Dr Alan Warren

Researcher

Department: Life Sciences
Division: LS Genomics and Microbial Biodiversity Division
Contact: email
Phone: +44 (0)20 7942 5149 5149

Summary

Main reserach interests are the systematics and biodiversity of ciliated protists (ciliates) inlcuding: (1) morphology and morphogenesis; (2) molecular phylogeny; (3) use as bioindicators of water quality. The main focus of this reserach is the marine ciliate fauna of coastal waters of China.

Qualifications

Degrees

PhD, University of Surrey, UK, 1977 - 1983

Employment history

Academic

Researcher, Protists, Natural History Museum, Department of Life Sciences, UK, 1979 - ongoing

Publications

Esteban G, Finlay BJ, Warren A (2015) Free-living Protozoa. , Thorp J, Ecology and General Biology: Thorp and Covich’s Freshwater Invertebrates., pp. 111 - 132. Academic Press.

Chen L, Zhao X, Ma H, Warren A, Shao C, Huang J (2015) Morphology, morphogenesis and molecular phylogeny of a soil ciliate, Pseudouroleptus caudatus caudatus Hemberger, 1985 (Ciliophora, Hypotricha), from Lhalu Wetland, Tibet. European Journal of Protistology, 51 (1): 1 - 14. doi: 10.1016/j.ejop.2014.09.001

Shao C, Chen L, Pan Y, Warren A, Miao M (2014) Morphology and phylogenetic position of the oxytrichid ciliates, Urosoma salmastra (Dragesco and Dragesco-Kernéis, 1986) Berger, 1999 and U. karinae sinense nov. sspec. (Ciliophora, Hypotrichia). European Journal of Protistology, 50 (5): 593 - 605. doi: 10.1016/j.ejop.2014.08.004

Lu X, Gao F, Shao C, Hu X, Warren A (2014) Morphology, morphogenesis and molecular phylogeny of a new marine ciliate, Trichototaxis marina n. sp. (Ciliophora, Urostylida). European Journal of Protistology, 50 (5): 524 - 537. doi: 10.1016/j.ejop.2014.08.002

Xu G, Zhong X, Wang Y, Warren A, Xu H (2014) A multivariate approach to the determination of an indicator species pool for community-based bioassessment of marine water quality. Marine Pollution Bulletin, 87 : 147 - 151. doi: 10.1016/j.marpolbul.2014.07.068

Shi X, Meng Q, Liu G, Qi G, Jiang C, Meng X, Warren A (2014) Morphology and morphogenesis of Epistylis plicatilis ehrenberg, 1831 (Ciliophora, Peritrichia) from Wuhan, China. Journal of Morphology, 275 (8): 882 - 893. doi: 10.1002/jmor.20265

Li J, Liu W, Gao S, Warren A, Song W (2014) Multigene-Based Analyses of the Phylogenetic Evolution of Oligotrich Ciliates, with Consideration of the Internal Transcribed Spacer 2 Secondary Structure of Three Systematically Ambiguous Genera. Eukaryotic Cell, 13 (7): 947 - 947. doi: 10.1128/EC.00120-14

Wu L, Chen R, Yi Z, Li J, Warren A, Lin X (2014) The Morphology of Three Loxophyllum Species (Ciliophora, Pleurostomatida) from Southern China, L. lembum sp. n., L. vesiculosum sp. n. and L. perihoplophorum Buddenbrock, 1920, with Notes on the Molecular Phylogeny of Loxophyllum. Journal of Eukaryotic Microbiology, 61 (2): 115 - 125. doi: 10.1111/jeu.12089

Gong J, Qing Y, Guo X, Warren A (2014) “Candidatus Sonnebornia yantaiensis”, a member of candidate division OD1, as intracellular bacteria of the ciliated protist Paramecium bursaria (Ciliophora, Oligohymenophorea). Systematic and Applied Microbiology, 37 (1): 35 - 41. doi: 10.1016/j.syapm.2013.08.007

Shao C, Lv Z, Jin L, Warren A (2014) Morphogenesis of a unique pseudourostylid ciliate, Trichototaxis songi (Ciliophora, Urostylida). European Journal of Protistology, 50 (1): 68 - 77. doi: 10.1016/j.ejop.2013.09.002

Jiang Y, Xu H, Warren A (2014) Insights into discriminating environmental quality status using taxonomic distinctness based on a small species pool of ciliated protozoa in marine ecosystems. Science of The Total Environment, : 663 - 670. doi: 10.1016/j.scitotenv.2013.08.083

Dunthorn M, Stoeck T, Clamp J, Warren A, Mahé F (2014) Ciliates and the Rare Biosphere: A Review. Journal of Eukaryotic Microbiology, 61 (4): 404 - 409. doi: 10.1111/jeu.12121

Fan Y, Pan Y, Huang J, Lin X, Hu X, Warren A (2014) Molecular Phylogeny and Taxonomy of Two Novel Brackish Water Hypotrich Ciliates, with the Establishment of a New Genus, Antiokeronopsis gen. n. (Ciliophora, Hypotrichia). Journal of Eukaryotic Microbiology, 61 (5): 449 - 462. doi: 10.1111/jeu.12125

Warren A (2014) Protozoa. eLS, doi: 10.1002/9780470015902.a0001924.pub3

Zhang Q, Yi Z, Fan X, Warren A, Gong J, Song W (2014) Further insights into the phylogeny of two ciliate classes Nassophorea and Prostomatea (Protista, Ciliophora). Molecular Phylogenetics and Evolution, 70 : 162 - 170. doi: 10.1016/j.ympev.2013.09.015

Liu W, Yi Z, Li J, Warren A, Al-Farraj SA, Lin X (2013) Taxonomy, morphology and phylogeny of three new oligotrich ciliates (Protozoa, Ciliophora, Oligotrichia) from southern China. International Journal of Systematic and Evolutionary Microbiology, 63 : 4805 - 4817. doi: 10.1099/ijs.0.052878-0

Ma B, Lv X, Warren A, Gong J (2013) Shifts in diversity and community structure of endophytic bacteria and archaea across root, stem and leaf tissues in the common reed, Phragmites australis, along a salinity gradient in a marine tidal wetland of northern China. Antonie Van Leeuwenhoek International Journal of General and Molecular Microbiology, 104 (5): 759 - 768. doi: 10.1007/s10482-013-9984-3

Lv Z, Chen L, Chen L, Shao C, Miao M, Warren A (2013) Morphogenesis and Molecular Phylogeny of a New Freshwater Ciliate, Notohymena apoaustralis n. sp (Ciliophora, Oxytrichidae). Journal of Eukaryotic Microbiology, 60 (5): 455 - 466. doi: 10.1111/jeu.12053

Xu Y, Li J, Song W, Warren A (2013) Phylogeny and Establishment of a New Ciliate Family, Wilbertomorphidae fam. nov (Ciliophora, Karyorelictea), a Highly Specialized Taxon Represented by Wilbertomorpha colpoda gen. nov., spec. nov. Journal of Eukaryotic Microbiology, 60 (5): 480 - 489. doi: 10.1111/jeu.12055

Yan Y, Xu Y, Yi Z, Warren A (2013) Redescriptions of three trachelocercid ciliates (Protista, Ciliophora, Karyorelictea), with notes on their phylogeny based on small subunit rRNA gene sequences. International Journal of Systematic and Evolutionary Microbiology, 63 : 3506 - 3514. doi: 10.1099/ijs.0.053959-0

Jiang Y, Xu H, Hu X, Warren A, Song W (2013) Functional groups of marine ciliated protozoa and their relationships to water quality. Environmental Science and Pollution Research, 20 (8): 5272 - 5280. doi: 10.1007/s11356-013-1525-0

Jiang J, Xing Y, Miao M, Shao C, Warren A, Song W (2013) Two New Marine Ciliates, Caryotricha rariseta n. sp and Discocephalus pararotatorius n. sp (Ciliophora, Spirotrichea), with Phylogenetic Analyses Inferred from the Small Subunit rRNA Gene Sequences. Journal of Eukaryotic Microbiology, 60 (4): 388 - 398. doi: 10.1111/jeu.12046

Chen X, Yan Y, WARREN A, Ma H, Hu X (2013) Morphology and morphogenesis of a soil ciliate, Cyrtohymena candens (Kahl, 1932) Foissner, 1989 (Ciliophora, Hypotrichia, Oxytrichidae), with notes on its molecular phylogeny based on SSU rDNA sequence data. International Journal of Systematic and Evolutionary Microbiology, 63 : 1912 - 1921. doi: 10.1099/ijs.0.048611-0

Fan X, Lin X, Liu W, Xu Y, Al-Farraj SA, Al-Rasheid KAS, Warren A (2013) Morphology of three new marine Frontonia species (Ciliophora; Peniculida) with note on the phylogeny of this genus. European Journal of Protistology, 49 (2): 312 - 323. doi: 10.1016/j.ejop.2012.06.003

Pan Y, Li J, Li L, Hu X, Al-Rasheid KAS, Warren A (2013) Ontogeny and molecular phylogeny of a new marine ciliate genus, Heterokeronopsis g. n. (Protozoa, Ciliophora, Hypotricha), with description of a new species. European Journal of Protistology, 49 (2): 298 - 311. doi: 10.1016/j.ejop.2012.08.008

