x

Loader SP, Measey GJ, De SaRO, Malonza PK (2009) A new brevicipitid species (Brevicipitidae: Callulina) from the fragmented forests of the Taita Hills, Kenya. ZOOTAXA, 55 - 68.

Loader SP, Menegon M, Müller H, Gower DJ, Wilkinson M, Howell KM, Orton F (2009) Remarkable amphibian diversity in the South Nguru Mountains, Tanzania. Froglog, 8 - 10.

Menegon M, Burgess N, Loader SP, Doggart N, Owen N (2009) The South Nguru Mountains: a new Jewel in the Eastern Arc crown. Oryx, 43 : 174 - 176.

Poynton J, Menegon M, Loader SP (2008) A new giant species of Arthroleptis (Amphibia: Anura) from the forests of the Nguru Mountains, Tanzania. AFRICAN JOURNAL OF HERPETOLOGY, 57 (2) : 63 - 74.

Poynton JC, Loader SP (2008) Clinal Variation and Its Taxonomic Consequences in the Common Tanzanian Forest Frog, Arthroleptis affinis. Copeia, 2008 (3) : 517 - 526. doi: 10.1643/CH-07-057

Andreone F, Channing A, Drewes R, Gerlach J, Glaw F, Howell K, Largen M, Loader S, Lötters S, Minter L (2008) . AMPHIBIANS OF THE AFROTROPICAL REALM. null,

Loader SP, Pisani D, Cotton JA, Gower DJ, Day JJ, Wilkinson M (2007) Relative time scales reveal multiple origins of parallel disjunct distributions of African caecilian amphibians. Biology Letters, 3 (5) : 505 - 508. doi: 10.1098/rsbl.2007.0266

Müller H, Loader SP, Ngalason W, Howell KM, Gower DJ (2007) Reproduction in Brevicipitid Frogs (Amphibia: Anura: Brevicipitidae)—Evidence from Probreviceps M. Macrodactylus. Copeia, 2007 (3) : 726 - 733. doi: 10.1643/0045-8511(2007)2007[726:RIBFAA]2.0.CO;2

Poynton JC, Loader SP, Sherratt E, Clarke BT (2007) Amphibian Diversity in East African Biodiversity Hotspots: Altitudinal and latitudinal Patterns. Biodiversity and Conservation, 16 (4) : 1103 - 1118. doi: 10.1007/s10531-006-9074-1

Roelants K, Gower DJ, Wilkinson M, Loader SP, Biju SD, Guillaume K, Moriau L, Bossuyt F (2007) Global patterns of diversification in the history of modern amphibians. Proceedings of the National Academy of Sciences, 104 (3) : 887 - 892. doi: 10.1073/pnas.0608378104

Burgess ND, Butynski TM, Cordeiro NJ, Doggart NH, Fjeldså J, Howell KM, Kilahama FB, Loader SP, Lovett JC, Mbilinyi B, Menegon M, Moyer DC, Nashanda E, Perkin A, Rovero F, Stanley WT, Stuart SN (2007) The biological importance of the Eastern Arc Mountains of Tanzania and Kenya. Biological Conservation, 134 (2) : 209 - 231. doi: 10.1016/j.biocon.2006.08.015

Menegon M, Salvidio S, Ngalason W, Loader SP (2007) A new dwarf forest toad (Amphibia : Bufonidae : Nectophrynoides) from the Ukaguru Mountains, Tanzania. ZOOTAXA, 31 - 40.

Loader SP, Channing A, Menegon M, Davenport TRB (2006) A new species of Probreviceps (Amphibia : Anura) from the Eastern Arc Mountains, Tanzania. ZOOTAXA, 45 - 60.

Jones DT, Loader SP, Gower DJ (2006) Trophic ecology of East African caecilians (Amphibia: Gymnophiona), and their impact on forest soil invertebrates. Journal of Zoology, 269 (1) : 117 - 126. doi: 10.1111/j.1469-7998.2006.00045.x

Loader SP (2005) Systematics and biogeography of amphibians of the African Eastern Arc mountains. null,

Muller H, Measey GJ, Loader SP, Malonza PK (2005) A new species of Boulengerula tornier (Amphibia : Gymnophiona : Caeciliidae) from an isolated mountain block of the Taita Hills, Kenya. ZOOTAXA, 37 - 50.

Menegon M, Salvidio S, Loader SP (2004) Five new species of Nectophrynoides Noble 1926 (Amphibia Anura Bufonidae) from the Eastern Arc Mountains, Tanzania. Tropical Zoology, 17 (1) : 97 - 121. doi: 10.1080/03946975.2004.10531201

De SáRO, Loader SP, Channing A (2004) A New Species of Callulina (Anura: Microhylidae) from the West Usambara Mountains, Tanzania. Journal of Herpetology, 38 (2) : 219 - 224. doi: 10.1670/209-03A

Gower DJ, Rasmussen JB, Loader SP, Wilkinson M (2004) The caecilian amphibian Scolecomorphus kirkii Boulenger as prey of the burrowing asp Atractaspis aterrima Gunther: trophic relationships of fossorial vertebrates. African Journal of Ecology, 42 (2) : 83 - 87. doi: 10.1111/j.1365-2028.2004.00495.x

Loader SP, Poynton JC, Mariaux J (2004) Herpetofauna of Mahenge Mountain, Tanzania: a window on African biogeography. AFRICAN ZOOLOGY, 39 (1) : 71 - 76.

