Dr Harald Schneider

Dr Harald Schneider

Researcher, Pteridophytes

Department: Life Sciences
Division: LS Plants Division
Contact: email
Phone: +44 7942 6058
Research Gate
google scholar citations list

Summary

My research is focused on all aspects of the diversification of land plants with focus on seed-free land plants such as ferns and liverworts. I am especially interested in macroevolutionary patterns and their link to underlying microevolutinary processes. In my current research I am focusing in particular on the following key questions: - Diversification of land plants through time investigated by integrating new fossil evidence and DNA based divergence time estimates. - Assembly of species diversity and morphological disparity with focus on the “coupling” of the two parameters in the tree of life and its consequences for taxonomic practice - Evolution of polyploidy in ferns - Early divergence of land plant and the consequences of terrestriazation of plants - Cretaceous Terrestrial Revolution and its consequences for plant life today - Accumulation of plant diversity in SE Asia and Eurasia since the late Eocene - Evolution of insect-plant interactions with focus on the widely ignored evidence for fern-insect interactions My research integrates (1) state-of-the-art phylogenetics, (2) exhaustive expertise in plant anatomy and morphology based on the traditional expertise of the 19th and 20th century, (2) cytology, (3) biogeography and phylogeography, (4) taxonomy, (5) fossil evidence, (6) population genetics, and other related research approaches.

Qualifications

Degrees

Habilitation / venia legendi, Georg August University Goettingen, Germany, 2002 - 2006

PhD, University of Zuerich, Switzerland, 1992 - 1996

Diploma Biology, University of Kaiserslautern, Germany, 1986 - 1992

Employment history

Academic

Research Leader, Natural Hostory Museum, Life Sciences, United Kingdom, 2006 - ongoing

Lecturer, Albrecht-von-Haller Institute, Georg-August University, Goettingen, Systematic Botany, Germany, 2002 - 2006

Postdoctoral Research Assistent, Duke University, Biology, United States, 2001 - 2001

Postdoctoral Research Assisent, University of California at Berkeley, Integrative Biology, United States, 2000 - 2000

Postdoctoral Research Assisent, Field Museum of Natural History, Chicago, Botany, United States, 1999 - 1999

Marie-Curie Postdoctoral Fellow, Rijksuniversiteit Leiden, Rijksherbarium, Netherlands, 1996 - 1998

Publications

Wang A-H, Sun Y, Schneider H, Zhai J-W, Liu D-M, Zhou J-S, Xing F-W, Chen H-F, Wang F-G (2015) Identification of the relationship between Chinese Adiantum reniforme var. sinense and Canary Adiantum reniforme. BMC Plant Biology, 15 (1)doi: 10.1186/s12870-014-0361-9

Heinrichs J, Feldberg K, Bechteler J, Scheben A, Czumaj A, Pócs T, Schneider H, Schäfer-Verwimp A (2015) Integrative taxonomy of Lepidolejeunea (Jungermanniopsida: Porellales): Ocelli allow the recognition of two neglected species. Taxon, 64 (2): 216 - 228. doi: 10.12705/642.5

Schneider H, Liu H, Clark J, Hidalgo O, Pellicer J, Zhang S, Kelly LJ, Fay MF, Leitch IJ (2015) Are the genomes of royal ferns really frozen in time? Evidence for coinciding genome stability and limited evolvability in the royal ferns. New Phytologist, 207 (1): 10 - 13. doi: 10.1111/nph.13330

Schneider H, Schmidt AR, Nascimbene PC, Heinrichs J (2015) A new Dominican amber fossil of the derived fern genus Pleopeltis confirms generic stasis in the epiphytic fern diversity of the West Indies. Organisms Diversity & Evolution, 15 (2): 277 - 283. doi: 10.1007/s13127-015-0200-3

Laenen B, Shaw B, Schneider H, Goffinet B, Paradis E, Désamoré A, Heinrichs J, Villarreal JC, Gradstein SR, McDaniel SF, Long DG, Forrest LL, Hollingsworth ML, Crandall-Stotler B, Davis EC, Engel J, Von Konrat M, Cooper ED, Patiño J, Cox CJ, Vanderpoorten A, Shaw AJ (2014) Extant diversity of bryophytes emerged from successive post-Mesozoic diversification bursts. Nature Communications, 5 : 5134 - 5134. doi: 10.1038/ncomms6134

Hennequin S, Kessler M, Lindsay S, Schneider H (2014) Evolutionary patterns in the assembly of fern diversity on the oceanic Mascarene Islands. Journal of Biogeography, 41 (9): 1651 - 1663. doi: 10.1111/jbi.12339

Loriga J, Schmidt AR, Moran RC, Feldberg K, Schneider H, Heinrichs J (2014) The first fossil of a bolbitidoid fern belongs to the early-divergent lineages of Elaphoglossum (Dryopteridaceae). American Journal of Botany, 101 (9): 1466 - 1475. doi: 10.3732/ajb.1400262

Vikulin SV, Bystriakova N, Schneider H, Jolley D (2014) Plant macrofossils from Boltysh crater provide a window into early Cenozoic vegetation. Geological Society of America Special Papers, 505 : 505 - 507.

Feldberg K, Schneider H, Stadler T, Schäfer-Verwimp A, Schmidt AR, Heinrichs J (2014) Epiphytic leafy liverworts diversified in angiosperm-dominated forests. Scientific Reports, 4 doi: 10.1038/srep05974

Zhou S, Dong W, Chen X, Zhang X, Wen J, Schneider H (2014) How many species of bracken (Pteridium) are there? Assessing the Chinese brackens using molecular evidence. Taxon, 63 (3): 509 - 521. doi: 10.12705/633.9

Yu Y, Heinrichs J, Schäfer-Verwimp A, Zhu R-L, Schneider H (2014) Inferring the accumulation of morphological disparity in epiphyllous liverworts. Organisms Diversity & Evolution, 14 (2): 151 - 162. doi: 10.1007/s13127-014-0166-6

Schaefer-Verwimp A, Feldberg K, Dong S, Van Melick H, Peralta DF, Schmidt AR, Schneider H, Heinrichs J (2014) Towards a monophyletic classification of Lejeuneaceae III: the systematic position of Leiolejeunea. PHYTOTAXA, 170 (3): 187 - 198.