Shao C, Pan X, Jiang J, Ma H, Al-Rasheid KAS, Warren A, Lin X (2013) A redescription of the oxytrichid Tetmemena pustulata (Muller, 1786) Eigner, 1999 and notes on morphogenesis in the marine urostylid Metaurostylopsis salina Lei et al., 2005 (Ciliophora, Hypotrichia). European Journal of Protistology, 49 (2): 272 - 282. doi: 10.1016/j.ejop.2012.07.003

Fan Y, WARREN A, Al-Farraj SA, Chen X, Shao C (2013) Morphology and SSU rRNA gene-based phylogeny of two Diophrys-like ciliates from northern china, with notes on morphogenesis of Pseudodiophrys nigricans (Protozoa, Ciliophora). Journal of Morphology, 274 (4): 395 - 403. doi: 10.1002/jmor.20097

Li J, Liu W, Gao S, WARREN A, Song W (2013) Multigene-Based Analyses of the Phylogenetic Evolution of Oligotrich Ciliates, with Consideration of the Internal Transcribed Spacer 2 Secondary Structure of Three Systematically Ambiguous Genera. Eukaryotic Cell, 12 (3): 430 - 437. doi: 10.1128/ec.00270-12

Tang F, Zhao Y, WARREN A (2013) Phylogenetic Analyses of Trichodinids (Ciliophora, Oligohymenophora) Inferred from 18S rRNA Gene Sequence Data. Current Microbiology, 66 (3): 306 - 313. doi: 10.1007/s00284-012-0274-5

Xu HL, Jiang Y, Zhang W, Zhu MZ, Al-Rasheid KAS, WARREN A (2013) Annual variations in body-size spectra of planktonic ciliate communities and their relationships to environmental conditions: a case study in Jiaozhou Bay, northern China. Journal of the Marine Biological Association of the United Kingdom, 93 (1): 47 - 55. doi: 10.1017/s0025315412001075

Pan HB, Gao F, Lin XF, WARREN A, Song WB (2013) Three New Loxophyllum Species (Ciliophora: Pleurostomatida) from China with a Brief Review of the Marine and Brackish Loxophyllum Species. Journal of Eukaryotic Microbiology, 60 (1): 44 - 56. doi: 10.1111/jeu.12005

Pan X, Gao F, Liu W, Fan X, Warren A, Song W (2013) Morphology and SSU rRNA gene sequences of three Frontonia species, including a description of F. subtropica spec. nov (Ciliophora, Peniculida). European Journal of Protistology, 49 (1): 67 - 77. doi: 10.1016/j.ejop.2012.05.002

Shao C, Ding Y, Al-Rasheid KA, Al-Farraj SA, Warren A, Song W (2013) Establishment of a new hypotrichous genus, Heterotachysoma n. gen. and notes on the morphogenesis of Hemigastrostyla enigmatica (Ciliophora, Hypotrichia). European Journal of Protistology, 49 (1): 93 - 105. doi: 10.1016/j.ejop.2012.04.002

Warren A (2013) Protozoology. eLS, doi: 10.1002/9780470015902.a0001923.pub3

Wu L, Chen R, Yi Z, Li J, WARREN A, Lin X (2013) Morphology and Phylogeny of Three New Loxophyllum Species (Ciliophora, Pleurostomatida) from Mangrove Wetlands of Southern China. Journal of Eukaruyotic Microbiology, 60 (3): 267 - 281. doi: 10.1111/jeu.12032

Xu Y, Li J, WARREN A, Song W (2013) Molecular phylogeny and establishment of a new ciliate family, Wilbertomorphidae fam. nov. (Ciliophora, Karyorelictea), a highly specialized taxon represented by Wilbertomorpha colpoda gen. nov., spec. nov.. Protist,

Xu D, Sun P, WARREN A, Noh JH, Choi DL, Shin MK, Kim YO (2013) Phylogenetic Investigations on Ten Genera of Tintinnid Ciliates (Ciliophora: Spirotrichea: Tintinnida), Based on Small Subunit Ribosomal RNA Gene Sequences. Journal of Eukaryotic Microbiology, 60 (2): 192 - 202. doi: 10.1111/jeu.12023

Appeltans W, Ahyong ST, Anderson G, Angel MV, Artois T, Bailly N, Bamber R, Barber A, Bartsch I, Berta A, Blazewicz-Paszkowycz M, Bock P, BOXSHALL G, Boyko CB, Brandao SN, BRAY RA, Bruce NL, Cairns SD, Chan T, Cheng L, Collins AG, Cribb T, Curini-Galletti M, Dandouh-Guebas F, Davie PJF, Dawson MN, De Clerck O, Decock W, De Grave S, de Voogd NJ, Domning DP, Emig CC, Erseus C, Eschmeyer W, Fauchald K, Fautin DG, Feist SW, Fransen CHJM, Furuya H, Garcia-Alvarez O, Gerken S, GIBSON D, Gittenberger A, Gofas S, Gomez-Daglio L, Gordon DP, Guiry MD, Hernandez F, Hoeksema BW, Hopcroft RR, Jaume D, Kirk P, Koedam N, Koenemann S, Kolb JB, Kristensen RM, Kroh A, Lambert G, Lazarus DB, Lemaitre R, Longshaw M, Lowry J, Macpherson E, Madin LP, Mah C, MAPSTONE G, McLaughlin PA, Mees J, Meland K, Messing CG, Mills CE, Molodtsova TN, Mooi R, Neuhaus B, Ng PKL, Nielsen C, Norenburg J, Opresko DM, Osawa M, Paulay G, Perrin W, Pilger JF, Poore GCB, Pugh P, Read GB, Reimer JD, Rius M, Rocha RM, Saiz-Salinas JI, Scarabino V, Schierwater B, Schmidt-Rhaesa A, Schnabel KE, Schotte M, Schuchert P, Schwabe E, Segers H, Self-Sullivan C, Shenkar N, Siegel V, Sterrer W, Stohr S, Swalla B, Tasker ML, Thuesen EV, Timm T, Todaro MA, Turon X, Tyler S, Uetz P, van der Land J, Vanhoorne B, van Ofwegen LP, van Soest RWM, Vanaverbeke J, Walker-Smith G, Walter TC, WARREN A, Williams GC, Wilson SP, Costello MJ (2012) The Magnitude of Global Marine Species Diversity. Current Biology, 22 (23): 2189 - 2202. doi: 10.1016/j.cub.2012.09.036

HU X, Huang J, WARREN A (2012) The morphology and phylogeny of two euplotid ciliates, Diophrys blakeneyensis spec. nov and Diophrys oligothrix Borror, 1965 (Protozoa, Ciliophora, Euplotida). International Journal of Systematic and Evolutionary Microbiology, 62 : 2757 - 2773. doi: 10.1099/ijs.0.039503-0

Jiang Y, Yang J, Al-Farraj SA, WARREN A, Lin X (2012) Redescriptions of three tintinnid ciliates, Tintinnopsis tocantinensis, T. radix, and T. cylindrica (Ciliophora, Spirotrichea), from coastal waters off China. European Journal of Protistology, 48 (4): 314 - 325. doi: 10.1016/j.ejop.2012.02.001

Liu WW, Yi ZZ, Lin XF, WARREN A, Song WB (2012) Phylogeny of three choreotrich genera (Protozoa, Ciliophora, Spirotrichea), with morphological, morphogenetic and molecular investigations on three strobilidiid species. Zoologica Scripta, 41 (4): 417 - 434. doi: 10.1111/j.1463-6409.2012.00542.x

Du YF, Xu KD, WARREN A, Lei YL, Dai RH (2012) Benthic ciliate and meiofaunal communities in two contrasting habitats of an intertidal estuarine wetland. Journal of Sea Research, 70 : 50 - 63. doi: 10.1016/j.seares.2012.03.004

Chen XR, Gao S, Liu WW, Song WB, Al-Rasheid KAS, WARREN A (2012) Taxonomic descriptions of three marine colepid ciliates, Nolandia sinica spec. nov., Apocoleps caoi spec. nov and Tiarina fusa (Claparède & Lachmann, 1858) Bergh, 1881 (Ciliophora, Prorodontida). International Journal of Systematic and Evolutionary Microbiology, 62 : 735 - 744. doi: 10.1099/ijs.0.026302-0

Mazei Y, WARREN A (2012) A survey of the testate amoeba genus Difflugia Leclerc, 1815 based on specimens in the E. Penard and C.G. Ogden collections of the Natural History Museum, London. Part 1: Species with shells that are pointed aborally and/or have aboral protuberances. Protistology, 7 : 121 - 171.

