Dr Paul Clark

Dr Paul Clark

Researcher

Department: Life Sciences
Division: LS Invertebrates
Contact: email
Phone: +44 (1) 207 942 5564
Staff Directory

Specialisms

Anguilla anguilla; Eriocheir sinensis, Chinese mitten crab, UK invasion and commercial exploitation; Thames submerged plastic litter, decapod taxonomy and systematics; brachyuran and Achelata larvae; decapod invasives; entrainment of the European eel

Summary

Started as an Assistance Scientific Officer in the Crustacea Section, Natural History Museum on 19 November 1974 and therefore I have just completed my fortieth year at South Kensington. During this period, duties undertaken include the curation of decapod collection, contract work, research (mainly brachyuran crabs) and rearing brachyuran larvae resulting in the publication of well over 200 research papers, books, edited works, joint reports and other publications. For the last 10 years work has focused mainly on research; taxonomy and systematics of decapods, brachyuran larvae, some eel studies, invasive Decapoda and riverine plastic litter.

Qualifications

Degrees

PhD, University Marine Biological Station, Millport, University of London (part time), United Kingdom, 1994 - 2007

MSc Modern Taxonomy (evening classes), Polytechnic of Central London, United Kingdom, 1977 - 1979

BSc General (external) Physiology & Zoology, Portsmouth Polytechnic, United Kingdom, 1969 - 1973

Employment history

Academic

Grade 4, level 3, Researcher in the Crustacea Section, The Natural History Museum, Department of Zoology and Life Sciences, United Kingdom, 2008 - ongoing

Grade 4, Decapoda Worker and Contract Support, The Natural History Museum, Department of Zoology, United Kingdom, 2001 - 2008

Grade 4, Curation Manager, Invertebrates II, The Natural History Museum, Department of Zoology, United Kingdom, 1992 - 2001

Senior Scientific Officer, Crustacea Section, British Museum (Natural History), Department of Zoology, United Kingdom, 1990 - 1992

Higher Scientific Officer, Crustacea Section, British Museum (Natural History), Department of Zoology, United Kingdom, 1985 - 1990

Scientific Officer, Crustacea Section, British Museum (Natural History), Department of Zoology, United Kingdom, 1977 - 1985

Assistant Scientific Officer, Crustacea Section, British Museum (Natural History), Department of Zoology, United Kingdom, 1974 - 1977

Laboratory Assistant, Wellcome Research Laboratories, Beckenham, Kent, Chemotherapy, England, 1968 - 1969

Non-academic

Computer Operator, Seismograph Services Ltd (UK), Keston, Kent, 1973 - 1974

Publications

Ng PK, Clark PF (2014) The type of the land crab Cardisoma hirtipes Dana, 1851 (Crustacea: Brachyura: Gecarcinidae).. Zootaxa, 3860 (6): 596 - 599. doi: 10.11646/zootaxa.3860.6.8

Huys R, Oidtmann B, Pond M, Goodman H, Clark PF (2014) Invasive crayfish and their symbionts in the Greater London area: new data and the fate of Astacus leptodactylus in the Serpentine and Long Water Lakes. Ethology Ecology & Evolution, 26 : 320 - 347. doi: 10.1080/03949370.2014.903433

Morritt D, Stefanoudis PV, Pearce D, Crimmen OA, Clark PF (2014) Plastic in the Thames: A river runs through it. Marine Pollution Bulletin, 78 : 196 - 200. doi: 10.1016/j.marpolbul.2013.10.035

Macpherson E, Amon D, Clark PF (2014) A new species of Munidopsis from a seamount of the Southwest Indian Ocean Ridge (Decapoda: Munidopsidae). Zootaxa, 3753 (3): 291 - 296. doi: 10.11646/zootaxa.3753.3.8

HUYS RONY, LOW MARTYNEY, GRAVE SAMMYDE, NG PETERKL, CLARK PAULF (2014)

On two reports associated with James Wood-Mason and Alfred William Alcock published by the Indian Museum and the Indian Marine Survey between 1890 and 1891: implications for malacostracan nomenclature

. Zootaxa, 3757 (1): 1 - 1. doi: 10.11646/zootaxa.3757.1.1

Lai JCY, Thoma BP, Clark PF, Felder DL, Ng PKL (2014) Phylogeny of eriphioid crabs (Brachyura, Eriphioidea) inferred from molecular and morphological studies. Zoologica Scripta, 43 (1): 52 - 64. doi: 10.1111/zsc.12030

Palero F, Clark PF, Guerao G (2014) Infraorder Achelata. , Martin JW, Olesen J, Høeg JT, Atlas of Crustacean Larvae., pp. 272 - 278. John Hopkins University Press. Baltimore.

Palero F, Guerao G, Hall M, Chan TY, Clark PF (2014) The ‘giant phyllosoma’ are larval stages of Parribacus antarcticus (Decapoda : Scyllaridae). Invertebrate Systematics, 28 (3): 258 - 258. doi: 10.1071/IS13037

Richard J, Clark PF (2014) Caridina simoni Bouvier, 1904 (Crustacea: Decapoda: Caridea: Atyoidea: Atyidae) and the synonymy by Johnson, 1963.. Zootaxa, 3841 (3): 301 - 338. doi: 10.11646/zootaxa.3841.3.1

Minchin D, Cook EJ, Clark PF (2013) Alien species in British brackish and marine waters. Aquatic Invasions, 8 (1): 3 - 19. doi: 10.3391/ai.2013.8.1.02

Karhan SU, Yokeş MB, Clark PF, Galil BS (2013) First Mediterranean record of Actaea savignii (H. Milne Edwards, 1834) (Crustacea: Decapoda: Brachyura: Xanthidae), an additional Erythrean alien crab. BioInvasions Records, 2 (2): 145 - 148.