Vikulin SV, Bystriakova N, Schneider H, Jolley D (2014) Plant macrofossils from Boltysh crater provide a window into early Cenozoic vegetation. Geological Society of America Special Papers, 505

Yan-Fen CHANG, Jie LI, Shu-Gang LIU, Schneider H (2014) Systematic Position and Polyploid Origin of the Fern Asplenium kiangsuense (Aspleniaceae). Acta Botanica Yunnanica, 36 (1): 7 - 12.

Heinrichs J, Dong S, Schaefer-Verwimp A, Peralta DF, Feldberg K, Schmidt AR, Schneider H (2014) Towards a monophyletic classification of Lejeuneaceae II: subtribes Pycnolejeuneinae and Xylolejeuneinae subtr. nov., transfer of Otolejeunea to Lepidolejeuninae, and generic refinements. PHYTOTAXA, 163 (2): 61 - 76.

Heinrichs J, Czumaj A, Dong S, Scheben A, Schaefer-Verwimp A, Peralta DF, Feldberg K, Schmidt AR, Schneider H (2014) The Bromeliaceae tank dweller Bromeliophila (Lejeuneaceae, Porellales) is a member of the Cyclolejeunea-Prionolejeunea clade. Plant Systematics and Evolution, 300 (1): 63 - 73. doi: 10.1007/s00606-013-0860-4

LIU H-M, HE L-J, SCHNEIDER H (2014) Towards the natural classification of tectarioid ferns: Confirming the phylogenetic relationships of Pleocnemia and Pteridrys (eupolypods I). Journal of Systematics and Evolution, 52 (2): 161 - 174. doi: 10.1111/jse.12073

Rumsey FJ, Robba L, Schneider H, Carine MA (2014) Taxonomic uncertainty and a continental conundrum: Polypodium macaronesicum reassessed. Botanical Journal of the Linnean Society, 174 (3): 449 - 460. doi: 10.1111/boj.12140

Bystriakova N, Ansell SW, Russell SJ, Grundmann M, Vogel JC, Schneider H (2014) Present, past and future of the European rock fern Asplenium fontanum: combining distribution modelling and population genetics to study the effect of climate change on geographic range and genetic diversity. Annals of Botany, 113 (3): 453 - 465. doi: 10.1093/aob/mct274

Heinrichs J, Schaefer-Verwimp A, Czumay A, Dong S, Scheben A, Feldberg K, Schneider H (2014) Towards a monophyletic classification of Lejeuneaceae I: subtribe Leptolejeuneinae subtr. nov. Phytotaxa, 156 (3): 165 - 174.

Heinrichs J, Dong S, Schaefer-Verwimp A, Pocs T, Feldberg K, Czumaj A, Schmidt AR, Reitner J, Renner MAM, Hentschel J, Stech M, Schneider H (2013) Molecular Phylogeny of the Leafy Liverwort Lejeunea (Porellales): Evidence for a Neotropical Origin, Uneven Distribution of Sexual Systems and Insufficient Taxonomy. Plos One, 8 (12)doi: 10.1371/journal.pone.0082547

Yu Y, Heinrichs J, Zhu R, Schneider H (2013) Empirical Evidence Supporting Frequent Cryptic Speciation in Epiphyllous Liverworts: A Case Study of the Cololejeunea lanciloba Complex. Plos One, 8 (12)doi: 10.1371/journal.pone.0084124

Chang Y, Li J, Lu S, Schneider H (2013) Species diversity and reticulate evolution in the Asplenium normale complex (Aspleniaceae) in China and adjacent areas. Taxon, 62 (4): 673 - 687. doi: 10.12705/624.6

Liu H-M, Jiang R-H, Guo J, Hovenkamp P, Perrie LR, Shepherd L, Hennequin S, Schneider H (2013) Towards a phylogenetic classification of the climbing fern genus Arthropteris. Taxon, 62 (4): 688 - 700. doi: 10.12705/624.26

Wei R, Schneider H, Zhang X (2013) Toward a new circumscription of the twinsorus-fern genus Diplazium (Athyriaceae): A molecular phylogeny with morphological implications and infrageneric taxonomy. Taxon, 62 (3): 441 - 457. doi: 10.12705/623.8

Feldberg K, Heinrichs J, Schmidt AR, Vana J, SCHNEIDER H (2013) Exploring the impact of fossil constraints on the divergence time estimates of derived liverworts. Plant Systematics and Evolution, 299 (3): 585 - 601. doi: 10.1007/s00606-012-0745-y

SCHNEIDER H, Navarro-Gomez A, RUSSELL S, Pedersen N, ANSELL SW, VOGEL JC (2012) Exploring the utility of three nuclear regions to reconstruct the reticulate evolution in the fern genus Asplenium. Journal of Systematics and Evolution, 51 (2): 142 - 153. doi: 10.1111/j.1759-6831.2012.00226.x

SCHNEIDER H, He L, Hennequin S, Zhang X (2013) Towards a natural classification of Pteridaceae: inferring the relationships of enigmatic pteridoid fern species occurring in the Sino-Himalaya and Afro-Madagascar. Phytotaxa, 77 (4): 49 - 60. doi: 10.11646/phytotaxa.77.4.1

Dong SS, Schaefer-Verwimp A, Pocs T, Feldberg K, Czumaj A, Schmidt AR, SCHNEIDER H, Heirnichs J (2013) Size doesn’t matter - recircumscription of Microlejeunea (Lejeuneaceae, Porellales). Phytotaxa, 85 (2): 41 - 55. doi: 10.11646/phytotaxa.85.2.2

Dyer RJ, Pellicer J, Savolainen V, Leitch IJ, Schneider H (2013) Genome size expansion and the relationship between nuclear DNA content and spore size in the Asplenium monanthes fern complex (Aspleniaceae). BMC Plant Biology, 13 (1): 219 - 219. doi: 10.1186/1471-2229-13-219

Liu H, Schneider H (2013) Evidence supporting Davallia canariensis as a Late Miocene relict endemic to Macaronesia and Atlantic Europe. Australian Systematic Botany, 26 (5): 378 - 385. doi: 10.1071/sb13029