Pan Y, Li LQ, Shao C, Hu XZ, Ma HG, Al-Rasheid KAS, WARREN A (2012) Morphology and Ontogenesis of a Marine Ciliate, Euplotes balteatus (Dujardin, 1841) Kahl, 1932 (Ciliophora, Euplotida) and Definition of Euplotes wilberti nov spec. Acta Protozoologica, 51 (1): 29 - 38. doi: 10.4467/16890027ap.12.003.0386

Xu Y, Miao M, WARREN A, Song WB (2012) Diversity of the karyorelictid ciliates: Remanella (Protozoa, Ciliophora, Karyorelictida) inhabiting intertidal areas of Qingdao, China, with descriptions of three species. Systematics and Biodiversity, 10 (2): 207 - 219. doi: 10.1080/14772000.2012.681713

Zhao Y, Gao F, Li J, Yi Z, WARREN A (2012) Phylogenetic Analyses on the Tintinnid Ciliates (Protozoa, Ciliophora) Based on Multigene Sequence Data. Acta Protozoologica, 51 (4): 319 - 328. doi: 10.4467/16890027ap.12.025.0785

Chen R, Lin X, Warren A (2011) A new pleurostomatid ciliate, Amphileptus salignus n. sp (Protozoa, Ciliophora), from mangrove wetlands in southern China. ZOOTAXA, (3052): 62 - 68.

Chen RM, Lin XF, WARREN A (2011) A new pleurostomatid ciliate, Amphileptus salignus n. sp (Protozoa, Ciliophora), from mangrove wetlands in southern China. Zootaxa, 3048 (3048): 62 - 68.

Pan HB, HU XZ, Gong J, Lin XF, Al-Rasheid KAS, Al-Farraj SA, WARREN A (2011) Morphological redescriptions of four marine ciliates (Ciliophora: Cyrtophorida: Dysteriidae) from Qingdao, China. European Journal of Protistology, 47 (3): 197 - 207. doi: 10.1016/j.ejop.2011.03.002

Vasudevan P, Griffin P, WARREN A, Thapliyal A, Srivastava RK, Tandon M (2011) Localized domestic wastewater treatment: Part II - Irrigation potential in Indian scenario. Journal of Scientific & Industrial Research, 70 (8): 595 - 600.

Xu Y, Huang J, HU XZ, Al-Rasheid KAS, Song WB, WARREN A (2011) Taxonomy, ontogeny and molecular phylogeny of Anteholosticha marimonilata spec. nov (Ciliophora, Hypotrichida) from the Yellow Sea, China. International Journal of Systematic and Evolutionary Microbiology, 61 (8): 2000 - 2014. doi: 10.1099/ijs.0.024638-0

Xu Y, Huang J, WARREN A, Al-Rasheid KAS, Al-Farraj SA, Song WB (2011) Morphological and molecular information of a new species of Geleia (Ciliophora, Karyorelictea), with redescriptions of two Kentrophoros species from China. European Journal of Protistology, 47 (3): 172 - 185. doi: 10.1016/j.ejop.2011.03.003

Wang YG, HU XZ, Huang J, Al-Rasheid KAS, WARREN A (2011) Characterization of two urostylid ciliates, Metaurostylopsis flavicana spec. nov. and Tunicothrix wilberti (Lin & Song, 2004) Xu et al., 2006 (Ciliophora, Stichotrichia), from a mangrove nature protection area in China. International Journal of Systematic and Evolutionary Microbiology, 61 (7): 1740 - 1750. doi: 10.1099/ijs.0.024935-0

Fan X, Chen X, Song W, Al-Rasheid KAS, WARREN A (2011) Two novel marine Frontonia species, Frontonia mengi spec. nov and Frontonia magna spec. nov (Protozoa; Ciliophora), with notes on their phylogeny based on small-subunit rRNA gene sequence data. International Journal of Systematic and Evolutionary Microbiology, 61 (6): 1476 - 1486. doi: 10.1099/ijs.0.024794-0

Xu H, Jiang Y, Al-Rasheid KAS, Song W, WARREN A (2011) Spatial variation in taxonomic distinctness of ciliated protozoan communities at genus-level resolution and relationships to marine water quality in Jiaozhou Bay, northern China.. Hydrobiologia, 665 (1): 67 - 78. doi: 10.1007/s10750-011-0604-9

Jiang Y, Xu H, Al-Rasheid KAS, WARREN A, Xiaozhong H (2010) Planktonic ciliate communities in a semi-enclosed bay of Yellow Sea, northern China: annual cycle. Journal of the Marine Biological Association of the United Kingdom, 91 (1): 97 - 105. doi: 10.1017/S002531541000175X

Jiang Y, Xu H, Hu X, Zhu M, Al-Rasheid KAS, WARREN A (2011) An approach to analyzing spatial patterns of planktonic ciliate communities for monitoring water quality in Jiaozhou Bay, nothern China. Marine Pollution Bulletin, 62 (2): 227 - 235. doi: 10.1016/j.marpolbul.2010.11.008

Fan XP, Lin XF, Al-Rasheid KAS, Al-Farraj SA, WARREN A, Song WB (2011) The diversity of scuticociliates (Protozoa, Ciliophora): a report on eight marine forms found in coastal waters of China, with a description of one new species. Acta Protozoologica, 50 (3): 219 - 234.

Jiang JM, WARREN A, Song WB (2011) Morphology and molecular phylogeny of two new marine euplotids, Pseudodiophrys nigricans n. g., n. sp., and Paradiophrys zhangi n. sp (Ciliophora: Euplotida). Journal of Eukaryotic Microbiology, 58 (5): 437 - 445. doi: 10.1111/j.1550-7408.2011.00567.x

Liu W, Yi Z, WARREN A, Al-Rasheid KAS, Al-Farraj SA, Lin X, Song W (2011) Taxonomy, morphology and molecular systematics of a new oligotrich ciliate, Williophrya maedai gen. nov., sp. nov., with redescriptions of Strombidium basimorphum and pseudotontonia simplicidens (Protozoa, Ciliophora, Oligotrichia). Systematics and Biodiversity, 9 (3): 247 - 258. doi: 10.1080/14772000.2011.605812

Qin YH, Qiu ZJ, Shao C, WARREN A, Shen Z (2011) Morphological redescription and morphogenesis of Urosoma macrostyla (Wrzesniowski, 1866) Berger, 1999 (Ciliophora, Hypotrichida). Acta Protozoologica, 50 (3): 163 - 174.

Shao C, Gao F, HU X, Al-Rasheid KA, WARREN A (2011) Ontogenesis and molecular phylogeny of a new marine Urostylid ciliate, Anteholosticha petzi n. sp. (Ciliophora, Urostylida). Journal of Eukaryotic Microbiology, 58 (3): 254 - 265. doi: 10.1111/j.1550-7408.2011.00542.x

Shen Z, Yi Z, WARREN A (2011) The morphology, ontogeny, and small subunit rRNA gene sequence analysis of Diophrys parappendiculata n. sp. (Protozoa, Ciliophora, Euplotida), a new Marine ciliate from coastal waters of southern China. Journal of Eukaryotic Microbiology, 58 (3): 242 - 248. doi: 10.1111/j.1550-7408.2011.00536.x

Xu Y, Esaulov A, Lin XF, Mazei Y, HU XZ, Al-Rasheid KAS, WARREN A (2011) Morphological studies on five trachelocercids from the Yellow Sea Coast of China, with a description of Tracheloraphis huangi spec. nov. (Ciliophora, Karyorelictea). Acta Protozoologica, 50 (3): 205 - 218.

Xu H, Zhang W, Jiang Y, Zhu M, Al-Rasheid KAS, WARREN A, Song W (2011) An approach to determining the sampling effort for analyzing biofilm-dwelling ciliate colonization using an artificial substratum in coastal waters. Biofouling, 27 (4): 357 - 366. doi: 10.1080/08927014.2011.576340

Chen X, Wang Y, Long H, Al-Rasheid KAS, WARREN A, Song W (2009) Morphological studies on two marine colepid ciliates from Qingdao, China, Nolandia orientalis spec.nov. and Pinacocoleps similis (Kahl, 1933) comb.nov. (Ciliophora, Colepidae). European Journal of Protistology, 46 (4): 254 - 262. doi: 10.1016/j.ejop.2010.06.002

Zhang Q, Yi Z, Song W, Al-Rasheid KAS, WARREN A (2010) The systematic positon of Paraspathidium Noland, 1937 (Ciliophora, Litostomatea?) inferred from primary SSU rRNA gene sequences and predicted secondard rRNA struture.. European Journal of Protistology, 46 (4): 280 - 288. doi: 10.1016/j.ejop.2010.05.001

Fan X, Chen X, Song W, Al-Rasheid KAS, Warren A (2010) Two new marine scuticociliates, Sathrophilus planus n. sp. and Pseudoplatynematum dengi n. sp., with improved definition of Pseudoplatynematum (Ciliophora, Oligohymenophora). European Journal of Protistology, 46 (3): 212 - 220. doi: 10.1016/j.ejop.2010.03.002

Xu H, WARREN A, Al-Rasheid KAS, Zhu M, Song W (2010) Plantonic protist communities in semi-enclosed mariculture waters: temporal dynamics of functional groups and their responses to environmental conditions.. Acta Oceanolica Sinica, 29 (4): 106 - 115. doi: 10.1007/s13131-010-0055-6

Jiang J, Zhang Q, WARREN A, Al-Rasheid KAS, Song W (2010) Morphology and SSU rRNA gene-based phylogeny of two marine Euplotes species, E.orientalis spec. nov. and E.raikovi Agamaliev, 1966 (Ciliophora, Euplotida).. European Journal of Protistology, 46 (2): 121 - 132. doi: 10.1016/j.ejop.2009.11.003

Li L, Song W, Al-Rasheid KAS, WARREN A, Li Z, Xu Y, Shao C (2010) Morphology and morphogenesis of a new marine hypotrichous ciliate (Protozoa, Ciliophora, Pseudoamphisiellidae), including a report on the small subunit rRNA gene sequence. Zoological Journal of the Linnean Society, 158 (2)doi: 10.1111/j.1096-3642.2009.00546.x

Chen XR, Gao S, Song WB, Al-Rasheid KAS, WARREN A, Gong J, Lin XF (2010) Parabirojimia multinucleata spec. nov and Anteholosticha scutellum (Cohn, 1866) Berger, 2003, marine ciliates (Ciliophora, Hypotrichida) from tropical waters in southern China, with notes on their small-subunit rRNA gene sequences. International Journal of Systematic and Evolutionary Microbiology, 60 (1): 234 - 243. doi: 10.1099/Ijs.0.008037-0

Fan X, Chen X, Song W, Al-Rasheid KAS, WARREN A (2010) Two marine scuticociliates, Sathrophilus planus n.sp. and Pseudoplatynematum, dengi n. sp., with improved definition of Pseudoplatymematum (Ciliophora, Oligohymenophora). European Journal of Protistology, 46 : 212 - 220.