Morritt D, Mills H, Hind K, Clifton-Dey D, Clark PF (2013) Monitoring downstream migrations of Eriocheir sinensis H. Milne Edwards, 1853 (Crustacea: Brachyura: Grapsoidea: Varunidae) in the River Thames using capture data from a water abstraction intake. Management of Biological Invasions, 4 (2): 139 - 147.

Ng PKL, CLARK PF (2012) Danielita Ng, Clark & Tan, 2010 (Crustacea: Decapoda: Brachyura: Ocypodoidea: Camptandriidae), preoccupied by Danielita Kiriakoff, 1970 (Lepidoptera: Glossata: Heteroneura: Noctuoidea: Notodontidae) [Letter]. Zootaxa, (3490): 48 - 48.

Clark PF, Fujita Y, Ball AD, Ng PKL (2012) The first zoeal stage morphology of Crossotonotus spinipes (De Man, 1888) and Pseudopalicus serripes (Alcock & Anderson, 1895), with implications for palicoid systematics (Crustacea: Brachyura: Palicoidea). Zootaxa, (3367): 191 - 203.

Mendoza JCE, CLARK PF, Ng PKL (2012) The identity of Pilumnoplax acanthomerus Rathbun, 1911 (Crustacea: Decapoda: Brachyura: Xanthidae), with new records from the central and western Pacific. Zootaxa, (3367): 211 - 221.

RICHARD J, de Grave S, CLARK PF (2012) A new atyid genus and species from Madagascar (Crustacea: Decapoda: Caridea). Zootaxa, 3162 (3162): 31 - 38.

Cumberlidge N, Clark PF (2012) The freshwater crabs of Ethiopia, northeastern Africa, with the description of a new Potamonautes cave species (Brachyura: Potamonautidae). Contributions to Zoology, 81 (4): 235 - 251.

Cumberlidge N, CLARK PF (2012) The freshwater crabs of Ethiopia, northeastern Africa, with the description of a new Potamonautes cave species (Brachyura: Potamonautidae). Contributions to Zoology, 81 (4): 235 - 251.

Galil BS, Ashelby CW, Clark PF (2012) New records of four leucosiid species (Crustacea: Decapoda: Brachyura: Leucosiidae) from Qatar, Persian Gulf.. Marine Biodiversity Records, 5 : 1 - 4.

Mortimer K, Rowson R, Mackie ASY, CLARK PF, Maslen C, A. S, Harrower C (2012) Steep Holm Island, Bristol Channel, UK: evidence of Larus fuscus Linnaeus, 1758 (lesser black-backed gull) feeding on the invasive signal crayfish, Pacifastacus leniusculus Dana, 1852. BioInvasions Records, 1 : 201 - 208.

Naser MD, Page TJ, Ng NK, Apel M, Yasser AG, Bishop JM, Ng PKL, CLARK PF (2012) Invasive records of Eriocheir hepuensis Dai, 1991 (Crustacea: Brachyura: Grapsoidea: Varunidae): implications and taxonomic consideration. BioInvasions Records, 1 : 71 - 86.

Stebbing P, Johnson P, Delahunty A, CLARK PF, McCollin T, Hale C, Clark S (2012) Reports of American lobsters, Homarus americanus (H. Milne Edwards, 1837) (Crustacea: Decapoda: Astacidea: Nephropoidea) in Great British waters.. BioInvasions Records, 1 : 17 - 23.

CLARK PF, Ng PKL (2011) A new species of Notonyx A. Milne-Edwards, 1873 (Crustacea: Decapoda: Brachyura: Goneplacidae) from the Philippines. Zootaxa, (2982): 27 - 32.

Froglia C, CLARK PF (2011) The forgotten Narrative of HMS Challenger and the implications for decapod nomenclature. Zootaxa, 2788 (2788): 45 - 56.

Palero F, Guerao G, CLARK PF, Abelló P (2010) Scyllarus arctus (Crustacea: Decapoda: Scyllaridae) final stage phyllosoma identified by DNA analysis, with morphological description. Journal of Marine Biological Association, UK, 91 (2): 485 - 492. doi: 10.1017/S0025315410000287

Allen CJ, CLARK PF, PATERSON GLJ, Hawkins LE, Aryuthaka C (2011) New record of Dotilla intermedia (Brachyura: Ocypodidae) from Thailand. Marine Biodiversity Records, 4 doi: 10.1017/S1755267211000091

Clark PF (2011) The commercial exploitation of the Chinese mitten crab, Eriocheir sinensis in the River Thames, London: damned if we don’t and damned if we do. , Galil BS, Clark PF, Carlton JD, Alien Marine Crustaceans: Distribution, Biology and Impacts., pp. 537 - 580. Springer.

CLARK PF (2011) The commercial exploitation of the Chinese mitten crab, Eriocheir sinensis in the River Thames, London: damned if we don’t and damned if we do. , Galil BS, CLARK PF, Carlton JD, In the Wrong Place - Alien Marine Crustaceans: Distribution, Biology and Impacts, Invading Nature., pp. 537 - 580. Springer Series in Invasion Ecology 6, Springer Science+Business Media B.V.