SCHNEIDER H, He L, MARQUARDT J, WANG L, Heinrichs J, Hennequin S, Zhang X (2013) Exploring the origin of the latitudinal diversity gradient: Contrasting the sister fern genera Phegopteris and Pseudophegopteris. Journal of Systematics and Evolution, 51 (1): 61 - 70. doi: 10.1111/j.1759-6831.2012.00234.x

Yu Y, Pocs T, Schaefer-Verwimp A, Heinrichs J, Zhu R, Schneider H (2013) Evidence for Rampant Homoplasy in the Phylogeny of the Epiphyllous Liverwort Genus Cololejeunea (Lejeuneaceae). Systematic Botany, 38 (3): 553 - 563. doi: 10.1600/036364413x670304

DYER RJ, Savolainen V, SCHNEIDER H (2012) Apomixis and reticulate evolution in the Asplenium monanthes fern complex. Annals of Botany, 110 (8): 1515 - 1529. doi: 10.1093/aob/mcs202

Wang L, SCHNEIDER H, Zhang X, Xiang Q (2012) The rise of the Himalaya enforced the diversification of SE Asian ferns by altering the monsoon regimes. Bmc Plant Biology, 12 (1): 210 - 210. doi: 10.1186/1471-2229-12-210

Dong SS, Schafer-Verwimp A, Meinecke P, Feldberg K, Bombosch A, Pocs T, Schmidt AR, Reitner J, SCHNEIDER H, Heinrichs J (2012) Tramps, narrow endemics and morphologically cryptic species in the epiphyllous liverwort Diplasiolejeunea. Molecular Phylogenetics and Evolution, 65 (2): 582 - 594. doi: 10.1016/j.ympev.2012.07.009

Wang L, SCHNEIDER H, Wu Z, He L, Zhang X, Xiang Q (2012) Indehiscent sporangia enable the accumulation of local fern diversity at the Qinghai-Tibetan Plateau. Bmc Evolutionary Biology, 12 (1): 158 - 158. doi: 10.1186/1471-2148-12-158

Christenhusz MJM, SCHNEIDER H (2012) (2054) Proposal to conserve the name Drynaria against Aglaomorpha (Polypodiaceae). Taxon, 61 (2): 465 - 466.

De Groot GA, During HJ, ANSELL SW, SCHNEIDER H, Bremer P, Wubs ERJ, Maas JW, Korpelainen H, Erkens RHJ (2012) Diverse spore rains and limited local exchange shape fern genetic diversity in a recently created habitat colonized by long-distance dispersal. Annals of Botany, 109 (5): 965 - 978. doi: 10.1093/aob/mcs013

Kenrick P, Wellman CH, Schneider H, Edgecombe GD (2012) A timeline for terrestrialization: consequences for the carbon cycle in the Palaeozoic. Philosophical Transactions of the Royal Society B: Biological Sciences, 367 (1588): 519 - 536. doi: 10.1098/rstb.2011.0271

BYSTRIAKOVA N, Peregrym M, Erkens RHJ, Bezsmertna O, SCHNEIDER H (2012) Sampling bias in geographic and environmental space and its effect on the predictive power of species distribution models. Systematics and Biodiversity, 10 (3): 305 - 315. doi: 10.1080/14772000.2012.705357

Heinrichs J, Dong SS, Yu Y, Schaefer-Verwimp A, Pocs T, Feldberg K, Hentschel J, Schmidt AR, SCHNEIDER H (2012) A 150 year-old mystery solved: transfer of the rheophytic endemic liverwort Mycriocolea irrorata to Colura. Phytotaxa, 66 : 55 - 64.

Liu HM, DYER RJ, Go ZY, Meng Z, Li JH, SCHNEIDER H (2012) The evolutionary dynamic of apomixis in ferns: a case study from polystichoid ferns. Journal of Botany, 2012 doi: 10.1155/2012/510478

SCHNEIDER H (2012) Evolutioanry morphology of ferns (moilophytes). Annual Plant Reviews, 45 : 115 - 140.

Fiz-Palacios O, SCHNEIDER H, Heinrichs J, Savolainen V (2011) Diversification of land plants: insights from a family-level phylogenetic analysis. BMC Evolutionary Biology, 11 (1): 341 - 341. doi: 10.1186/1471-2148-11-341

Eiserhardt WL, Rohwer JG, RUSSELL SJ, YESILYURT JC, SCHNEIDER H (2011) Evidence for radiations of cheilanthoid ferns in the Greater Cape Floristic Region. Taxon, 60 (5): 1269 - 1283.

GUEIDAN C, Ruibal C, De Hoog GS, SCHNEIDER H (2011) Rock-inhabiting fungi originated during periods of dry climate in the late Devonian and middle Triassic. Fungal Biology, 115 (10): 987 - 996. doi: 10.1016/j.funbio.2011.04.002

ANSELL SW, Stenøien HK, GRUNDMANN M, RUSSELL SJ, Koch MA, SCHNEIDER H, VOGEL JC (2011) The importance of Anatolian mountains as the cradle of global diversity in Arabis alpina, a key arctic-alpine species. Annals of Botany, 108 (2): 241 - 252. doi: 10.1093/aob/mcr134

HUNT HV, ANSELL SW, RUSSELL SJ, SCHNEIDER H, VOGEL JC (2011) Dynamics of polyploid formation and establishment in the allotetraploid rock fern Asplenium majoricum. Annals of Botany, 108 (1): 143 - 157. doi: 10.1093/aob/mcr118

CHRISTENHUSZ MJM, Zhang XC, SCHNEIDER H (2011) A linear sequence of extant families and genera of lycophytes and ferns. Phytotaxa, 19 : 7 - 54.