Foissner W, Shi X, Wang R, WARREN A (2010) A reinvestigation of Neokeronopsis populations, including the description of N.asiatica n.sp. (Ciliophora, Hypotricha). Acta Protozoologica, 49 (2): 87 - 105.

Shao C, Zhang QQ, Al-Rasheid KAS, WARREN A, Song WB (2010) Ontogenesis and Molecular Phylogeny of the Marine Ciliate Diophryopsis hystrix: Implications for the Systematics of the Diophrys-like Species (Ciliophora, Spirotrichea, Euplotida). Journal of Eukaryotic Microbiology, 57 (1): 33 - 39. doi: 10.1111/j.1550-7408.2009.00442.x

WARREN A, Salvado H, CURDS CR, ROBERTS DM (2010) Protozoa in activated sludge. , Serviour R, Nielsen PH, The Microbiology of Activated Sludge.. IWA Press. London, UK.

Yi Z, Lin X, WARREN A, Al-Rasheid KAS, Song W (2010) Molecular phylogeny of Nothoholosticha (Protozoa, Ciliophora, Urostylida) and systematic relationships of the Holosticha-complex.. Systematics and Biodiversity, 8 (1): 149 - 155. doi: 10.1080/14772000903523493

Shao C, Song W, WARREN A, Al-Rasheid KA (2009) Morphogenesis of Kiitrichia marine Nozawa, 1941 (Ciliophora, Spirotrichea), a possible model for the ancestor of hypotrichs s. l.. European Journal of Protistology, 45 (4): 292 - 304. doi: 10.1016/j.ejop.2009.07.002

Li L, Zhang Q, Hu X, WARREN A, Al-Rasheid KAS, Al-Khedheiry AA, Song W (2009) A redescription of the marine hypotrichous ciliate, Nothoholosticha fasciola (Kahl, 1932) nov. gen., nov. comb.(Ciliophora:Urostylida) with brief notes on its cellular reorganization and SSrRNA gene sequence. European Journal of Protistology, 45 (3): 237 - 248. doi: 10.1016/j.ejop.2009.01.004

GONG J, Stoeck T, Yi Z, Miao M, Zhang Q, ROBERTS DM, WARREN A, Song W (2009) Small subunit rRNA phylogenies show that the Class Nassophorea is not monophyletic (Phylum Ciliophora). Journal of Eukaryotic Microbiology, 56 (4): 339 - 347. doi: 10.1111/j.1550-7408.2009.00413.x

Li L, Song W, Shin MK, Warren A, Al-Rasheid KAS, Al-Khedhairy AA, Al-Arifi S (2009) Reconsideration of the phylogenetic positions of three stichotrichous genera Holosticha, Anteholosticha and Pseudokeronopsis (Spirotrichea: Ciliophora) inferred from complete SSU rRNA gene sequences. Progress in Natural Science, 19 (6): 769 - 773. doi: 10.1016/j.pnsc.2008.06.027

Yi Z, Song W, Gong J, WARREN A, Al-Rasheid , K. A. S. , AL-Arifi S, Al-Khedhairy AA (2009) Phylogeny of six oligohymenophoreans (Protozoa, Ciliophora) inferred from small subunit rRNA gene sequences. Zoologica Scripta, 38 (3): 323 - 331. doi: 10.1111/j.1463-6409.2008.00371.x

Chen X, WARREN A, Song W (2009) Taxonomic studies on a new marine ciliate, Apocoleps magnus gen. nov., spec. nov. (Ciliophora, Colepidae), isolated from Qingdao, China. Journal of the Ocean University of China, 8 : 317 - 321.

Chen X, WARREN A, Song W (2009) Suctoria. , Song W, WARREN A, Hu X, Free-living Ciliates in the Bohai and Yellow Seas, China., pp. 135 - 153. Science Press. Beijing.

GONG J, WARREN A, Song W (2009) Cyrtophorids. , Song W, WARREN A, Hu X, Free-living Ciliates in the Bohai and Yellow Seas., pp. 49 - 92. China, Science Press. Beijing.

Hu X, Song W, WARREN A (2009) Scuticociliatids. , Song W, WARREN A, Hu X, Free-living Ciliates in the Bohai and Yellow Seas, China., pp. 171 - 216. Science Press. Beijing.

Hu X, WARREN A, Song W (2009) Hypotrichs. , Song W, WARREN A, Hu X, Free-living Ciliates in the Bohai and Yellow Seas, China.., pp. 353 - 413. Science Press. Beijing.

Ji D, Sun P, WARREN A, Song W (2009) Colonial sessilid peritrichs. , Song W, WARREN A, Hu X, Free-living Ciliates in the Bohai and Yellow Seas, China., pp. 257 - 286. Science Press. Beijing.

Ji D, Xu H, Choi JK, WARREN A, Song W (2009) Taxonomic characterization of three marine peritrichous ciliates from China: Zoothamnium alrasheidi spec. nov., Z. marinum Kahl, 1933 and Z. vermicola Precht, 1935 (Ciliophora: Peritrichia). Acta Protozoologica, 48 (2): 171 - 183.

Lin X, WARREN A, Song W (2009) Pleurostomatids. , Song W, WARREN A, Hu X, Free-living Ciliates in the Bohai and Yellow Seas, China., pp. 93 - 134. Science Press. Beijing.

Long H, Song W, WARREN A (2009) Haptorid and other lower Kinetofragminophoran ciliates. , Song W, WARREN A, Hu X, Free-living Ciliates in the Bohai and Yellow Seas, China., pp. 19 - 47. Science Press. Beijing.

Mazei Y, Gao S, WARREN A, Li L, Li J, Song W, Esaulov A (2009) A reinvestigation of the marine ciliate Trachelocerca ditis (Wright, 1982) Foissner and Dragesco, 1996 (Ciliophora, Karyorelictea) from the Yellow Sea and an assessment of its phylogenetic position inferred from the small subunit rRNA gene sequence. Acta Protozoologica, 48 (3): 213 - 221.

Shao C, WARREN A, Song W (2009) Heterotrichs and related groups. , Song W, WARREN A, Hu X, Free-living Ciliates in the Bohai and Yellow Seas, China., pp. 287 - 306. Science Press. Beijing.

Song W, Shao C, Yi Z, Li L, WARREN A, Al-Rasheid KAS, Yang J (2009) The morphology, morphogenesis and SSrRNA gene sequence of a new marine ciliate, Diophrys apoligothrix spec. nov. (Ciliophora; Euplotida). European Journal of Protistology, 45 (1): 38 - 50. doi: 10.1016/j.ejop.2008.06.002

Song W, WARREN A (2009) Protohypotrichids, discocephalids and euplotids. , Song W, WARREN A, Hu X, Free-living Ciliates in the Bohai and Yellow Seas, China., pp. 415 - 459. Science Press. Beijing.

Sun P, Ji D, WARREN A, Song W (2009) Solitary sessilid peritrichs. , Song W, WARREN A, Hu X, Free-living Ciliates in the Bohai and Yellow Seas, China., pp. 217 - 256. Science Press. Beijing.

Wang Y, Song W, WARREN A, Al-Rasheid KAS, Al-Quraishy SA, Al-Farraj SA, Hu X, Pan H (2009) Descriptions of two new marine scuticociliates, Pleuronema sinica n. sp. and P. wilberti n. sp. (Ciliophora: Scuticociliatida), from the Yellow Sea, China. European Journal of Protistology, 45 (1): 29 - 37. doi: 10.1016/j.ejop.2008.06.001

Wang Y, WARREN A, Song W (2009) Hymenostomatids. , Song W, WARREN A, Hu X, Free-living Ciliates in the Bohai and Yellow Seas, China., pp. 155 - 169. Science Press. Beijing.

Xu D, WARREN A, Song W (2009) Oligotrichs. , Song W, WARREN A, Hu X, Free-living Ciliates in the Bohai and Yellow Seas, China., pp. 307 - 351. Science Press. Beijing.