Lai JCY, Mendoza JCE, Guinot D, CLARK PF, Ng PKL (2011) Xanthidae MacLeay, 1838 (Decapoda: Brachyura: Xanthoidea) systematics: A multi-gene approach with support from adult and zoeal morphology. Zoologischer Anzeiger, 250 (4)doi: 10.1016/j.jcz.2011.07.002

CLARK PF, Ng PKL (2010) Description of the first zoea of Domecia glabra Alcock, 1899 (Crustacea: Brachyura, Domeciidae) and implications for the systematics of the Trapezioidea. Proceedings of the Biological Society of Washington, 123 (4): 258 - 273. doi: 10.2988/10-17.1

Low MEY, CLARK PF (2010) Kraussia Dana, 1852 (Crustacea: Decapoda: Brachyura), the senior homonym of Kraussia Davidson, 1852 (Brachiopoda: Terebratulida: Terebratellidina): confirmation and authorship of the replacement name Kraussina Suess, 1859.. Zootaxa, 2628 (2628): 66 - 68.

Palero F, Hall S, CLARK PF, Johnston DG, MACKENZIE-DODDS J, Thatje S (2010) DNA extraction from formalin-fixed tissue: New light from the Deep Sea. Biotechniques.. Scientia Marina, 74 (3): 465 - 470. doi: 10.3989/scimar.2010.74n3465

Ng PKL, CLARK PF (2010) On two new species of Notonyx A. Milne-Edwards, 1873 (Crustacea: Decapoda: Brachyura: Goneplacidae) from the Philippines and Fiji.. Zootaxa, 2509 (2509)

Palero F, Guerao G, CLARK PF, Abelló PC (2010) Final-stage phyllosoma of Palinustus A. Milne Edwards, 1880 (Crustacea: Decapoda: Achelata: Palinuridae) - The first complete description.. Zootaxa, 2403 (2403): 42 - 58.

RICHARD J, CLARK PF (2010) Caridina H. Milne Edwards, 1837 (Crustacea: Decapoda: Caridea: Atyoidea: Atyidae) - freshwater shrimps from eastern and southern Africa. Zootaxa, 2372 (2372): 305 - 337.

Allen CJ, Paterson GLJ, Hawkins LE, Hauton C, Clark PF, Aryuthaka C (2010) Zonation on sandy tropical beaches: a case study using Dotilla intermedia (Brachyura: Ocypodidae). Marine Ecology-Progress Series, 408 : 97 - 107. doi: 10.3354/meps08565

Cumberlidge N, Clark PF (2010) A redescription of Potamonautes loveni (Colosi, 1924), a medically important freshwater crab from western Kenya and eastern Uganda, east Africa (Brachyura: Potamoidea: Potamonautidae). , Castro P, Davie PJF, Ng PKL, Richer de Forges B, Studies on Brachyura: a homage to Danièle Guinot., pp. 61 - 74.

Cumberlidge N, CLARK PF (2010) Two new species of freshwater crabs from Mount Elgon, East Africa (Brachyura: Potamoidea: Potamonautidae) and a redescription of Potamonautes niloticus (A. Milne-Edwards, 1887) with notes on their natural history and onchocerciasis. Journal of Natural History, 44 : 1807 - 1829. doi: 10.1080/00222931003678792

Fujita Y, CLARK PF (2010) The larval development of Chirostylus stellaris Osawa, 2007 (Crustacea: Anomura: Galatheoidea: Chirostylidae) described from laboratory reared material.. Crustacean Research, 39 : 55 - 66.

Ng PKL, Clark PF (2010) The identity of Euryplax bevisi Stebbing, 1921 (Decapoda, Brachyura). , Fransen CHJM, de Grave S, Ng PKL, Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume., pp. 573 - 584.

Ng PKL, CLARK PF, Cuesta JA (2010) Establishment of a new subfamily for Shenius anomalus (Shen, 1935) (Crustacea: Decapoda: Brachyura: Dotillidae). Journal of Natural History, 44 : 1531 - 1553. doi: 10.1080/00222931003616362

Ng PKL, CLARK PF, Tan GS (2010) A new genus for Cleistostoma edwardsii Macleay, 1838 (Ocypodoidea: Camptandriidae) from South Africa. , Castro P, Davie PJF, Ng PKL, Richer de Forges B, Studies on Brachyura: a homage to Danièle Guinot. Crustacean Monographs. Vol. 11. pp. i-x, 1-366., pp. 251 - 260.

Richard J, Clark PF (2010) A new species of Caridina H. Milne Edwards, 1837 (Decapoda, Caridea, Atyidae) from the Betsiboka River basin, Madagascar. , Fransen CHJM, de Grave S, Ng PKL, Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume., pp. 637 - 644.

CLARK PF, Schram FR (2009) Raymond B. Manning: An appreciation. Journal of Crustacean Biology, 29 (4): 431 - 457. doi: 10.1651/09-3158.1

CLARK PF, Mortimer DN, Law RJ, Averns JM, Cohen BA, Wood D, Rose MD, Fernandes AR, RAINBOW PS (2009) Dioxin and PCB contamination in Chinese mitten crabs: human consumption as a control mechanism for an invasive species. Environmental Science and Technology, 43 (5): 1624 - 1629. doi: 10.1021/es802935a

Naruse T, CLARK PF (2009) Establishment of a new genus for Asthenognathus gallardoi Serène & Soh, 1976 within Gaeticinae Davie & N.K. Ng, 2007 (Crustacea: Decapoda: Brachyura: Varunidae). Zootaxa, 1987 (1987): 61 - 68.