De Groot GA, During HJ, Maas JW, SCHNEIDER H, VOGEL JC, Erkens RHJ (2011) Use of rbcL and trnL-F as a two-locus DNA barcode for identification of NW-European ferns: an ecological perspective. PLoS ONE, 6 (1)doi: 10.1371/journal.pone.0016371

BYSTRIAKOVA N, SCHNEIDER H, Coomes D (2011) The evolution of climatic niche in scaly tree ferns (Cyatheaceae, Polypodiopsoda). Botanical Journal of the Linnean Society, 165 (1): 1 - 19. doi: 10.1111/j.1095-8339.2010.01092.x

Challen G, VORONTSOVA MS, SCHNEIDER H, Cheek M (2011) Phylogenetically distinct and critically endangered new tree species of Phyllanthus from Cameroon (Phyllanthaceae, Euphorbiaceae s. l.). Systematic Botany, 36 (4): 933 - 938. doi: 10.1600/036364411X604949

Christenhusz MJM, Schneider H, others (2011) Corrections to Phytotaxa 19: Linear sequence of lycophytes and ferns. Phytotaxa, 28 : 50 - 52.

Hovenkamp PH, Fraser-Jenkins CR, SCHNEIDER H, Zhang XC (2011) (2002) Proposal to conserve the name Lepisorus against Belvisia, Lemmaphyllum, Paragramma, Drymotaenium & Neocheiropteris (Pteridophyta, Polypodiaceae). Taxon, 60

Sprunt SV, SCHNEIDER H, Watson LE, RUSSELL SJ, NAVARRO-GOMEZ A, Hickey RJ (2011) Exploring the molecular phylogeny and biogeography of Pleopeltis polypodioides (Polypodiaceae, Polypodiales) inferred from plastid DNA sequences. Systematic Botany, 36 (4): 862 - 869. doi: 10.1600/036364411X604886

WANG L, Wu ZQ, BYSTRIAKOVA N, ANSELL SW, Xiang QP, Heinrichs J, SCHNEIDER H, Zhang XC (2011) Phylogeography of the Sino-Himalayan Fern Lepisorus clathratus on "The Roof of the World". PLoS ONE, 6 (9)doi: 10.1371/journal.pone.0025896

ANSELL SW, Stenøien HK, GRUNDMANN M, SCHNEIDER H, Hemp A, Bauer N, RUSSELL SJ, VOGEL JC (2010) Population structure and historical biogeography of European Arabidopsis lyrata. Heredity, 105 (6)doi: 10.1038/hdy.2010.10

HENNEQUIN SABINE, HOVENKAMP PETER, CHRISTENHUSZ MAARTENJM, SCHNEIDER HARALD (2010) Phylogenetics and biogeography of Nephrolepis - a tale of old settlers and young tramps. Botanical Journal of the Linnean Society, 164 (2): 113 - 127. doi: 10.1111/j.1095-8339.2010.01076.x

Wubs ERJ, de Groot GA, During HJ, VOGEL JC, Grundmann M, Bremer P, SCHNEIDER H (2010) Mixed mating system in Asplenium scolopendrium L.: implications for colonization potential. Annals of Botany, 106 (4)doi: 10.1093/aob/mcq157

Yesilyurt JC, Schneider H (2010) The new fern genus Calciphilopteris (Pteridaceae). PHYTOTAXA, 7 : 52 - 59.

Schmidt AR, Perrichot V, Svotjtka M, Anderson KB, Belete KH, Doerfelt H, Jancke S, Mohr B, Mohrmann E, Nascimbeme PC, Nel A, Nel P, Pagazzi E, Roghi G, Saupe EE, Schmidt K, SCHNEIDER H, Selden PA, Vavra N (2010) Cretaceous African life captured in amber. Proceedings National Academy of Sciences, USA, 107 (16): 7329 - 7334. doi: 10.1073/pnas.1000948107

HENNEQUIN S, Ebihara A, Dubuisson JY, SCHNEIDER H (2010) Chromosome number evolution in Hymenophyllum (Hymenophyllaceae), with special reference to the subgenus Hymenophylum. Molecular Phylogenetics and Evolution, 55 (1)doi: 10.1016/j.ympev.2010.01.001

Grundmann M, Rumsey FJ, Ansell SW, Russell SJ, Darwin SC, Vogel JC, Spencer M, Squirrell J, Hollingsworth PM, Ortiz S, Schneider H (2010) Phylogeny and taxonomy of the bluebell genus Hyacinthoides, Asparagaceae [Hyacinthaceae]. TAXON, 59 (1): 68 - 82.

STRICKLUND-CONSTABLE R, SCHNEIDER H, ANSELL SW, RUSSELL SJ, KNAPP S (2010) Species identity in the Solanum bahamense complex (Solanaceae, Solanum subgenus Leptostemum). Taxon, 59 (1): 209 - 226.

SCHNEIDER H, Janssen T, BYSTRIAKOVA N, Heinrichs J, Hennequin S, Rakontondrainibe F (2010) Rapid radiations and neoendemism in the Madagascan biodiversity hotspot. , Glaubrecht M, Evolution in Action - Adaptive Radiations and the Origins of Biodiversity., pp. 3 - 15. Springer. Stuttgart.

SCHNEIDER H, Kreier HP, Janssen T, Otto E, Muth H, Heinrichs J (2010) Key innovations versus key opportunities: identifying causes of rapid radiations in derived ferns. , Glaubrecht M, Evolution in Action - Adaptive Radiations and the Origins of Biodiversity., pp. 61 - 75. Springer. Stuttgart.

Wang L, Qi X-P, Xiang Q-P, Heinrichs J, Schneider H, Zhang X-C (2010) Phylogeny of the paleotropical fern genus Lepisorus (Polypodiaceae, Polypodiopsida) inferred from four chloroplast DNA regions. Molecular Phylogenetics and Evolution, 54 (1): 211 - 225. doi: 10.1016/j.ympev.2009.08.032

WANG L, Wu ZQ, Xiang QP, Heinrichs J, SCHNEIDER H, Zhang XC (2010) A molecular phylogeny and a revised classification of tribe Lepisoreae (Polypodiaceae) based on an analysis of four plastid DNA regions. Botanical Journal of the Linnean Society, 162 (1): 28 - 38. doi: 10.1111/j.1095-8339.2009.01018.x

Hunt HV, ANSELL SW, RUSSELL SJ, SCHNEIDER H, VOGEL JC (2009) Genetic diversity and phylogeography in two diploid ferns, Asplenium fontanum subsp. fontanum and A. petrachae subsp. bivalens, in the western Mediterranean. Molecular Ecology, 18 (23): 4940 - 4954. doi: 10.1111/j.1365-294X.2009.04402.x