Zhan Z, Xu K, WARREN A, Gong Y (2009) Reconsideration of phylogenetic relationships of the subclass Peritrichia (Ciliophora, Oligohymenophorea) based on small subunit rRNA gene sequences, with the establishment of a new subclass Mobilia Kahl, 1933. Journal of Eukaryotic Microbiology, 56 (6): 552 - 558. doi: 10.1111/j.1550-7408.2009.00435.x

Shao C, Song W, Li L, WARREN A, Al-Rasheid KAS, Al-Quraishy SA, Al-Farraj SA, Lin X (2008) Systematic position of Discocephalus-like ciliates (Ciliophora: Spirotrichea) inferred from SSU rDNA and ontogenetic information. International Journal of Systematic and Evolutionary Microbiology, 58 (12): 2962 - 2972. doi: 10.1099/ijs.0.65781-0

Xu H, Song W, WARREN A, Al-Rasheid KAS, Al-Farraj SA, GONG J, Hu X (2008) Planktonic protist communities in a semi-closed mariculture pond: structural variation and correlation with environmental conditions. Journal of the Marine Biological Association of the UK, 88 (7): 1353 - 1362. doi: 10.1017/S0025315408002129

Yi Z, Song W, Shao C, WARREN A, Al-Rasheid KAS, ROBERTS DM, Miao M, Al-Quraishy SA, Chen Z (2008) Phylogeny of some systematically uncertain urostyloids– Apokeronopsis, Metaurostylopsis, Thigmokeronopsis (Ciliophora, Stichotrichia)estimated with small subunit rRNA gene sequence information: Discrepancies and agreements with morphological data. European Journal of Protistology, 44 (4): 254 - 262. doi: 10.1016/j.ejop.2007.12.002

Miao M, WARREN A, Song W, Wang S, Shang H, Chen Z (2008) Analysis of the internal transcriber spacer 2 (ITS2) region of scuticociliates and related taxa (Ciliophora, Oligohymenophorea) to infer their evolution and phylogeny. Protist, 159 (4): 519 - 533. doi: 10.1016/j.protis.2008.05.002

Li L, Song W, WARREN A, Al-Rasheid KAS, ROBERTS D, Yi Z, Al-Farraj SA, Hua X (2008) Morphology and morphogenesis of a new marine ciliate, Apokeronopsis bergeri nov. spec. (Ciliophora, Hypotrichida), from the Yellow Sea, China. European Journal of Protistology, 44 (3): 208 - 219. doi: 10.1016/j.ejop.2008.01.001

Yi Z, Chen Z, WARREN A, ROBERTS D, Al-Rasheid KAS, Miao M, Gao S, Shao C, Song W (2008) Molecular phylogeny of Pseudokeronopsis (Protozoa, Ciliophora, Urostylida), with reconsideration of three closely related species at inter- and intra-specific levels inferred from the small subunit ribosomal RNA gene and the ITS1-5.8S-ITS2 region sequences. Journal of Zoology, 275 (3): 268 - 275. doi: 10.1111/j.1469-7998.2008.00438.x

Lin X, Li J, Gong J, WARREN A, Song W (2008) Taxonomic studies on three marine pleurostomatid ciliates, Litonotus bergeri nov. spec., L. blattereri nov. spec. and L. petzi nov. spec. (Ciliophora, Pleurostomatida) from North China Sea. European Journal of Protistology, 44 (2): 91 - 102. doi: 10.1016/j.ejop.2007.08.005

Hu X, WARREN A, Song W (2008) Stomatogenesis and morphological re-description of the marine ciliate, Philasterides armatalis (Protozoa: Ciliophora: Scuticociliatida). Journal of the Marine Biological Association of the United Kingdom, 881 (1): 29 - 34. doi: 10.1017/S0025315408000088

Yi Z, Song W, WARREN A, ROBERTS DM, Al-Rasheid KAS, Chen Z, Al-Farraj SA, Hu X (2008) A molecular phylogenetic investigation of Pseudoamphisiella and Parabirojimia (Protozoa, Ciliophora, Spirotrichea), two genera with ambiguous systematic positions. European Journal of Protistology, 44 (1): 45 - 53. doi: 10.1016/j.ejop.2007.08.002

Chen X, Miao M, Song W, WARREN A, Al-Rasheid KAS, Al-Farraj SA, Al-Quaraishy SA (2008) Redescriptions of two poorly known marine suctorian ciliates, Ephelota truncata Fraipont, 1878 and Ephelota mammillata Dons, 1918 (Protozoa, Ciliophora, Suctoria), from Qingdao, China. Acta Protozoologica, 47 (3): 247 - 256.

Chen X, Song W, WARREN A, Al-Rasheid KAS, Al-Farraj SA, Al-Quraishy SA, Gong J (2008) Redefinitions of two marine suctorian ciliates, Ephelota gemmipara (Hertwig, 1876) Bütschli, 1889 and E. crustaceorum Haller, 1880 (Ciliophora, Suctoria), with a brief description of the asexual reproduction process in E. gemmipara. Acta Protozoologica, 47 (2): 113 - 124.

Li L, Hu X, WARREN A, Al-Rasheid KAS, Al-Farraj SA, Shao C, Song W (2008) Divisional morphogenesis in the marine ciliate Anteholosticha manca (Kahl, 1932) Berger, 2003 (Ciliophora: Urostylida). Acta Oceanologica Sinica, 27 : 157 - 163.

Li L, Shao C, Yi Z, Song W, WARREN A, Al-Rasheid KAS, Al-Farraj SA, Al-Quaishy SA, Zhang Q, Hu X, Zhu M, Ma H (2008) Redescriptions and SSrRNA gene sequence analyses of two marine species of Aspidisca (Ciliophora, Euplotida), with notes on the morphogenesis in A. orthopogon. Acta Protozoologica, 47 (2): 83 - 94.

Li L, Song W, Al-Rasheid. KAS, WARREN A, Roberts D, Gong J, Zhang Q, Wang Y, Hu X (2008) Two discocephalid ciliates, Paradiscocephalus elongatus nov. gen., nov. spec. and Discocephalus ehrenbergi Dragesco, 1960, from the Yellow Sea, China (Ciliophora, Hypotrichida, Discocephalidae). Acta Protozoologica, 47 (4): 353 - 362.

Li L, Song W, WARREN A, Shin MK, Chen Z, Ji D, Sun P (2008) Reconsideration of the phylogenetic positions of five peritrich genera, Vorticella, Pseudovorticella, Zoothamnopsis, Zoothamnium, and Epicarchesium (Ciliophora, Peritrichia, Sessilida), based on small subunit rRNA gene sequences. Journal of Eukaryotic Microbiology, 55 (5): 448 - 456. doi: 10.1111/j.1550-7408.2008.00351.x

Shao C, Miao M, Song W, WARREN A, Al-Rasheid KAS, Al-Quraishy SA, Al-Farraj SA (2008) Studies on two marine Metaurostylopsis spp. from China with notes on morphogenesis in M. sinica nov. spec. (Ciliophora, Urostylida). Acta Protozoologica, 47 (2): 95 - 112.

Wang Y, Miao M, Zhang Q, Gao S, Song W, Al-Rasheid KAS, WARREN A, Ma H (2008) Three marine interstitial scuticociliates, Schizocalyptra similis sp. n., S. sinica sp. n. and Hippocomos salinus Small and Lynn, 1985 (Ciliophora: Scuticociliatida), isolated from Chinese coastal waters. Acta Protozoologica, 47 : 377 - 387.

Wang Y, Song W, Hu X, WARREN A, Chen X, Al-Rasheid KAS (2008) Descriptions of two new marine species of Pleuronema, P. czapikae sp. n. and P. wiackowskii sp. n. (Ciliophora: Scuticociliatida), from the Yellow Sea, North China. Acta Protozoologica, 47 (1): 35 - 45.

Xu D, Sun P, Song W, WARREN A (2008) Studies on a new endocommensal ciliate, Strombidium foissneri nov. sp. (Ciliophora, Oligotrichida), from the intestine of the sea urchin Hemicentrotus pulcherrimus (Camarodonta, Echinoida). Denisia, 23 : 273 - 278.

Peng X, Shi X, WARREN A (2007) Establishment of a new peritrich genus, Pseudepistylis n. gen. (Ciliophora: Peritrichia: Epistylididae), with a description of a new freshwater species, Pseudepistylis songi n. sp. from Wenzhou, China. Zootaxa, 1524 (1524): 35 - 45.

Gong J, Song W, WARREN A, Lin X, ROBERTS DM (2007) Microscopical observations on four marine Dysteria species (Ciliophora, Cyrtophorida). European Journal of Protistology, 43 (2): 147 - 161. doi: 10.1016/j.ejop.2007.01.002

Chen X, Song W, Al-Rasheid KAS, WARREN A, Long H, Shao C, Al-Farraj SA, GONG J, Hu X (2007) The morphology of three marine heterotrichous ciliates, Condylostentor auriculatus (Kahl, 1932) Jankowski, 1978, Condylostoma minutum Bullington, 1940 and C. spatiosum Ozaki & Yagiu in Yagiu, 1944 (Ciliophora: Heterotrichida). Acta Protozoologica, 46 (4): 289 - 309.