Clark PF (2009) The bearing of larval morphology on brachyuran phylogeny. , Martin JW, Crandeall KA, Felder DL, Decapod Crustacean Phylogenetics., pp. 221 - 241. CRC Press.

HUYS R, CLARK P (2009) Case 3467. ZOSIMIDAE Seifried, 2003 (Crustacea, Copepoda, Harpacticoida): proposed emendation of spelling to ZOSIMEIDAE to remove homonymy with ZOSIMINAE Alcock, 1898 (Crustacea, Decapoda, XANTHIDAE). Bulletin of Zoological Nomenclature, 66 (1): 24 - 29.

Palero F, Guerao G, CLARK PF, Abello P (2009) The true identities of the slipper lobsters Nisto laevis and Nisto asper (Crustacea: Decapoda: Scyllaridae) and verified by DNA analysis. Invertebrate Systematics, 23 (1): 77 - 85. doi: 10.1071/IS08033

RICHARD J, CLARK PF (2009) African Caridina (Crustacea: Decapoda: Caridea: Atyidae): redescriptions of C. africana Kingsley, 1882, C. togoensis Hilgendorf, 1893, C. natalensis (Bouvier, 1925) and C. roubaudi (Bouvier, 1925) with descriptions of 14 new species. Zootaxa, 1995 : 1 - 75.

Rose MD, CLARK PF, RAINBOW PS, Mortimer DN, Averns JM, Smith F, Fernandes AR, Panton S (2009) Chinese mitten crabs in European rivers: contamination with dioxins, PCBs, PBBs and PBDEs and implications for human consumption. Organohalogen Compounds, 71 : 901 - 904.

CLARK PF, Guerao G (2008) A description of Calocarcinus africanus Calman, 1909 (Brachyura, Xanthoidea) first zoeal stage morphology with implications for Trapeziidae systematics. Proceedings of the Biological Society of Washington, 121 (4): 475 - 500. doi: 10.2988/06-11.1

Ng PKL, CLARK PF (2008) A new species of Notonyx A. Milne-Edwards, 1873 (Crustacea: Decapoda: Brachyura: Goneplacidae) from Indonesia. Zootaxa, 1897 (1897): 20 - 26.

Palero F, Guerao G, CLARK PF (2008) Palinustus mossambicus Barnard, 1926 (Crustacea: Decapoda: Achelata: Palinuridae); morphology of the puerulus stage. Zootaxa, 1857 (1857): 44 - 54.

CLARK PF, Ng PKL (2008) The lecithotrophic zoea of Chirostylus ortmanni Miyake & Baba, 1968 (Crustacea: Anomura: Galatheoidea: Chirostylidae) described from laboratory hatched material. Raffles Bulletin of Zoology, 56 (1): 85 - 94.

CLARK PF, Ng PKL, Fransen CHJM, Mclaughlin PA, Dworschak PC, Baba K (2008) A checklist of Crustacea Decapoda collected from Conic Island Cave and adjacent areas of Hong Kong. Journal of Natural History, 42 doi: 10.1080/00222930701850570

Ng PKL, CLARK PF (2008) A revision of Latopilumnus Türkay & Schuhmacher, 1985, and Aniptumnus Ng, 2002 (Crustacea: Decapoda: Brachyura: Pilumnidae). Journal of Natural History, 42 : 885 - 912. doi: 10.1080/00222930701869422

Cook EJ, CLARK PF (2007) Invasive Crustacea. Symposium 7 at the Sixth International Crustacean Congress (ICC6), held at the University of Glasgow, UK, 18-22 July 2005. Hydrobiologia, 590 (1): 1 - 121. doi: 10.1007/s10750-007-0751-1

CLARK PF, Galil BS, Poore GCB (2007) A new species of Calaxius Sakai & de Saint Laurent, 1989, from West Africa (Crustacea, Decapoda, Axiidae). Proceedings of the Biological Society of Washington, 120 (1): 63 - 73. doi: 10.2988/0006-324X(2007)120[63:ANSOCS]2.0.CO;2

Shakirova FM, Panov VE, CLARK PF (2007) New records of the Chinese mitten crab, Eriocheir sinensis H. Milne Edwards, 1853, from the Volga River, Russia. Aquatic Invasions, 2 : 169 - 173.

CLARK PF, Ng PKL (2006) First stage zoeas of Quadrella Dana, 1851 (Crustacea: Decapoda: Brachyura: Xathoidea: Trapeziidae) and their affinities with those of Tetralia Dana, 1851 and Trapezia Latreille, 1828. Hydrobiologia, 560 (1): 267 - 294. doi: 10.1007/s10750-004-0636-5

Clark PF (2006) Eriocheir sinensis H. Milne Edwards: 1853 or 1854 – Grapsidae or Varunidae?. Aquatic Invasions, 1 : 17 - 27.

CLARK PF, Abdul-Sahib IT, Al-Asid MS (2006) The first record of Eriocheir sinensis H. Milne Edwards, 1853 (Crustacea: Brachyura: Varunidae) from the Basrah region of southern Iraq. Aquatic Invasions, 1 (2): 51 - 54.

CLARK PF, Ng PKL (2006) A new species of Notonyx A Milne-Edwards, 1873 (Crustacea: Braychura: Goneplacidae) from the intertidal zone of Phuket, Thailand. Zoosystema, 28 (2)

Galil BS, CLARK PF, Ng PKL, Falciai L (2006) Actumnus globulus (Crustacea: Decapoda: Pilumnidae) - another Erythrean pilumnid in the Mediterranean Sea. Journal of the Marine Biological Association of the United Kingdom, Biodiversity Records, 2 : 1 - 4.