Cohen BL, Wilkinson M, Loader S, Howell K, De SáRO, Gower D, Clarke BT, Doggart N, Rödel MO (2004) Phylogenetic relationships of African Microhylid frogs inferred from DNA sequences of mitochondrial 12S and 16S ribosomal rRNA genes. Org, Divers Evol 4: 227-235. null,

Gower DJ, Loader SP, Moncrieff CB, Wilkinson M (2004) Niche separation and comparative abundance of Boulengerula boulengeri and Scolecomorphus vittatus (Amphibia: Gymnophiona) in an East Usambara forest, Tanzania. African Journal of Herpetology, 53 (2) : 183 - 190. doi: 10.1080/21564574.2004.9635510

Gower DJ, Loader SP, Wilkinson M (2004) Assessing the conservation status of soil‐dwelling vertebrates: Insights from the rediscovery of Typhlops uluguruensis (Reptilia: Serpentes: Typhlopidae). Systematics and Biodiversity, 2 (1) : 79 - 82. doi: 10.1017/S1477200004001343

Wilkinson M, Loader SP, Müller H, Gower DJ (2004) Taxonomic status and phylogenetic relationships of Boulengerula denhardtiNieden, 1912 (Amphibia, Gymnophiona, Caeciliidae). Zoosystematics and Evolution, 80 (1) : 41 - 51.

Loader SP, Wilkinson M, Gower DJ, Msuya CA (2003) A remarkable young Scolecomorphus vittatus (Amphibia: Gymnophiona: Scolecomorphidae) from the North Pare Mountains, Tanzania. Journal of Zoology, 259 (1) : 93 - 101. doi: 10.1017/S0952836902003060

Wilkinson M, Loader SP, Gower DJ, Sheps JA, Cohen BL (2003) Phylogenetic relationships of African caecilians (Amphibia: Gymnophiona): Insights from mitochondrial rRNA gene sequences. African Journal of Herpetology, 52 (2) : 83 - 92. doi: 10.1080/21564574.2003.9635483

Gower DJ, Kupfer A, Oommen OV, Himstedt W, Nussbaum RA, Loader SP, Presswell B, Muller H, Krishna SB, Boistel R, Wilkinson M (2002) A molecular phylogeny of ichthyophiid caecilians (Amphibia: Gymnophiona: Ichthyophiidae): out of India or out of South East Asia?. Proceedings of the Royal Society B: Biological Sciences, 269 (1500) : 1563 - 1569. doi: 10.1098/rspb.2002.2050

Wilkinson M, Loader S, Msuya CA, Gower D (2002) A remarkable young Scolecomorphus vittatus (Amphibia: Gymnophiona Scolecomorphidae) from the North Pare Mountains, Tanzania. null,

Krause J, Loader SP, Kirkman E, Ruxton GD (1999) Refuge use by fish as a function of body weight changes. acta ethologica, 2 (1) : 29 - 34. doi: 10.1007/PL00012229

Krause J, Loader SP, Mcdermott J, Ruxton GD (1998) Refuge use by fish as a function of body length-related metabolic expenditure and predation risks. Proceedings of the Royal Society B: Biological Sciences, 265 (1413) : 2373 - 2379. doi: 10.1098/rspb.1998.0586

Barratt CD, Bwong BA, Onstein RE, Rosauer DF, Menegon M, Doggart N, Nagel P, Kissling WD, Loader SP (2017) Environmental correlates of phylogenetic endemism in amphibians and the conservation of refugia in the Coastal Forests of Eastern Africa. Diversity and Distributions, 23 (8) : 875 - 887. doi: 10.1111/ddi.12582

Barratt CD, Lawson LP, Bittencourt-silva GB, Doggart N, Morgan-brown T, Nagel P, Loader SP (null) Herpetological Journal FULL PAPER. null,

Mengistu AA, Loader SP (null) Amphibian Conservation in the Ethiopian Highlands: The Case of Endemic Tree Frogs (Leptopelis, Arthroleptidae) and Ridged Frogs (Ptychadena, Ptychadenidae). AMPHIBIAN DIVERSITY, DISTRIBUTION AND CONSERVATION IN THE ETHIOPIAN HIGHLANDS,

Mengistu AA, Nagel P, Getahun A, Saber SA, Loader SP (null) Current Status of Amphibian Diversity, Distribution and Conservation in Ethiopia: with emphasis on Leptopelis and Ptychadena (Anura). AMPHIBIAN DIVERSITY, DISTRIBUTION AND CONSERVATION IN THE ETHIOPIAN HIGHLANDS,