Day JG, Achilles-Day U, Brown S, WARREN A (2007) Cultivation of algae and protozoa. , Hurst CJ, Crawford RL, Garland JL, Lipson DA, Mills AL, Stetzenbach LD, Manual of Environmental Microbiology, 3rd edition., pp. 79 - 92. ASM Press. Washington.

GONG J, Kim SJ, Kim SY, Min GS, ROBERTS DM, WARREN A, Choi JK (2007) Taxonomic redescriptions of two ciliates, Protogastrostyla (Gastrostyla) pulchra nov. gen. nov. comb. and Hemigastrostyla enigmatica (Ciliophora: Spirotrichea: Stichotrichia), with phylogenetic analyses based on 18S and 28S rRNA gene sequences. Journal of Eukaryotic Microbiology, 54 (0): 468 - 478. doi: 10.1111/j.1550-7408.2007.00288.x

Holland T, Warren A, JOHANSON Z, Long JA, Parker K, Garvey J (2007) A new species of Barameda (Rhizodontida) and heterochrony in the rhizodontid pectoral fin. Journal of Vertebrate Paleontology, 27 : 295 - 315.

Long H, Song W, WARREN A, Al-Rasheid KAS, Gong J, Chen X (2007) Two new ciliates from the north China seas, Schizocalyptra aeschtae nov. spec. and Sathrophilus holtae nov. spec., with a new definition of the genus Sathrophilus (Ciliophora, Oligohymenophora). Acta Protozoologica, 46 (3): 229 - 245.

Shao C, Hu X, WARREN A, Al-Rasheid KAS, Al-Quraishy SA, Song W (2007) Morphogenesis in the marine spirotrichous ciliate Apokeronopsis crassa (Claparede & Lachmann, 1858) nov. comb. (Ciliophora: Stichotrichia), with the establishment of a new genus, Apokeronopsis n.g., and redefinition of the genus Thigmokeronopsis. Journal of Eukaryotic Microbiology, 54 (4): 392 - 401. doi: 10.1111/j.1550-7408.2007.00278.x

Shao C, Song W, Li L, WARREN A, Hu X (2007) Morphological and morphogenetic redescriptions of the stichtrich ciliate Diaxonemma trimarginata Jankowski, 1979 (Ciliophora, Stichtrichia, Urosylida). Acta Protozoologica, 46 (1): 25 - 39.

Shi X, Hu X, WARREN A, Liu G (2007) Redescription of morphology and morphogenesis of the freshwater ciliate Pseudokeronopsis similis (Stokes, 1886) Borror & Wicklow, 1983 (Ciliophora: Urostylida). Acta Protozoologica, 46 (1): 41 - 54.

SONG WEIBO, WILBERT NORBERT, AL-RASHEID KAHLED, WARREN ALAN, SHAO CHEN, LONG HONGAN, YI ZHENZHEN, LI LIQIONG (2007) Redescriptions of Two Marine Hypotrichous Ciliates, Diophrys irmgard and D. hystrix (Ciliophora, Euplotida), with a Brief Revision of the Genus Diophrys. The Journal of Eukaryotic Microbiology, 54 (3): 283 - 296. doi: 10.1111/j.1550-7408.2007.00266.x

Song W, Wilbert N, Al-Rasheid KAS, WARREN A, Shao C, Long H, Yi Z, Li L (2007) Redscriptions of two marine hypotrichous ciliates, Diophrys irmgard and D. hystrix (Ciliophora, Euplotida), with a brief revision of the genus Diophrys. Journal of Eukaryotic Microbiology, 54 : 283 - 296.

Wang R, Qiu Z, Chen J, WARREN A, Song W (2007) Morphogenesis of the freshwater ciliate Neokeronopsis spectabilis (Kahl, 1932) Warren et al., 2002, based on a China population (Ciliophora: Urostylidae). Journal of Eukaryotic Microbiology, 54 (2): 184 - 190. doi: 10.1111/j.1550-7408.2007.00248.x

Xu D, Song W, WARREN A, ROBERTS D, Hu X (2007) Redescriptions of two marine planktonic ciliates from China, Parastrombidium faurei (Kahl, 1932) Maeda, 1986 and Strombidium capitum (Leegaard, 1915) Kahl, 1932 (Ciliophora, Oligotrichea). European Journal of Protistology, 43 (1): 27 - 35. doi: 10.1016/j.ejop.2006.09.005

Fyda J, Kennaway G, Adamus K, WARREN A (2006) Ultrastructural events in the predator induced defence response of Colpidium kleini (Ciliphora: Hymenostomatia). Acta Protozoologica, 45 (4): 461 - 464.

Sun P, Song W, WARREN A (2006) Taxonomic characterisation of two marine peritrichous ciliates, Epicarchesium corlissi n.sp. and Pseudovorticella jiangi n.sp. (Ciliophora: Peritrichia), from northern China. European Journal of Protistology, 42 (4): 281 - 289. doi: 10.1016/j.ejop.2006.07.004

Xu D, Song W, Lin XK, WARREN A (2006) On two marine oligotrich ciliates, Spirostomum agathae n. sp. and S. schizostomum (Kahl, 1932) n. comb. from China, with a key to the identification of seven well-characterized Spirostrombidium spp. (Ciliophora: Oligotrichida). Acta Protozoologica, 45 (4): 433 - 442.

Ma H, Song W, WARREN A, ROBERTS D, Gong J, Al-Rasheid K (2006) Redescription of the marine scuticociliate Glauconema trihymene Thompson, 1966 (Protozoa: Ciliophora): life cycle and stomatogenesis. Zootaxa, 1296 (1296): 1 - 17.

Song W, WARREN A, ROBERTS D, Wilbert N, Li L, Sun P, Hu X, Ma H (2006) Comparison and redefinition of four marine, coloured Pseudokeronopsis spp. (Ciliophora, Hypotrichida), with emphasis on their living morphology. Acta Protozoologica, 45 (3): 271 - 287.

Li L, Song W, WARREN A, Wang Y, Ma H, Hu X, Chen Z (2006) Phylogenetic position of the marine ciliate Cardiostomatella vermiforme (Kahl, 1928) Corliss, 1960 inferred from complete SSrRNA gene sequence, with establishment of a new order Loxocephalida n. ord. (Protozoa, Ciliophora, Oligohymenophorea). European Journal of Protistology, 42 (2): 107 - 114. doi: 10.1016/j.ejop.2006.01.004

Shang H, Song W, WARREN A, Li L, Chen Z (2006) Phylogenetic positions of two marine ciliates, Metanophrys similis and Pseudocohnilembus hargisi (Protozoa: Ciliophora: Scuticociliata), inferred from complete small subunit rRNA gene sequences. Progress in Natural Science, 16 (4): 373 - 378. doi: 10.1080/10020070612330007

Ji D, Song W, WARREN A (2006) Redescriptions of three marine peritrichous ciliates, Zoothamnium alternans Claparède and Lachmann, 1859, Z. sinense Song, 1991 and Z. commune Kahl, 1933 (Ciliophora, Peritrichia), from north China. Acta Protozoologica, 45 (1): 27 - 39.

Shao C, Song W, WARREN A, Al-Rasheid KAS, Yi Z, Gong J (2006) Morphogenesis of the marine ciliate, Pseudoamphisiella alveolata (Kahl, 1932) Song & Warren, 2000 (Ciliophora, Stichotrichia, Urostylida) during binary fission. Journal of Eukaryotic Microbiology, 53 (5): 388 - 396. doi: 10.1111/j.1550-7408.2006.00117.x

Xu D, Song W, WARREN A (2006) Morphology and infraciliature of two new species of marine oligotrich ciliates (Ciliophora: Oligotrichida) from China. Journal of Natural History, 40 : 1287 - 1299. doi: 10.1080/00222930600913925

Lin X, Song W, WARREN A (2005) Two new marine pleurostomatid ciliates from China, Amphileptus gui nov. spec. and Amphileptus yuianus nov. spec. (Ciliophora, Pleurostomatida). European Journal of Protistology, 41 (3): 163 - 174. doi: 10.1016/j.ejop.2005.01.002

Song WB, Warren A, Roberts D, Shang HM, Ma HW, Wilbert N, Miao M, Yi ZZ (2005) Re-evaluation of the phylogenetic position of the genus Dexiotrichides (Protozoa, Ciliophora, Scuticociliatida) inferred from stomatogenetic and molecular information for Dexiotrichides pangi. PROGRESS IN NATURAL SCIENCE, 15 (8): 700 - 707.

Ramirez E, Robles E, Bonilla P, Sainz G, Lopez M, De La Cerda JM, WARREN A (2005) Occurrence of pathogen free-living amoeba and bacterial indicators in a constructed wetland treating domestic wastewater from a single household. Engineering in Life Sciences, 5 (3): 253 - 258. doi: 10.1002/elsc.200420071

Vickerman K, Warren A, Preston T (2005) Meeting Report: Evolution of Protozoa and Other Protists, Linnean Society, London, September 13, 2004. Protist, 156 (1): 9 - 17. doi: 10.1016/j.protis.2005.02.003

Gong J, Song W, WARREN A (2005) Periphytic ciliate colonization: annual cycle and responses to environmental conditions. Aquatic Microbial Ecology, 39 (2): 159 - 170. doi: 10.3354/ame039159

Lin X, Song W, WARREN A (2005) Taxonomic studies on three marine pleurostomatid ciliates: Kentrophyllum verrucosum (Stokes, 1893) Petz, Song & Wilbert, 1995, Epiphyllum soliforme (Fauré-Frémiet, 1908) gen. n., comb. n. and Amphileptus sikorai sp. n., with the establishment of a new genus Epiphyllum (Ciliophora: Pleurostomatida). Acta Protozoologica, 44 (2): 129 - 145.