Hartnoll RG, CLARK PF (2006) A mass recruitment event in the land crab Gecarcinus ruricola (Linnaeus, 1758) (Brachyura: Grapsoidea: Gecarcinidae), and a description of the megalop. Zoological Journal of the Linnean Society, 146 (2): 146 - 164. doi: 10.1111/j.1096-3642.2006.00195.x

Ingle RW, CLARK PF (2006) First reported occurences of the marbled crab, Pachygrapsus marmoratus (Crustacea: Brachyura:Grapsoidea) in southern coastal waters of the British Isles. Journal of The Marine Biological Association, Biodiversity Records, 2 : 1 - 5.

ROBBINS RS, Sakari MH, Baluchi SN, CLARK PF (2006) The occurrence of Eriocheir sinensis H Milne Edwards, 1853 (Crustaceca: Brachyura: Varunidae) from the Caspian Sea region, Iran. Aquatic Invasions, 1 : 32 - 34.

Richard J, CLARK PF (2005) Caridina nilotica (P. Roux, 1833) (Crustacuea: Decapoda: Caridea: Atyidae) from East Africa, with descriptions from four new species. Proceedings of the Biological Society of Washington, 118 (4): 706 - 730. doi: 10.2988/0006-324X(2005)118[706:CNPRCD]2.0.CO;2

CLARK PF (2005) The evolutionary significance of heterochrony in the abbreviated zoeal development of pilumine crabs (Crustacea: Xanthoidea). Zoological Journal of the Linnean Society, 143 (3): 417 - 446. doi: 10.1111/j.1096-3642.2005.00149.x

CLARK PF, Ng PKL, Noho H, Shokita S (2005) The first-stage zoeas of Carpilus convexus (ForskDl, 1775), and Carpilius maculatus (Linnaeus, 1758) (Crustacea: Decapoda: Brachyura: Zanthoidea: Carpiliidae): an example of heterochrony. Journal of Plankton Research, 27 (2): 211 - 219. doi: 10.1093/plankt/fbh169

CLARK PF, Ng PKL (2005) The Indo-Pacific Pilumnidae XVIII: On the taxonomy of Pilumnus scabriusculus Adams and White, 1849, and P. sluiteri De Man, 1892, with a note on Cancer incanus Forskal, 1775 (Brachyura: Xanthoidea). Zootaxa, 841 (841): 1 - 14.

Afonso-Dias M, Pires A, Clark PF (2005) Occurrence of Chaceon gordonae (Ingle, 1985) and C. sanctaehelenae Manning & Holthuis, 1989 (Crustacea: Brachyura: Geryonidae) off the Island of São Tomé. Biodiversity Records, : 1 - 2.

CLARK PF (2005) Eriocheir sinensis H Milne Edwards: 1853 or 1854 - Grapsidae or Varunidae?. Aquatic Invasions, 1 : 17 - 27.

CLARK PF, Ng PKL, Ho PH (2004) Atergatis subdentatus (de Haan, 1835) Atergatopsis germaini A Milne Edwards, 1865 and Platypoda eydouxi (A Milne Edwards, 1865) (Crustacea: Decapoda: Xanthoidea: Xanthidae: Zosiminae) - first stage zoeal descriptions with implications for the subfamily. The Raffles Bulletin of Zoology, 52 (2): 563 - 592.

CLARK PF, Ng PKL (2004) Two zoeal stages and megalop of Pilumnus sluiteri De Man, 1892 (Crustacea: Brachyura: Xanthoidea: Pilumnidae) described from laboratory-reared material. Invertebrate Reproduction and Development, 45 (3): 205 - 219. doi: 10.1080/07924259.2004.9652592

Bamber RN, Clark PF (2004) A new species of Lophogaster (Crustacea, Mysidacea, Lophogastrida) from the equatorial eastern Atlantic. Zoosystema, 26 (3)

Clark PF, Ng PKL (2004) The larval development of Actumnus setifer (de Haan, 1835) (Brachyura: Xanthoidea: Pilumnidae) described from laboratory reared material. Crustacean Research, 33 : 27 - 50.

CLARK PF, Dionisio MA, Costa AC (2004) Microcassiope minor (Dana, 1852): a description of the first stage zoea (Crustacea: Brachyura: Zanthidae). Mediterranean Marine Science, 5 : 23 - 33.

CLARK PF, Ng P (2004) The larval development of Actumnus setifer (de Haan, 1835) (Brachyura: Xanthanoidea: Pilumnidae) described from laboratory reared material. Crustacean Research, 33 : 27 - 50.

Dornelas M, Clark PF, Paula J (2004) The larval development of Nanocassiope melanodactyla (A. Milne-Edwards. 1867) (Crustacea: Decopoda: Brachyura: Xanthidae). Journal of Natural History, 38

Ming-Shou J, CLARK PF, Ng PKL (2004) The first zoea, and first crab stage of the hydrothermal vent crab Xenograpsus testudinatus (Decapoda: Brachyura: Grapsoidea) and the systematic implications for the Varunidae. Journal of Crustacean Biology, 24 (1): 118 - 212. doi: 10.1651/C-2359

CLARK PF, Paula J (2003) Descriptions of ten Xanthoidean (Crustacea: Decapoda: Brachyura) first stage zoeas from Inhaca Island, Mozambique. The Raffles Bulletin of Zoology, 51 (2): 323 - 378.

