Stem group echinoids with:
Lower Carboniferous (Visean) to Upper Permian, USA, Europe.
The presence of specialised adoral pore-pairs (and presumably tube-feet) characterizes this clade. Interambulacral plating is more imbricate than in Palaechinidae, and there are primary tubercles, which are never seen in Palaechinidae. Lepidesthidae have expanded ambulacra with many columns, but the pore-pairs remain simple throughout.