Lin X, Song W, Wilbert N, WARREN A (2005) Two new marine pleurostomatid ciliates from China, Loxophyllum jinni and L. qiuianum (Ciliophora: Pleurostomatida). Acta Protozoologica, 44 (2): 147 - 157.

Fyda J, WARREN A, Wolińska J (2005) An investigation of predator-induced defence responses in ciliated protozoa. Journal of Natural History, 39 (18): 1431 - 1422. doi: 10.1080/00222930400004396

Gong J, Song W, WARREN A (2005) Updating the ciliate genus Chlamydodon Ehrenberg, 1835, with redescriptions of three species (Ciliophora: Cyrtophorida). Acta Protozoologica, 44 : 19 - 32.

Ji DD, Song W, WARREN A (2005) Two new marine sessile peritrichous ciliates (Protozoa: Ciliophora). Acta Zootaxonomica Sinica, 30 : 702 - 705.

Ma HW, Song W, Gong J, WARREN A (2005) Reconsideration of stomatogenesis in Uronemia marinum Dujardin, 1841 during asexual division (Protozoa: Scuticociliatida). Acta Zoologica Sinica, 50 : 823 - 827.

Song W, WARREN A, ROBERTS D, Shang H, Ma H (2005) Re-evaluation of the phylogenetic position of the genus Dexiotrichides inferred from stomatogenetic and molecular information for Dexiotrichides pangi. Progress in Natural Science, 15 : 700 - 707.

Vickerman K, WARREN A, Preston T (2005) Evolution of protozoa and other protists. Meeting Report. Protist, 156 : 9 - 17.

Xu H, Song W, Lu L, WARREN A (2005) Tolerance of ciliated protozoan Paramecium bursaria to ammonia and nitrites. Chinese Journal of Oceanology and Limnology, 23 : 349 - 353.

Ji DD, Song W, WARREN A (2004) Rediscovery and redescription of the marine peritrichous ciliate Epicarchesium abrae (Precht, 1935) nov. comb. (Protozoa, Ciliophora, Peritrichia). European Journal of Protistology, 40 (3): 219 - 224. doi: 10.1016/j.ejop.2004.03.002

Lin X, Song W, WARREN A (2004) Redescription of the rare marine ciliate, Prodiscocephalus borrori (Wicklow, 1982) from shrimp-culturing waters near Qingdao, China, with redefinitions of the genera Discocephalus, Prodiscocephalus and Marginotricha (Ciliophora, Hypotrichida, Discocephalidae). European Journal of Protistology, 40 (2): 137 - 146. doi: 10.1016/j.ejop.2003.12.003

Shi X, WARREN A, Yu Y, Shen Y (2004) Infraciliature and myoneme system of Campanella umbellaria (Protozoa, Ciliophora, Peritrichida). Journal of Morphology, 261 (1): 43 - 51. doi: 10.1002/jmor.10225

Hu XZ, WARREN A, Song W (2004) Observations on the morphology and morphogenesis of a new marine hypotrich ciliate (Ciliophora, Hypotrichida) from China. Journal of Natural History, 38 (9): 1059 - 1069. doi: 10.1080/0022293031000069256

Xu H, Song W, WARREN A (2004) An investigation of the tolerance to ammonia of the marine ciliate Euplotes vannus (Protozoa: Ciliophora). Hydrobiologia, 519 : 189 - 195. doi: 10.1023/B:HYDR.0000026505.91684.ab

Ji DD, Song WB, WARREN A (2004) Pseudovorticella paracratera n. sp., a new marine peritrich ciliate (Ciliophora : Peritrichida) from north China. Hydrobiologia, 515 : 49 - 57. doi: 10.1023/B:HYDR.0000027317.26498.6a

Hu X, WARREN A, Suzuki T (2004) Morphology and morphogenesis of two marine ciliates, Pseudokeronopsis pararubra sp.n. and Amphistella annulata from China and Japan (Protozoa: Ciliophora). Acta Protozoologica, 43 : 351 - 368.

Shi X, Hu X, WARREN A (2003) Rediscovery of Gastrostyla setifera (Engelmann, 1862) Kent, 1882: morphology and divisional morphogenesis (Ciliophora, Hypotrichida). Journal of Natural History, 37 (12): 1411 - 1422. doi: 10.1080/00222930110121006

Chen Z, Song W, WARREN A (2003) Species separation and identification of Uronychia spp. (Hypotrichia: Ciliophora) using RAPD Fingerprinting and ARDRA Riboprinting. Acta Protozoologica, 42 (2): 83 - 90.

Ma H, Song W, Hu Z, WARREN A (2003) Morphology and stomatogenesis of Pseudocohnilembus hargisi (Ciliophora: Scuticociliatida). Journal of The Marine Biological Association of the United Kingdom, 83 (2): 399 - 405. doi: 10.1017/S0025315403007240h

Shang H, Song W, WARREN A (2003) Phylogenetic positions of two ciliates, Paranophrys magna and Mesanophrys carcini (Protozoa, Ciliophora, Oligohymenophorea), within the subclass Scuticociliatia inferred from complete small subunit rRNA gene sequences. Acta Protozoologica, 42 : 171 - 181.

Song WB, WARREN A, Ji DD, Wang M, Al-Rasheid KAS (2003) New contributions to two heterotrichous ciliates, Folliculina simplex (Dons, 1917), Condylostoma curva Burkovsky,1970 and one licnophorid, Licnophora lyngbycola Faure-Fremiet,, 1937 (Protozoa, Ciliophora): Descriptions of morphology and infraciliature. Journal of Eukaryotic Microbiology, 50 (6): 449 - 462. doi: 10.1111/j.1550-7408.2003.tb00271.x

Hu XZ, Song WB, Warren A (2002) Observations on the morphology and morphogenesis of a new marine urostylid ciliate, Parabirojimia similis nov gen., nov spec. (Protozoa, Ciliophora, Hypotrichida). EUROPEAN JOURNAL OF PROTISTOLOGY, 38 (4): 351 - 364. doi: 10.1078/0932-4739-00880

Shi XL, Warren A, Song WB (2002) Studies on the morphology and morphogenesis of Allotricha curdsi sp n. (Ciliophora : Hypotrichida). ACTA PROTOZOOLOGICA, 41 (4): 397 - 405.

Kinner NE, Harvey RW, Shay DM, Metge DW, Warren A (2002) Field Evidence for a Protistan Role in an Organically-Contaminated Aquifer. Environmental Science & Technology, 36 (20): 4312 - 4318. doi: 10.1021/es020611m

Gong J, Song WB, Warren A (2002) Redescriptions of two marine cyrtophorid ciliates, Dysteria cristata (Gourret and Roeser,, 1888) Kahl,, 1931 and Dysteria monostyla (Ehrenberg, 1838) Kahl, 1931 (Protozoa, Ciliophora, Cyrtophorida), from China. EUROPEAN JOURNAL OF PROTISTOLOGY, 38 (3): 213 - 222. doi: 10.1078/0932-4739-00862

Warren A, Fyda J, Song WB (2002) The morphology of the poorly-known freshwater urostylid ciliate Neokeronopsis spectabilis (Kahl, 1932) nov gen. nov comb., (Ciliophora : Urostylidae), with notes on its morphogenesis. EUROPEAN JOURNAL OF PROTISTOLOGY, 38 (2): 195 - 206. doi: 10.1078/0932-4739-00820

Xu K, Song W, Warren A (2002) Taxonomy of trichodinids from the gills of marine fishes in coastal regions of the Yellow Sea, with descriptions of two new species of Trichodina Ehrenberg, 1830 (Protozoa: Ciliophora: Peritrichia). Systematic Parasitology, 51 (2): 107 - 120. doi: 10.1023/A:1014066915539

Decamp O, Warren A (2001) ABUNDANCE, BIOMASS AND VIABILITY OF BACTERIA IN WASTEWATERS: IMPACT OF TREATMENT IN HORIZONTAL SUBSURFACE FLOW CONSTRUCTED WETLANDS. Water Research, 35 (14): 3496 - 3501. doi: 10.1016/S0043-1354(01)00064-1

Song WB, Petz W, Warren A (2001) Morphology and morphogenesis of the poorly-known marine urostylid ciliate, Metaurostylopsis marina (Kahl, 1932) nov gen., nov comb. (Protozoa, Ciliophora, Hypotrichida). EUROPEAN JOURNAL OF PROTISTOLOGY, 37 (1): 63 - 76. doi: 10.1078/0932-4739-00802

Hodson M (2001) Effect of bone meal (calcium phosphate) amendments on metal release from contaminated soils — a leaching column study. Environmental Pollution, 112 (2): 233 - 243. doi: 10.1016/S0269-7491(00)00116-0