Ng PKL, CLARK PF (2003) Three new genera of Indo-West Pacific Xanthidae (Crustacea: Decapoda: Brachyura: Xanthoidea). Zoosystema, 25 (1): 131 - 147.

RAINBOW PS, ROBBINS SM, CLARK PF (2003) Alien invaders: Chinese mitten crabs in the Thames and spreading. Biologist, 50 : 227 - 230.

Ko HS, Clark PF (2002) The zoeal development of Nanocassiope granulipes (Sakai, 1939) (Crustacea: Decapoda: Brachyura: Xanthidae) described from laboratory-reared material. Journal of Natural History, 36 (12): 1463 - 1488. doi: 10.1080/00222930110047685

Clark PF (2002) Alain Georges Paul Crosnier - Recipient of the Arthur G. Humes Award for Excellence in Research. JOURNAL OF CRUSTACEAN BIOLOGY, 22 (1): 210 - 212. doi: 10.1651/0278-0372(2002)022[0210:AGPCRO]2.0.CO;2

Cumberlidge N, Clark PF (2002) A new species of freshwater crab (Brachyura: Potamoidea: Potamonautidae) from Príncipe, Gulf of Guinea, Central Africa. Bulletin of the Natural History Museum, 68 (1): 13 - 18.

Ng PKL, Clark PF (2002) A description of a new species of Paramedaeus Guinot, 1967 (Decapoda: Brachyura: Xanthoidea: Xanthidae) with notes on xanthid species from New Caledonia. Crustaceana, 75 : 527 - 538.

Clark P, Ng P (2002) Description of a new species of Paramedaeus Guinot, 1967, with notes on Paramedaeus simplex (A. Milne-Edwards, 1873) and Metaxanthops acutus Serène, 1984 (Decapoda, Brachyura, Xanthoidea, Xanthidae). Crustaceana, 75 (3): 527 - 538. doi: 10.1163/156854002760095570

Umetsu K, Iwabuchi N, Yuasa I, Clark PF, Boxshall GA, Osawa M, Igarashi K (2002) Complete mitochondrial DNA sequence of tadpole shrimp (Triops cancriformis) and the analysis of museum samples. Electrophoresis, 23 : 4080 - 4084.

Dineen JF, Clark PF, Hines AH, Reed SA, Walton HP (2001) Life history, larval description, and natural history of Charybdis hellerii (Decapoda, Brachyura, Portunidae), an invasive crab in the western Atlantic. JOURNAL OF CRUSTACEAN BIOLOGY, 21 (3): 774 - 805. doi: 10.1651/0278-0372(2001)021[0774:LHLDAN]2.0.CO;2

Clark PF, Presswell B (2001) Adam White: The Crustacean years. RAFFLES BULLETIN OF ZOOLOGY, 49 (1): 149 - 166.

Clark PF, Neale M, Rainbow PS (2001) A morphometric analysis of regional variation in Carcinus Leach, 1814 (Brachyura : Portunidae : Carcininae) with particular reference to the status of the two species C-maenas (Linnaeus, 1758) and C-aestuarii Nardo, 1847. JOURNAL OF CRUSTACEAN BIOLOGY, 21 (1): 288 - 303. doi: 10.1651/0278-0372(2001)021[0288:AMAORV]2.0.CO;2

Clark PF (2001) Interpreting patterns in chaetotaxy and segmentation associated with abbreviated brachyuran zoeal development. Invertebrate Reproduction and Development, 38 (3): 171 - 181.

Ng PKL, Clark PF (2001) The eumedonid file: a case study of systematic compatibility using larval and adult characters (Crustacea: Decapoda: Brachyura). Invertebrate Reproduction and Development, 38 (3)

Ng PKL, Clark PF (2000) The Indo-Pacific Pilumnidae XII. On the familial placement of Chlorodiella bidentata (Nobili, 1901) and Tanaocheles stenochilus Kropp, 1984 using adult and larval characters with the establishment of a new subfamily, Tanaochelinae (Crustacea: Decapoda: Brachyura). Journal of Natural History, 34 (2): 207 - 245. doi: 10.1080/002229300299615

Clark PF, Ng PKL (1999) The identity of Chlorodius miliaris A. Milne Edwards, 1873, and the establishment of a new genus of Chlorodiinae (Crustacea: Decapoda: Brachyura: Xanthoidea: Xanthidae) from New Caledonia. Zoosystema, 21 (2): 353 - 365.

Clark PF, Rainbow PS, Robbins RS, Smith B, Yeomans WE, Thomas M, Dobson G (1998) The alien Chinese mitten crab, Eriocheir sinensis (Crustacea : Decapoda : Brachyura), in the Thames catchment. JOURNAL OF THE MARINE BIOLOGICAL ASSOCIATION OF THE UNITED KINGDOM, 78 (4): 1215 - 1221.

Ingle RW, Clark PF (1998) A swimming crab new to the British fauna, Liocarcinus vernalis (Crustacea : Brachyura : Portunidae). JOURNAL OF THE MARINE BIOLOGICAL ASSOCIATION OF THE UNITED KINGDOM, 78 (1): 223 - 229.

Clark PF, Galil BS (1998) The first stage zoea of Pseudoliomera speciosa (Dana, 1853) [Crustacea: Decapoda: Brachyura: Xanthidae], a description from laboratory hatched material. Zoosystema, 20 (2): 193 - 200.