Heinrichs J, Klugmann F, Hentschel J, SCHNEIDER H (2009) DNA taxonomy, cryptic speciation and diversification of the Neotropical-African liverwort, Marchesinia brachiata (Lejeuneaceae, Porellales). Molecular Phylogenetics and Evolution, 53 (1)doi: 10.1016/j.ympev.2009.05.032

Otto EM, Janssen T, Kreier HP, SCHNEIDER H (2009) New insights into the phylogeny of Pleopeltis and related Neotropical genera (Polypodiaceae, Polypodiopsida). Molecular Phylogenetics and Evolution, 53 (1)doi: 10.1016/j.ympev.2009.05.001

Heinrichs J, Hentschel J, Feldberg K, Bombosch A, SCHNEIDER H (2009) Phylogenetic biogeography and taxonomy of disjunct distributed bryophytes. Journal of Systematics and Evolution, 47 (5)doi: 10.1111/j.1759-6831.2009.00028.x

Dubuisson JY, SCHNEIDER H, HENNEQUIN S (2009) Epiphytism in ferns: diversity and history. Comptes Rendus Biologies, 332 : 120 - 128. doi: 10.1016/j.crvi.2008.08.018

HENNEQUIN S, SCHNEIDER H, Ranker TA, Haufler CH (2009) Biology and Evolution of Ferns and Lycophytes (Book Review). Botanical Journal Linnean Society, 2009

Hentschel J, Von Konrat MJ, Pócs T, Schäfer-Verwimp A, Shaw AJ, SCHNEIDER H, Heinrichs J (2009) Molecular insights into the phylogeny and subgeneric classification of Frullania Raddi (Frullaniaceae, Porellales). Molecular Phylogenetics and Evolution, 52 : 142 - 156.

Hollingsworth P, Forrest L, Spouge J, Hajibabaei M, Ratnasingham S, Van der Bank M, Chase M, Cowan R, Erickson D, Fazekas A, Graham S, JAMES KE, Kim KJ, John W, SCHNEIDER H, Van Alphen Stahl J, Barrett S, Van den Berg C, Bogarin D, Burgess K, Cameron K, CARINE M, Chacón J, Clark A, Clarkson J, Conrad F, Devey D, Ford C, Hedderson T, Hollingsworth M, Husband B, Kelly L, Kesanakurti P, Kim J, Kim Y, Lahaye R, Lee HL, Long D, Madriñán S, Maurin O, Meusnier I, Newmaster S, Park CW, Percy D, Petersen G, Richardson J, Salazar G, Savolainen V, Seberg O, WILKINSON M, Yi DK, Little D (2009) A DNA Barcode for Land Plants. CBOL Plant Working Group. Proceedings of the National Academy of Sciences, USA, 106 : 12794 - 12797.

SCHNEIDER H, Smith AR, Pryer KM (2009) Is morphology really at odds with molecules in estimating fern phylogeny?. Systematic Botany, 34 (3): 455 - 475. doi: 10.1600/036364409789271209

SCHNEIDER H (2009) Taschenlehrbuch-Reihe Biologie [Chapter 7 Evolution]. , Munk K, Band 6 Evolution-Ökologie.. Georg-Thieme Verlag. Stuttgart.

Kreier HP, Zhang XR, Muth H, SCHNEIDER H (2008) The microsoroid ferns: inferring the relationships of a highly diverse lineage of Paleotropical epiphytic ferns (Polypodiacaee, Polypdiidae). Molecular Phylogenetics and Evolution, 48 (3): 1155 - 1167. doi: 10.1016/j.ympev.2008.05.001

Janssen T, Bystriakova N, Rakotondrainibe F, Coomes D, Labat JN, SCHNEIDER H (2008) Neoendemism in Madagascan tree ferns results from recent, coincident diversification bursts. Evolution, 62 (8): 1876 - 1889. doi: 10.1111/j.1558-5646.2008.00408.x

Kreier H-P, Rex M, Weising K, Kessler M, Smith AR, Schneider H (2008) Inferring the diversification of the epiphytic fern genus Serpocaulon (Polypodiaceae) in South America using chloroplast sequences and amplified fragment length polymorphisms. Plant Systematics and Evolution, 274 : 1 - 16. doi: 10.1007/s00606-008-0021-3

SCHNEIDER H, Kreier HP, Janssen T, Hovenkamp P (2008) Phylogenetic relationships of the fern genus Chrstiopteris shed new light onto the classification and biogeography of drynarioid ferns. Botanical Journal Linnean Society, 157 (4): 645 - 656. doi: 10.1111/j.1095-8339.2008.00825.x

ANSELL SW, GRUNDMANN M, RUSSELL SJ, SCHNEIDER H, VOGEL JC (2008) Genetic discontinuity, breeding-system change and population history of Arabis alpina in the Italian Peninsular and adjacent Alps. Molecular Ecology, 17 (9): 2245 - 2257. doi: 10.1111/j.1365-294X.2008.03739.x

JAMES KE, SCHNEIDER H, ANSELL SW, Evers M, ROBBA L, Uszynski G, PEDERSEN N, NEWTON AE, RUSSELL SJ, VOGEL JC, Kilian A (2008) Diversity Arrays Technology (DArT) for Pan-Genomic Evolutionary Studies of Non-Model Organisms. PLoS ONE, 3 (2)doi: 10.1371/journal.pone.0001682

Salino A, Almeida TE, Smith AR, Gómez AN, Kreier H-P, Schneider H (2008) A New Species of Microgramma (Polypodiaceae) from Brazil and Recircumscription of the Genus Based on Phylogenetic Evidence. Systematic Botany, 33 (4): 630 - 635. doi: 10.1600/036364408786500208

SCHNEIDER H, Dietz KJ (2008) Taschenlehrbuch-Reihe Biologie. [Chapter 1: Das Pflanzenreich, chapter 2: Landgang, chapter 3: Samenpflanzen and chapter 4: Fortpflanzung und Vermehrung (Co. authored by U. Rothe]. , Munk K, Band 3 Botanik.. Georg-Thieme Verlag. Stuttgart.

