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1b. Aboral interambulacral plates firmly sutured. |
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3a. Interambulacral plates with confluent, ovate areoles more or less throughout. Spines with smooth cortex |
3b. Interambulacral plates at ambitus and adapically with relatively separated, circular areoles; areoles confluent only adorally. Spines thorned or beaded If spines unknown 'Miocidaris' |
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4b. Primary spines with scattered coarse thorns along shaft. |