Clark PF, Ng PKL (1998) The larval development of the poisonous mosaic crab, Lophozozymus pictor (Fabricius, 1798) (Crustacea: Decapoda: Brachyura: Xanthidae: Zosiminae), with comments on familial characters for first stage zoeas. Zoosystema, 20 (2)

Crosnier A, Clark PF (1998) Recherches zoologiques pour servir à l’histoire de la faune de l’Amérique centrale et du Mexique: fivth part – Études sur les xiphosures et les crustacés de la région mexicaine by A. Milne Edwards. Titles, contents and publication dates of the different issues, with remarks on the other parts. Archives of Natural History, 25 : 87 - 101.

Lincoln RJ, Boxshall GA, Clark PF. , (1998) A Dictionary of Ecology, Evolution, and Systematics. null. 361

Godley BJ, Smith SM, Clark PF, Taylor JD (1997) Molluscan and crustacean items in the diet of the loggerhead turtle, Caretta caretta (Linnaeus, 1758) [Testudines: Chelonidae] in the eastern Mediterranean. Journal of Molluscan Studies, 63 (3): 474 - 476. doi: 10.1093/mollus/63.3.474

McLaughlin PA, Clark PF (1997) A review of the Diogenes (Crustacea, Paguridea) hermit crabs collected by Bedford and Lanchester from Singapore and from the Skeat voyage to the Malay Peninsular, with a description of a new species and notes on Diogenes intermedius. Bulletin of the British Museum (Natural History). (Zoology), 63 (1): 33 - 49.

GALIL BELLAS, CLARK PAULF (1996) A revision of Cryptosoma Brullé, 1837 and Cycloes de Haan, 1837 (Crustacea: Brachyura: Calappidae). Zoological Journal of the Linnean Society, 117 (2): 175 - 204. doi: 10.1111/j.1096-3642.1996.tb02155.x

Galil BS, Clark PF (1995) A new species of Etisus (Crustacea, Brachyura, Xanthoidea, Xanthidae) from Nouvelle-Caledonie. Bulletin du Muséum national d'Histoire naturelle, 17

Clark PF, Crimmen OA, Naggs FC, Wahl A, Mansfield MC (1994) Transportation of fluid-preserved natural history specimens stored in glass containers: new solutions to an old problem. Collection Forum, 10 (1): 1 - 9.

Galil BS, Clark PF (1994) A revision of the genus Matuta Weber, 1795 (Crustacea: Brachyura: Calappidae). Zoologische Verhandelingen, 294 : 1 - 55.

Ng PKL, Clark PF (1994) The first stage zoea of Gaillardiellus orientalis (Odhner, 1925) (Crustacea: Decapoda: Brachyura: Xanthidae: Actaeinae). Bulletin of the Raffles Museum, 24 (4)

Boxshall GA, Evstigneeva TD, Clark PF (1993) A new interstitial cyclopoid copepod from a sandy beach on the western shore of Lake Baikal, Siberia. Hydrobiologia, 268 (2): 99 - 107. doi: 10.1007/BF00006880

Boxshall G, Evstigneeva T, Clark PF (1993) A new interstitial cyclopoid copepod from a sandy beach on the western shore of Lake Baikal, Siberia.. Hydrobiologia: the international journal on limnology and marine sciences, 268 : 99 - 107.

Clark PF, Galil BS (1993) A revision of the xanthid genus Pilodius Dana, 1851 (Crustacea: Brachyura: Xanthoidea). Journal of Natural History, 27 (5): 1119 - 1206. doi: 10.1080/00222939300770691

Clark PF, Galil BS (1993) A New Genus and Species of Axiid (Decapoda, Thalassinidea) From the Levantine Basin of the Mediterranean. Crustaceana, 64 (1): 48 - 55. doi: 10.1163/156854093X00054

Clark PF (1992) Ground glass stoppered jars for fluid collections. , Rose CL, De Torres AR, Storage of Natural History Collections: Ideas and Practical Solutions., pp. 221 - 223. Society for the Preservation of Natural History Collections.

Clark PF, Crosnier A (1992) Illustrations of the Zoology of the R.I.M.S. Investigator: authors, dates, issues, plates and titles. Archives of Natural History, 19 (3)

Cumberlidge N, Clark PF (1992) A new genus and species of freshwater crab from Cameroon, West Africa (Crustacea, Brachyura, Potamoidea, Potamonautidae). Bulletin of the British Museum (Natural History), (Zoology), 58 (2): 149 - 156.

Clark PF, Webber WR (1991) A redescription of Macrocheira kaempferi (Temminck, 1836) zoeas with a discussion of the classification of the Majoidea Samouelle, 1819 (Crustacea: Brachyura). Journal of Natural History, 25 (5): 1259 - 1279. doi: 10.1080/00222939100770781

Evstigneeva T, Clark PF, Boxshall GA (1991) Nocturnal emergence behaviour of the fauna of Lake Baikal, Siberia.. Progress in Underwater Science, 16 : 85 - 91.

Evstigneva T, Clark PF, Boxshall GA (1991) Nocturnal emergence behaviour of the fauna of Lake Baikal, Siberia. Progress in Underwater Science, 16 : 85 - 91.

Clark PF, Harrison K, Goodman SL (1990) Brock's Indian Archipelago Decapoda and Stomatopoda by De Man: 1887 or 1888?. Archives of Natural History, 17 (1): 81 - 82.

Galil BS, Clark PF (1990) Notes on trapeziid crabs (Crusctaea: Decapoda: Xanthoidea) from New Caledonia including descriptions of two new species. Memoires du Museum National D'Histoire Naturelle, 145 : 369 - 388.