Smith AR, Pryer KM, Schuettpelz E, Korall P, SCHNEIDER H, Wolf PG (2008) Fern Classification. , Haufler CH RTA, Biology and Evolution of Ferns and Lycophytes., pp. 417 - 467. Cambridge University Press.

ANSELL S, SCHNEIDER H, PEDERSEN N, GRUNDMANN M, RUSSEL S, VOGEL J (2007) Recombination diversifies chloroplast trnF pseudogenes in Arabidopsis lyrata.. Journal of Evolutionary Biology, 20 (6): 2400 - 2411. doi: 10.1111/j.1420-9101.2007.01397.x

Hentschel J, Zhu RL, Long DG, Davison PG, SCHNEIDER H, R. G, Heinrichs J (2007) Phylogeny of Porella (Porellaceae, Jungermanniopsida) based on nuclear and chloroplast DNA sequences. Molecular Phylogenetics and Evolution, 45 (2): 693 - 705. doi: 10.1016/j.ympev.2007.05.005

Metzgar J, SCHNEIDER H, Pryer KM (2007) Phylogeny and divergence time estimates for the fern genus Azolla (Salviniaceae). International Journal of Plant Sciences, 168 (7): 1045 - 1053. doi: 10.1086/519007

Gradstein RS, Schwertfeger M, Kessler , Wissemann V, SCHNEIDER H (2007) The Old Botanical Garden of the University of Göttingen - recent developments and scientific research. Museum Aktuell, 136 : 11 - 15.

Heinrichs J, Hentschel J, Wilson R, Feldberg K, SCHNEIDER H (2007) Evolution of leafy liverworts (Jungermanniidae, Marchantiophyta): estimating divergence times from chloroplast DNA sequences using penalized likelihood with integrated fossil evidence. Taxon, 56 : 31 - 44.

Hentschel J, Feldberg K, Zündorf HJ, Hellwig FH, SCHNEIDER H, Heinrichs J (2007) The systematic position of Pachyglossa and Clasmatocolea (Jungermanniopsida: Lophocoleaceae) inferred from nrDNA ITS sequences and morphology. Taxon, 56 : 1136 - 1142.

Hentschel J, Paton JG, SCHNEIDER H, Heinrichs J (2007) Acceptance of Liochlaena Nees and Solenostema Mitt., the systematic position of Eromonotus Pearson and notes on Jungermannia L. s.l. (Jungermanniidae) based on chloroplast DNA sequence data. Plant Systematics and Evolution, 268 : 147 - 157.

Janssen T, Kreier HP, SCHNEIDER H (2007) Origin and diversification of African ferns with special emphasis on Polypodiaceae. Brittonia, 59 : 159 - 181.

Korall P, Metzgar JS, Conant DS, SCHNEIDER H, Pryer KM (2007) Phylogeny of scaly tree ferns (Cyatheaceae) as revealed by five plastid loci. American Journal of Botany, 94 : 873 - 886.

Kreier HP, Rojas-Alvarado AF, Smith AR, SCHNEIDER H (2007) Hyalotrichopteris is indeed a Campyloneurum (Polypodiaceae). American Fern Journal, 97 (3): 127 - 135. doi: 10.1640/0002-8444(2007)97[127:HIIACP]2.0.CO;2

Nagalingum NS, SCHNEIDER H, Pryer KM (2007) Molecular phylogenetic relationships and morphological evolution in the heterosporous fern genus Marsilea. Systematic Botany, 32 : 16 - 25.

SCHNEIDER H (2007) Plant morphology as the cornerstone to the integration of fossil sand extant taxa in phylogenetic analyses. Species, Phylogeny and Evolution, 1 : 65 - 71.

Schuettplez E, SCHNEIDER H, Huiet L, Windham MD, Pryer KM (2007) A molecular phylogeny of the fern family Pteridaceae: assessing overall relationships and the affinities of previously unsampled genera. Molecular Phylogenetics and Evolution, 44 : 1172 - 1185.

Wilson R, Gradstein SR, SCHNEIDER H, Heinrichs J (2007) Unravelling the phylogeny of Lejeuneaceae (Jungermanniopsida): evidence for four main lineages. Molecular Phylogenetics and Evolution, 43 : 270 - 282.

Wilson R, Heinrichs J, Gradstein SR, SCHNEIDER H (2007) Steady diversification of derived liverworts under Tertiary climatic fluctuations. Biology Letters, 3

GRUNDMANN M, Schneider H, RUSSELL SJ, VOGEL JC (2006) Phylogenetic relationships of the moss genus Pleurochaete Lindb. (Bryales: Pottiaceae) based on chloroplast and nuclear genomic markers. Organisms Diversity and Evolution, 6 (1): 33 - 45. doi: 10.1016/j.ode.2005.04.005

Hartmann FA, Wilson R, Gradstein SR, Schneider H, Heinrichs J (2006) Testing hypotheses on species delimitations and disjunctions in the liverwort Bryopteris (Jungermanniopsida: Lejeuneaceae). International Journal of Plant Sciences, 167 : 1205 - 1214.

Heinrichs J, Lindner M, Groth H, Hentschel J, Feldberg K, Renker C, Engel JJ, Von Konrat M, Long DG, Schneider H (2006) Goodbye or welcome Gondwana?–insights into the phylogenetic biogeography of the leafy liverwort Plagiochila with a description of Proskauera, gen. nov.(Plagiochilaceae, Jungermanniales). Plant Systematics and Evolution, 258 : 227 - 250.

Hentschel J, Wilson R, Burghardt M, Zündorf H-J, Schneider H, Heinrichs J (2006) Reinstatement of Lophocoleaceae (Jungermanniopsida) based on chloroplast gene rbcL data: exploring the importance of female involucres for the systematics of Jungermanniales. Plant Systematics and Evolution, 258 : 211 - 226.

Hu S, Dilcher DL, Schneider H, Jarzen DM (2006) Eusporangiate ferns from the Dakota formation, Minnesota, USA. International Journal of Plant Sciences, 167 : 579 - 589.

Korall P, Conant DS, Schneider H, Ueda K, Nishida H, Pryer KM (2006) On the Phylogenetic Position of Cystodium: It’s Not a Tree Fern-It’s a Polypod!. American Fern Journal, 96 : 45 - 53.