Xu K, Song W, Warren A, Choi JK (2001) Trichodinid ectoparasites (Ciliophora: Peritrichida) of some marine fishes from coastal regions of the Yellow Sea and Bohai Sea. Systematic Parasitology, 50 (1): 69 - 79. doi: 10.1023/A:1011865124047

Song W, Warren A (2000) Pseudoamphisiella alveolata (Kahl, 1932) nov. comb., a large marine hypotrichous ciliate from China (Protozoa, Ciliophora, Hypotrichida). European Journal of Protistology, 36 (4): 451 - 457. doi: 10.1016/S0932-4739(00)80051-6

Hu X, Song W, Warren A (2000) Divisional morphogenesis in the marine ciliate Holosticha warreni (Ciliophora: Hypotrichida). Journal of the Marine Biological Association of the UK, 80 (5): 785 - 788. doi: 10.1017/S0025315400002757

Song W, Wang M, Warren A (2000) Redescriptions of three marine ciliates, Strombidium elegans Florentin, 1901, Strombidium sulcatum Claparède & Lachmann, 1859 and Heterostrombidium paracalkinsi Lei, Xu & Song, 1999 (Ciliophora, Oligotrichida). European Journal of Protistology, 36 (3): 327 - 342. doi: 10.1016/S0932-4739(00)80010-3

Chen ZG, Song WB, Warren A (2000) Studies on six Euplotes spp. (Ciliophora : Hypotrichida) using RAPD fingerprinting, including a comparison with morphometric analyses. ACTA PROTOZOOLOGICA, 39 (3): 209 - 216.

Decamp O, Warren A (2000) Investigation of Escherichia coli removal in various designs of subsurface flow wetlands used for wastewater treatment. Ecological Engineering, 14 (3): 293 - 299. doi: 10.1016/S0925-8574(99)00007-5

Xu K, Song W, Warren A (2000) Observations on trichodinid ectoparasites (Ciliophora: Peritricha) from the gills of maricultured molluscs in China, with descriptions of three new species of Trichodina Ehrenberg, 1838. Systematic Parasitology, 45 (1): 17 - 24. doi: 10.1023/A:1006284629307

Song W, Wilbert N, Warren A (1999) Three new entocommensal ciliates from digestive tract of sea urchins of the Weddell Sea, Antarctica (Protozoa, Ciliophora). Polar Biology, 22 (4): 232 - 240. doi: 10.1007/s003000050415

Xu K, Song W, Warren A (1999) Trichodinid ectoparasites (Ciliophora: Peritrichida) from the gills of mariculture molluscs in China, with the descriptions of four new species of Trichodina Ehrenberg, 1838. Systematic Parasitology, 42 (3): 229 - 237. doi: 10.1023/A:1006071106844

Xu K, Song W, Warren A (1999) Trichodinid ectoparasites (Ciliophora: Peritrichida) from the gills of cultured marine fishes in China, with the description of Trichodinella lomi n. sp.. Systematic Parasitology, 42 (3): 219 - 227. doi: 10.1023/A:1006067005936

Song W, Warren A (1999) Observations on Morphogenesis in a marine ciliate Tachysoma ovata (Protozoa: Ciliophora: Hypotrichida). Journal of the Marine Biological Association of the UK, 79 (1): 35 - 38. doi: 10.1017/S0025315498000034

Decamp O, Warren A (1998) Bacterivory in ciliates isolated from constructed wetlands (reed beds) used for wastewater treatment. Water Research, 32 (7): 1989 - 1996. doi: 10.1016/S0043-1354(97)00461-2

Song W, Warren A, Hu X (1997) Morphology and Morphogenesis of Pseudoamphisiella lacazei (Maupas, 1883) Song, 1996 with Suggestion of Establishment of a New Family Pseudoamphisiellidae nov. fam. (Ciliophora, Hypotrichida). Archiv für Protistenkunde, 147 : 265 - 276. doi: 10.1016/S0003-9365(97)80053-X

Campbell CD, Warren A, Cameron CM, Hope SJ (1997) Direct toxicity assessment of two soils amended with sewage sludge contaminated with heavy metals using a protozoan (Colpoda steinii) bioassay.. Chemosphere, 34 (3): 501 - 514. doi: 10.1016/S0045-6535(96)00389-X

Decamp O, Warren A (1997) Observations on the morphology of Caenomorpha uniserialis Levander, 1894 (Ciliophora, heterotrichida) isolated from a wastewater treatment plant. ACTA PROTOZOOLOGICA, 36 (2): 105 - 110.

Nagasawa S, Warren A (1996) Redescription of Vorticella oceanica Zacharias, 1906 (Ciliophora: Peritrichia) with notes on its host, the marine planktonic diatom Chaetoceros coarctatum Lauder,1864. Hydrobiologia, 337 : 27 - 36. doi: 10.1007/BF00028504

Pérez-Uz B, Song W, Warren A (1996) Morphogenetic processes in division of Uronema filificum Kahl 1931 (Ciliophora: Scuticociliatida) and implications for its systematic position in the family uronematidae. European Journal of Protistology, 32 (2): 262 - 270. doi: 10.1016/S0932-4739(96)80025-3

Song WB, Warren A (1996) A redescription of the marine ciliates Uroleptus retractilis (Claparede and Lachmann, 1858) comb n and Epiclintes ambiguus (Muller, 1786) Butschli, 1889 (Ciliophora, Hypotrichida). ACTA PROTOZOOLOGICA, 35 (3): 227 - 234.

Novarino G, Warren A, Kinner NE, Harvey RW (1994) Protists from a sewage‐contaminated aquifer on cape cod, Massachusetts. Geomicrobiology Journal, 12 (1): 23 - 36. doi: 10.1080/01490459409377968

Forge TA, Berrow ML, Darbyshire JF, Warren A (1993) Protozoan bioassays of soil amended with sewage sludge and heavy metals, using the common soil ciliate Colpoda steinii. Biology and Fertility of Soils, 16 (4): 282 - 286. doi: 10.1007/BF00369305

Ram�rez E, Warren A, Rivera FN, Bonilla P, Rodr�uez S, Calder�n A, Ort�z R, Gallegos E (1993) An investigation of the pathogenic and non-pathogenic free-living amoebae in an activated-sludge plant. Water, Air, & Soil Pollution, 69 : 135 - 139. doi: 10.1007/BF00478354

Rivera FN, Rodr�guez S, Warren A, Bonilla P, Ram�rez E, Calder�n A, Ort�z R (1993) An investigation of the pathogenic and non-pathogenic free-living amoebae from the root zone method of wastewater treatment. Water, Air, & Soil Pollution, 69 : 93 - 98. doi: 10.1007/BF00478350

WARREN A (1991) A SCANNING ELECTRON-MICROSCOPIC STUDY OF THE MORPHOLOGY OF SCYPHIDIA-PHYSARUM LACHMANN, 1856 (CILIOPHORA, PERITRICHIDA). SCANNING MICROSCOPY, 5 (1): 281 - 286.

Roberts DM, Warren A (1987) An inexpensive apparatus for automatic continuous dehydration.. Stain Technol, 62 (4): 211 - 215.

Warren A (1985) A Redescription of the freshwater loricate ciliate, Stentor barrettiBarrett, 1870. Archiv für Protistenkunde, 129 : 145 - 153. doi: 10.1016/S0003-9365(85)80016-6

CAREY P, WARREN A (1983) THE ROLE OF SURFACE-TOPOGRAPHY IN THE TAXONOMY OF PERITRICH CILIATES. PROTISTOLOGICA, 19 (1): 73 - 89.

ROBERTS D, WARREN A, CURDS C (1983) MORPHOMETRIC ANALYSIS OF OUTLINE SHAPE APPLIED TO THE PERITRICH GENUS VORTICELLA. SYSTEMATIC ZOOLOGY, 32 (4): 377 - 388. doi: 10.2307/2413165

WARREN A (1983) AN UNUSUAL LORICATE STENTOR, STENTOR-BARRETTI BARRETT, 1870. JOURNAL OF PROTOZOOLOGY, 30 (3)

WARREN A, CAREY P (1983) LORICA STRUCTURE OF THE FRESH-WATER CILIATE PLATYCOLA-DECUMBENS EHRENBERG, 1830 (PERITRICHIDA, VAGINICOLIDAE). PROTISTOLOGICA, 19 (1): 5 - 20.

WARREN A, CAREY P (1982) THE LORICA STRUCTURE OF PLATYCOLA-DECUMBENS EHRENBERG, 1830 (CILIOPHORA, PERITRICHIDA). JOURNAL OF PROTOZOOLOGY, 29 (4): 641 - 641.

WARREN A, CAREY P (1982) TOPOGRAPHIC PATTERN-ANALYSIS OF PERITRICH CILIATES. JOURNAL OF PROTOZOOLOGY, 29 (2): 323 - 323.

WARREN A, ROBERTS D (1982) DOES VORTICELLA HAVE VITAL STATISTICS - A MATHEMATICAL-ANALYSIS OF SHAPE, WITH TAXONOMIC IMPLICATIONS. JOURNAL OF PROTOZOOLOGY, 29 (4): 641 - 642.