Ingle RW, Clark PF (1989) Turkish crayfishes thrive in London canal. London Naturalist, 68 : 73 - 75.

CLARK P, GALIL B (1988) REDESCRIPTIONS OF TETRALIA-CAVIMANA HELLER, 1861 AND TRAPEZIA-CYMODOCE (HERBST, 1799) 1ST STAGE ZOEAS WITH IMPLICATIONS FOR CLASSIFICATION WITHIN THE SUPERFAMILY XANTHOIDEA (CRUSTACEA, BRACHYURA). PROCEEDINGS OF THE BIOLOGICAL SOCIETY OF WASHINGTON, 101 (4): 853 - 860.

Galil BS, Clark PF (1988) On a collection of Acropora-inhabiting trapeziids (Crustacea, Brachyura, Xanthoidea) from East Africa. Tropical Zoology, 1 : 137 - 151.

Clark PF, Clark PF (1987) A new species of Chasmagnathus from the Bodgaya Island Group (Semporna), Sabah, Borneo (Crustacea: Brachyura: Grapsidae). Malayan Nature Journal, 41 : 45 - 53.

Clark PF, Clark PF. , (1986) North-East Atlantic Crabs; an atlas of distribution. Marine Conservation Society.

Clark PF (1986) The larval stages of Maja squinado (Herbst, 1788) (Crustacea: Brachyura: Majidae) reared in the laboratory. Journal of Natural History, 20 (4): 825 - 836. doi: 10.1080/00222938600770591

Clark PF (1984) Recent records of alien crabs in Britain. London Naturalist, 109 : 111 - 112.

CLARK PAULF (1984) A comparative study of zoeal morphology in the genus Liocarcinus (Crustacea: Brachyura: Portunidae). Zoological Journal of the Linnean Society, 82 (3): 273 - 290. doi: 10.1111/j.1096-3642.1984.tb00644.x

Wheeler AC, Clark PF (1984) New records for the occurrence of Crenilabrus bailloni Valenciennes, 1839 (Osteichthyes-Perciformis-Labridae) in the waters of Northern Europe.. Journal of the Marine Biological Association of the United Kingdom. Plymouth, 64 (1): 1 - 6.

Clark PF (1983) The larval and first crab stages of three Inachus species (Crustacea: Decapoda: Majidae); a morphological and statistical analysis.. Bulletin of the British Museum (Natural History). (Zoology)., 44 (2): 179 - 190.

Ingle RW, Clark PF (1983) The larval development of the Angular Crab, Goneplax rhomboides (Linnaeus) (Decapoda: Brachyura). Bulletin of the British Museum (Natural History). (Zoology)., 44 (2): 163 - 177.

Ingle RW, Clark PF (1980) The larval and post-larval development of Gibbs' spider crab, Pisa armata (Latreille) (Family Majidae: sub-family Pisinae), reared in the laboratory.. Journal of Natural History., 14 (5): 723 - 735.

Ingle RW, Clark PF (1977) A laboratory module for rearing crab larvae. Crustaceana, 32 (2)

Clark PF, Crosnier A (-1) The Zoology of the Voyage au pole sud et dans l'Océanie sur les corvettes l'Astrolabe et la Zélée exécuté par ordre du roi pendant les années 1837-1838-1839-1840 sous le commandement de M. Dumont-D'Urville, 1842-1854: titles, volumes, plates, text, contents and proposed dates and anecdotal history of the publication. Archives of Natural History, 27 : 407 - 437.

Highlighted publications

Morritt D, Stefanoudis PV, Pearce D, Crimmen OA, Clark PF (2014) Plastic in the Thames: A river runs through it. Marine Pollution Bulletin, 78 : 196 - 200. doi: 10.1016/j.marpolbul.2013.10.035

Morritt D, Mills H, Hind K, Clifton-Dey D, Clark PF (2013) Monitoring downstream migrations of Eriocheir sinensis H. Milne Edwards, 1853 (Crustacea: Brachyura: Grapsoidea: Varunidae) in the River Thames using capture data from a water abstraction intake. Management of Biological Invasions, 4 (2): 139 - 147.

Naser MD, Page TJ, Ng NK, Apel M, Yasser AG, et al (2012) Invasive records of Eriocheir hepuensis Dai, 1991 (Crustacea: Brachyura: Grapsoidea: Varunidae): implications and taxonomic consideration. BioInvasions Records, 1 : 71 - 86.

Clark PF (2011) The commercial exploitation of the Chinese mitten crab, Eriocheir sinensis in the River Thames, London: damned if we don’t and damned if we do. , Galil BS, Clark PF, Carlton JD, Alien Marine Crustaceans: Distribution, Biology and Impacts., pp. 537 - 580. Springer.

CLARK PF (2011) The commercial exploitation of the Chinese mitten crab, Eriocheir sinensis in the River Thames, London: damned if we don’t and damned if we do. , Galil BS, CLARK PF, Carlton JD, In the Wrong Place - Alien Marine Crustaceans: Distribution, Biology and Impacts, Invading Nature., pp. 537 - 580. Springer Series in Invasion Ecology 6, Springer Science+Business Media B.V.

Lai JCY, Mendoza JCE, Guinot D, CLARK PF, Ng PKL (2011) Xanthidae MacLeay, 1838 (Decapoda: Brachyura: Xanthoidea) systematics: A multi-gene approach with support from adult and zoeal morphology. Zoologischer Anzeiger, 250 (4)doi: 10.1016/j.jcz.2011.07.002