Korall P, Pryer KM, Metzgar JS, Schneider H, Conant DS (2006) Tree ferns: monophyletic groups and their relationships as revealed by four protein-coding plastid loci. Molecular phylogenetics and evolution, 39 : 830 - 845.

Kreier H-P, Schneider H (2006) Phylogeny and biogeography of the staghorn fern genus Platycerium (Polypodiaceae, Polypodiidae). American Journal of Botany, 93 : 217 - 225.

Kreier H-P, Schneider H (2006) Reinstatement of Loxogramme dictyopteris, based on phylogenetic evidence, for the New Zealand endemic fern, Anarthropteris lanceolata (Polypodiaceae, Polypodiidae). Australian Systematic Botany, 19 : 309 - 314.

Nagalingum NS, Schneider H, Pryer KM (2006) Comparative morphology of reproductive structures in heterosporous water ferns and a reevaluation of the sporocarp. International Journal of Plant Sciences, 167 : 805 - 815.

Schneider H (2006) Aleuritopteris. , Johns RJ, Edwards PJ, Utteridge TMA, & Hopkins HCC RJ, Guide to the Alpine and Sub-Alpine Flora of Mount Jaya.. Royal Botanic Gardens Kew.

Schneider H, Kreier H-P, Perrie LR, Brownsey PJ (2006) The relationships of Microsorum (Polypodiaceae) species occurring in New Zealand. New Zealand Journal of Botany, 44 : 121 - 127.

Schneider H, Kreier H-P, Wilson R, Smith AR (2006) The Synammia enigma: evidence for a temperate lineage of polygrammoid ferns (Polypodiaceae, Polypodiidae) in southern South America. Systematic Botany, 31 : 31 - 41.

SCHNEIDER H, Schuettpelz E (2006) Identifying fern gametophytes using DNA sequences. Molecular Ecology Notes, 6 : 989 - 991.

Smith AR, Kreier HP, Haufler CH, Ranker TA, SCHNEIDER H (2006) Serpocaulon (Polypodiaceae), a new genus segregated from Polypodium. Taxon, 55 : 919 - 930.

Smith AR, Pryer KM, Schuettpelz E, Korall P, Schneider H, Wolf PG (2006) A classification for extant ferns. Taxon, : 705 - 731.

Schneider H, Ranker TA, Russell SJ, Cranfill R, Geiger JMO, Aguraiuja R, Wood KR, Grundmann M, Kloberdanz K, Vogel JC (2005) Origin of the endemic fern genus Diellia coincides with the renewal of Hawaiian terrestrial life in the Miocene. Proceedings of the Royal Society B: Biological Sciences, 272 (1561): 455 - 460. doi: 10.1098/rspb.2004.2965

Heinrichs J, Gradstein SR, Wilson R, Schneider H (2005) Towards a natural classification of liverworts (Marchantiophyta) based on the chloroplast gene rbcL. Cryptogamie. Bryologie, 26 : 131 - 150.

Janssen T, Schneider H (2005) Exploring the evolution of humus collecting leaves in drynarioid ferns (Polypodiaceae, Polypodiidae) based on phylogenetic evidence. Plant Systematics and Evolution, 252 : 175 - 197.

Schneider H, Smith AR, Cranfill R, Hildebrand TJ, Haufler CH, Ranker TA (2004) Unraveling the phylogeny of polygrammoid ferns (Polypodiaceae and Grammitidaceae): exploring aspects of the diversification of epiphytic plants. Molecular Phylogenetics and Evolution, 31 (3): 1041 - 1063. doi: 10.1016/j.ympev.2003.09.018

Groth-Malonek M, Heinrichs J, Schneider H, Gradstein SR (2004) Phylogenetic relationships in the Lejeuneaceae (Hepaticae) inferred using ITS sequences of nuclear ribosomal DNA. Organisms Diversity & Evolution, 4 : 51 - 57.

Pryer KM, Schneider H, Magallón S (2004) The radiation of vascular plants. Assembling the Tree of Life. Oxford Univ. Press, New York, : 138 - 153.

Pryer KM, Schuettpelz E, Wolf PG, Schneider H, Smith AR, Cranfill R (2004) Phylogeny and evolution of ferns (monilophytes) with a focus on the early leptosporangiate divergences. American Journal of Botany, 91 : 1582 - 1598.

Ranker TA, Smith AR, Parris BS, Geiger JMO, Haufler CH, Straub SCK, Schneider H (2004) Phylogeny and evolution of grammitid ferns (Grammitidaceae): a case of rampant morphological homoplasy. Taxon, 53 : 415 - 415.

Schneider H, Janssen T, Hovenkamp P, Smith AR, Cranfill R, Haufler CH, Ranker TA (2004) Phylogenetic relationships of the enigmatic Malesian fern Thylacopteris (Polypodiaceae, Polypodiidae). International Journal of Plant Sciences, 165 : 1077 - 1087.

Schneider H, RUSSELL SJ, Cox C, Bakker F, HENDERSON S, RUMSEY FJ, Barrett JA, Gibby M, VOGEL JC (2004) Chloroplast phylogeny of asplenioid ferns based on rbcL and trnL-F spacer sequences (Polypodiidae, Aspleniaceae) and its implications for the biogeography of these ferns. Systematic Botany, 29 (2): 260 - 274. doi: 10.1600/036364404774195476

Schneider H, Schuettpelz E, Pryer KM, Cranfill R, Magallón S, Lupia R (2004) Ferns diversified in the shadow of angiosperms. Nature, 428 : 553 - 557.

DUBUISSON J-Y, Hennequin S, Rakotondrainibe F, Schneider H (2003) Ecological diversity and adaptive tendencies in the tropical fern Trichomanes L.(Hymenophyllaceae) with special reference to climbing and epiphytic habits. Botanical Journal of the Linnean Society, 142 : 41 - 63.

Gradstein SR, REINER-DREHWALD ME, Schneider H (2003) A phylogenetic analysis of the genera of Lejeuneaceae (Hepaticae). Botanical Journal of the Linnean Society, 143 : 391 - 410.

Haufler CH, Grammer WA, Hennipman E, Ranker TA, Smith AR, Schneider H (2003) Systematics of the ant-fern genus Lecanopteris (Polypodiaceae): testing phylogenetic hypotheses with DNA sequences. Systematic Botany, 28 : 217 - 227.

Schneider H (2003) Pteris. , Amoroso V, Plant Resources of South East Asia - Volume 15.. Backhuys Publishers. Leiden.

Schneider H (2003) Taenitis blechnoides. , Amoroso V, Plant Resources of South East Asia - Volume 15.. Blackhuys Publisher. Leiden.

Pryer KM, Schneider H, Zimmer EA, Ann Banks J (2002) Deciding among green plants for whole genome studies. Trends in Plant Science, 7 (12): 550 - 554. doi: 10.1016/S1360-1385(02)02375-0

Pinter I, Bakker F, Barrett J, Cox C, Gibby M, Henderson S, Morgan-Richards M, Rumsey F, Russell S, Trewick S, Schneider H, Vogel J (2002) Phylogenetic and biosystematic relationships in four highly disjunct polyploid complexes in the subgenera Ceterach and Phyllitis in Asplenium (Aspleniaceae). ORGANISMS DIVERSITY & EVOLUTION, 2 (4): 299 - 311. doi: 10.1078/1439-6092-00050

Schneider H, Pryer KM (2002) Structure and function of spores in the aquatic heterosporous fern family Marsileaceae. International Journal of Plant Sciences, 163 : 485 - 505.

Schneider H, Pryer KM, Cranfill R, Smith AR, Wolf PG (2002) Evolution of vascular plant body plans: a phylogenetic perspective. SYSTEMATICS ASSOCIATION SPECIAL VOLUME, 65 : 330 - 364.

Schneider H, Smith AR, Cranfill R, Haufler CH, Ranker TA, Hildebrand T (2002) Gymnogrammitis dareiformis is a polygrammoid fern (Polypodiaceae)–resolving an apparent conflict between morphological and molecular data. Plant Systematics and Evolution, 234 : 121 - 136.

Schneider H, Kenrick P (2001) An Early Cretaceous root-climbing epiphyte (Lindsaeaceae) and its significance for calibrating the diversification of polypodiaceous ferns. Review of Palaeobotany and Palynology, 115 : 33 - 41. doi: 10.1016/S0034-6667(01)00048-3

Pryer KM, Schneider H, Smith AR, Cranfill R, Wolf PG, Hunt JS, Sipes SD (2001) Horsetails and ferns are a monophyletic group and the closest living relatives to seed plants. Nature, 409 : 618 - 622.

Wolf PG, Schneider H, Ranker TA (2001) Geographic distributions of homosporous ferns: does dispersal obscure evidence of vicariance?. Journal of Biogeography, 28 : 263 - 270.

Lupia R, Schneider H, Moeser GM, Pryer KM, Crane PR (2000) Marsileaceae sporocarps and spores from the Late Cretaceous of Georgia, USA. International Journal of Plant Sciences, 161 : 975 - 988.

SCHNEIDER H (2000) Morphology and anatomy of roots in the filmy fern tribe Trichomaneae H. Schneider (Hymenophyllaceae, Filicatae) and the evolution of rootless taxa. Botanical Journal of the Linnean Society, 132 : 29 - 46.

Wollenweber E, Schneider H (2000) Lipophilic exudates of Pteridaceae–chemistry and chemotaxonomy. Biochemical systematics and ecology, 28 : 751 - 777.

Schneider H (1999) YET ANOTHER FERN WITH STORAGE ROOTS-CHEILANTHES BOLBORRHIZA MICKEL & BEITEL (PTERIDACEAE: PTERIDOPHYTA) FROM MEXICO AND EL SALVADOR. Fern Gaz, 15 : 269 - 272.

Schneider H (1998) Biogeography, ecology and phylogeny of Pteridaceae (Filicatae) in Malesia: Taenitidoideae and Cheilanthoideae. Taxonomy: the Cornerstone of Biodiversity. Proceedings of the Fourth International Flora Malesiana Symposium, : 211 - 223.

Schneider H (1997) Root anatomy of Aspleniaceae and the implications for systematics of this fern family. Fern Gazette (United Kingdom),

Schneider H. , (1996) Vergleichende Wurzelanatomie der Farne. null. 171

Schneider H (1995) The root anatomy of ferns: a comparative study. Pteridology in perspective: pteridophyte symposium’95. Proceedings of the Holttum memorial pteridophyte symposium, Kew, : 271 - 283.

Highlighted publications

Schneider H, Liu H, Clark J, Hidalgo O, Pellicer J, et al (2015) Are the genomes of royal ferns really frozen in time? Evidence for coinciding genome stability and limited evolvability in the royal ferns. New Phytologist, 207 (1): 10 - 13. doi: 10.1111/nph.13330

Laenen B, Shaw B, Schneider H, Goffinet B, Paradis E, et al (2014) Extant diversity of bryophytes emerged from successive post-Mesozoic diversification bursts. Nature Communications, 5 : 5134 - 5134. doi: 10.1038/ncomms6134

Hennequin S, Kessler M, Lindsay S, Schneider H (2014) Evolutionary patterns in the assembly of fern diversity on the oceanic Mascarene Islands. Journal of Biogeography, 41 (9): 1651 - 1663. doi: 10.1111/jbi.12339

Feldberg K, Schneider H, Stadler T, Schäfer-Verwimp A, Schmidt AR, et al (2014) Epiphytic leafy liverworts diversified in angiosperm-dominated forests. Scientific Reports, 4 doi: 10.1038/srep05974

Bystriakova N, Ansell SW, Russell SJ, Grundmann M, Vogel JC, et al (2014) Present, past and future of the European rock fern Asplenium fontanum: combining distribution modelling and population genetics to study the effect of climate change on geographic range and genetic diversity. Annals of Botany, 113 (3): 453 - 465. doi: 10.1093/aob/mct274

SCHNEIDER H, Smith AR, Pryer KM (2009) Is morphology really at odds with molecules in estimating fern phylogeny?. Systematic Botany, 34 (3): 455 - 475. doi: 10.1600/036364409789271209